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Споровики
Паразитические простейшие. В связи с паразитическим образом жизни
происходит упрощение организации (исчезновение органоидов захвата и
приема пищи, пищеварительных и сократительных вакуолей). Происходит
усложнение жизненного цикла – смена хозяев, чередование бесполого и
полового размножения. Представители типа – малярийный плазмодий,
токсоплазма и др.
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202020202020202020202020202020202020202020202020202020202020202020202020
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40000002701ffff030000000000
Малярийный плазмодий вызывает у человека заболевание малярией. Заражение
человека происходит через укус малярийного комара (рода Anopheles),
который содержит возбудителя на стадии спорозоитов. Спорозоиты – тонкие,
червеобразные клетки, с током крови попадают в клетки печени, где
превращаются в шизонтов, размножающихся множественным делением –
шизогонией. Образуется следующая стадия паразита – мерозоиты. Они
внедряются в эритроциты и превращаются сначала в трофозоиты (питание
гемоглобином и рост), затем – в шизонты (размножение). Таким образом,
различают две формы шизогонии – в клетках печени и в эритроцитах. В
результате эритроцитарной шизогонии образуются 10-20 мерозоитов, которые
разрушают эритроцит, выходят в кровь и заражают следующие эритроциты.
Цикличность приступов малярии обусловлена цикличностью выходов
мерозоитов и продуктов их метаболизма из эритроцита в плазму крови.
После нескольких циклов шизогонии в эритроцитах образуются гамонты,
которые в организме комара превратятся в макрогаметы и микрогаметы.
Когда гамонты попадают в желудок комара, они превращаются в гаметы,
происходит копуляция, слияние гамет. Зигота подвижна и называется
оокинета. Оокинета мигрирует через стенку желудка комара и превращается
в ооцисту. Ядро ооцисты многократно делится и ооциста распадается на
огромное количество спорозоитов – до 10 000, этот процесс называется
спорогония. Спорозоиты мигрируют в слюнные железы комара. Таким
образом, в жизненном цикле малярийного плазмодия человек является
промежуточным хозяином (преэритроцитарная шизогония, эритроцитарная
шизогония, начало гаметогонии), а малярийный комар – окончательным
(завершение гаметогонии, оплодотворение и спорогония). У человека
в организме паразитируют четыре вида плазмодиев, период между выходами
мерозоитов в плазму крови у одного вида – 72 часа, заболевание получило
название четырехдневная малярия. У других видов – 48 часов – трехдневная
малярия. Кроме лихорадочного состояния развивается сильная анемия
(малокровие), содержание эритроцитов в 1 мм понижается, вместо 5 млн. их
количество уменьшается до 1 млн.
К типу споровиков относятся и токсоплазмы – внутриклеточные паразиты.
Развитие происходит со сменой хозяев, окончательным хозяином является
кошка, в ее кишечнике происходит половое размножение, промежуточными
хозяевами являются человек, грызуны, крупный и мелкий рогатый скот и
другие виды теплокровных животных. Токсоплазмоз проявляется в поражениях
лимфатической системы, тифоподобных заболеваниях, поражениях органов
зрения, нервной системы и др., у взрослых может быть бессимптомным.
Заражение человека происходит различными способами: а) перорально при
употреблении сырого или полусырого мяса или фарша; б) перорально с
овощами, фруктами, загрязненными ооциста-ми; в) через загрязненные
ооцистами руки и предметы; г) трансплацентарно от матери-носительницы к
плоду.
Инфузории
Класс инфузории
– наиболее высокоорганизованные простейшие. Органоидами движения служат
реснички, по строению сходные со жгутиками, но более короткие и
многочисленные. Тело покрыто прочной эластичной оболочкой, придающей ей
постоянную форму. У большинства инфузорий 2 ядра: большое и малое.
Большое ядро имеет полиплоидный набор хромосом и регулирует процессы
движения, питания, выделения, а также бесполое размножение,
осуществляемое поперечное деление клетки пополам. Малое ядро имеет
диплоидный набор хромосом и играет важную роль в половом процессе,
выступая в качестве носителя наследственной информации. Впервые
инфузорий обнаружили в воде, настоянной на различных травах («инфузум»
означает «настойка»).
Инфузория
– туфелька наиболее распространенный представитель, обитатель пресных
водоемов, длина тела 0,3 мм. Форма тела постоянная и напоминает подошву
туфли. Все тело равномерно покрыто ресничками, расположенными рядами, их
больше 10 тысяч. Работают они синхронно, совершая волнообразные
движения, обеспечивается это плотными цитоплазматическими нитями –
фибриллами (плавает тупым концом вперед). Между ресничками расположены
мелкие веретеновидные тельца – трихоцисты – органоиды защиты и
нападения, которые в ответ на раздражение с силой выбрасываются и
вонзаются в тело жертвы или врага. Питаются инфузории бактериями и
одноклеточными водорослями. Сбоку на теле имеется углубление –
предротовое углубление, ведущее в рот. На дне глотки в цитоплазме
образуется пищеварительная вакуоль, которая отделяется от глотки и
увлекается током цитоплазмы. При обилии пищи и нормальных температурных
условиях (15 градусов) пищеварительные вакуоли образуются, каждые 1-2
мин. В них пища переваривается и усваивается цитоплазмой, после чего
пищеварительная вакуоль, пройдя по часовой стрелке, подходит к заднему
концу тела, где через специальное отверстие в оболочке – порошицу
выбрасывает не переваренные остатки пищи наружу. Функцию осморегуляции
выполняют 2 сократительные вакуоли.
Размножается туфелька
Бесполым и половым способом. При бесполом размножении тело туфельки
вытягивается в длину, по экватору появляется перетяжка, которая делит
клетку пополам. Повторяется 1-2 раза в сутки, а через несколько
поколений бесполого размножения сменяется половым, протекающим по типу
конъюгации. В половом размножении большую роль играет малое ядро. В теле
каждого участника большое ядро разрушается, а малое ядро делится на 4
части (процесс мейоза, при котором число хромосом уменьшается вдвое).
Вскоре 3 новых ядра разрушаются, а четвертая вновь делится и образует в
каждой инфузории одно женское и одно мужское ядро. Мужское ядро
переходит в клетку своего партнера, где сливается с женским ядром. Таким
образом, при половом процессе происходит обмен ядерным материалом между
отдельными особями, которые получают новые признаки и свойства. Вскоре в
каждой из них ядро делится на большое и малое. При половом размножении
число особей не увеличивается, а обновляются наследственные свойства
организма, и возрастает его способность приспосабливаться к условиям
среды.
Характерной особенностью туфельки является раздражимость.
Это способность организма отвечать определенным образом на воздействия
окружающей среды. Это свойство характерно для всех живых существ.
Раздражитель – фактор среды; раздражение – воздействие раздражителя;
раздражимость – ответ организма на раздражение. Простейшие не имеют
нервной системы, они воспринимают раздражения всей клеткой и способны
отвечать на них движением, называемые таксисом, перемещаясь в
направлении раздражителя или от него. (Примеры с кристалликом соли и
бактериями в капле воды: поместим рядом на стекле каплю чистой воды и
каплю воды с инфузориями. Соединим обе капли тонким водяным каналом. В
каплю с инфузориями положим маленький кристаллик соли. По мере
растворения соли туфельки будут переплывать в каплю с чистой водой: для
инфузорий раствор соли вреден. Изменим условия опыта. В каплю с
инфузориями не будем прибавлять ничего. Зато в другую каплю добавим
немного настоя с бактериями. Тогда туфельки соберутся около бактерий –
своей обычной пищи. Эти опыты показывают, что инфузории могут отвечать
определенным образом на воздействия окружающей среды, т.е. обладает
раздражимостью). Среди паразитических форм у человека встречается
инфузория – балантидий. При попадании в слизистую оболочку она вызывает
ее изъявления и кровавый понос. Эта инфузория живет в кишечной свиней,
которые служат источником заражения людей, ухаживающих за животными.
Форма тела постоянная благодаря плотной пелликуле. Органоиды движения
реснички (9). Другой важный признак – наличие двух качественно различных
ядер: крупного полиплоидного вегетативного ядра – макронуклеуса (5) и
мелкого диплоидного – генеративного ядра – микронуклеуса (6). Захват
пищи осуществляется с помощью цитостома (клеточного рта) (1) и
цитофаринкса (клеточной глотки) (2), глотка открывается непосредственно
в эндоплазму. Переваривание пищи происходит в пищеварительных вакуолях
(3, 10), а не переваренные остатки выбрасываются через порошицу (4),
содержимое сократительных вакуолей (7, 8) изливается через выделительные
поры. В эктоплазме многих инфузорий находятся особые защитные
приспособления – трихоцисты. При раздражении животного они выстреливают
длинную упругую нить, парализующую добычу. При неблагоприятных условиях
способны к инцистированию.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3b3b2a1b0202020202020202020202020202020202020202020202020202020202020202
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2d10032d2d202d202d202d202d202d202d202d202d202d202d202d202d202d202d202d20
2d202d030202203e3b3b3b3b0c0202020202020202020202020202020202020202020202
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3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a3a270202253b1016363b0c020202
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40302a322a1c021828020202020202020202020202020202020202020202020202020202
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070202020202020202020202020202020202020202020202020202020202020202020202
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022b043327202f0e11021216073616020202020202020202020202020202020202020202
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1. Две инфузории соединяются друг с другом околоротовыми областями, в
этом месте происходит разрушение пелликулы, и образуется
цитоплазматический мостик, соединяющий обе инфузории.2. Макронуклеусы
разрушаются, микронуклеусы претерпевают мейотическое деление, образуются
четыре гаплоидных ядра. Три ядра разрушаются, чВ это время в каждой
инфузории по два гаплоидных ядра, одно женское, стационарное, ядро
остается на месте, второе, мужское, мигрирует по цитоплазматическому
мостику в другую инфузорию.3. Происходит слияние мужских и женских ядер.
После этого инфузории расходятся. Конъюгация продолжается несколько
часов.4. Диплоидное ядро претерпевает три митотических деления, в
результате образуется 8 диплоидный ядер, из которых 4 становятся
микронуклеусами, 4 – макронуклеусами. Три микронуклеуса разрушаются,
образуется инфузория с 5 ядрами: 1 – микронуклеус (диплоидный), 4 –
макронуклеусы, тоже диплоидный.5. Каждый из эксконъюгантов делится,
причем происходит митотическое деление генеративного ядра, а
макронуклеусы расходятся попарно в дочерние инфузории. Образуется 4
инфузории с одним генеративным ядром и двумя макронуклеусами.6. Каждая
из этих четырех инфузорий делится, причем происходит митотическое
деление генеративного ядра, а макронуклеусы расходятся в дочерние
инфузории. Образуется 8 инфузорий, в каждой из которых один макронуклеус
и один микронуклеус. Макронуклеус становится полиплоидный, микронуклеус
остается диплоидный. Таким образом, в конъюгации принимали участие две
особи, размножение закончилось образованием восьми особей. При
длительном бесполом размножении, в результате амитотического деления
макронуклеуса, наступает “депрессия”, чтобы восстановить нормальный
набор хромосом происходит аутогамия.1. Сначала происходит разрушение
макронуклеуса, микронуклеус делится с образованием восьми гаплоидных
ядер, из которых шесть разрушаются.2. Происходит слияние двух ядер,
образуется диплоидный синкариона.3. Происходит деление синкариона, два
ядра становятся микронуклеусами, два – макронуклеусами.4. Происходит
деление инфузории, образуются две особи с нормальными микронуклеусами и
макронуклеусами.Представители: – инфузория туфелька, инфузория – трубач,
инфузория-дидиниум, сдвойки. У человека в толстом кишечнике может
паразитировать инфузория балантидий, вызывает тяжелое заболевание –
балантидиаз. Заболевание проявляется в колитах (болях в кишечнике);
кровавом стуле, лихорадочном состоянии. Основным источником заражения
являются свиньи, зараженные балантидиями. Заражение происходит на стадии
цист.
Губки.
Губки – водные сидячие многоклеточные животные. Настоящих тканей и
органов нет. Нервная система у них отсутствует. Тело в виде мешка или
бокала состоит из разнообразных клеток, выполняющих различные функции, и
межклеточного вещества.
Стенка тела губок пронизана многочисленными порами и идущими от них
каналами, сообщающимися с внутренней полости. Полости и каналы выстланы
жгутиковыми клетками. За немногими исключениями губки имеют сложный или
минеральный или органический скелет. Ископаемые остатки губок известны
уже из протерозойских пород.
Описано около 5 тыс. видов губок, большинство из них обитает в морях.
Тип делится на четыре класса: известные губки (Calcarea), кремнероговые
или обыкновенные, губки (Demospongia), стеклянные или шестилучевые,
губки (Hexactinellida или Hyalospongia) и коралловые губки
(Sclerospongia). Последний класс включает небольшое число видов, которые
обитают в гротах и туннелях среди коралловых рифов и имеют скелет,
составляющий из массивного известного основания из карбоната кальция и
кремневых одноосных игл. В качестве примера рассмотрим строение
известных губок. Тело ее мешковидное, основание оно прикреплено к
субстрату, а отверстием или устьем, обращено к верху. Парагастральная
область тела сообщается с наружной средой многочисленными каналами,
начинающимися наружными порами.
В теле взрослой губки имеются два слоя клеток – экто – и энтодерма,
между которыми залегает прослойка бесструктурного вещества – мезоглеи –
с разбросанными в ней клетками. Мезоглея занимает большую часть тела,
содержит скелет и кроме прочих – половые клетки. Наружный слой образован
плоскими эктодермальными клетками, внутренний – воротничковыми клетками
– хоаноцитами, из свободного конца которых торчит длинный жгутик. Клетки
свободно рассеянные в мезоглее, подразделяются на неподвижные
звездчатые, выполняющие опорную функцию (колленциты) скелетные подвижные
(склеробласты) занятые перевариванием пищи (амебоциты), резервные
амебоядные, которые могут превращаться в любой из названных типов, и
половые. Способности клеточных элементов переходить друг в друга готовит
об отсутствии дифференцированных тканей.
По строению скелета стенки тела и канальной системы, а также месту
расположения участков жгутиконосного слоя различают три типа губок,
наиболее простой, из которых аксон и более сложные – сикон и лейкон.
Скелет губок образуется в мезоглее. Минеральный (известковый или
кремневый) скелет состоит из отдельных или спаянных между собой игл
(спикул), формирующихся внутри клеток – склеробластов. Органический
(спонгиновый) скелет слагается из сети волокон, близких по химическому
составу к шелку и образующихся межклеточно.
Губки относятся к организмам – фильтратом. Через их тело идет
непрерывный ток воды, вызываемым действиям воротничковых клеток, жгутики
которых бьют в одном направлении – к парагастральной полости.
Воротничковые клетки захватывают из проходящей мимо них воды пищевые
частицы (бактерий, одноклеточных и т. д.) и заглатывают их, часть пищи
переваривается на месте, часть передается амебоцитам. Отфильтрованная
вода выбрасывается из парагастральной полости через устье.
Размножаются губки как бесполые (почкованием), так и половым способом.
Большинство губок – гермафродиты. Половые клетки залегают в мезоглее.
Сперматозоиды выходят в каналы, выходят через устье, проникают в другие
особи губок и оплодотворяют их яйца. Зигота дробится, вследствие чего
появляются бластула. Второй зародышевый слой (фагоцитобласт) образуется
путем иммиграции либо инвагинации. У не известковых и некоторых
известных губок бластула состоит из более или менее одинаковых
жгутиковых клеток (целобластула). В дальнейшем часть клеток, терля
жгутики, погружаются внутрь, заполняя полость бластулы, и в итоге
возникает личинка – перенхимула.
Среди бластул губок, встречаются так называемые амфибластула у которых
анимальное полушарие состоит из всяких мягких жгутиковых клеток, а
вегетативное – из крупных клеток без жгутиков, но заполненных желтком.
Амфибластула проделывают гаструляцию в теле материнской губки: клетки
вегетативного полушария впячивается внутрь бластоцель. Однако при выходе
личинки в воду энтодермальные клетки снова выворачиваются наружу
(дегаструляция) возвращаясь к состоянию амфибластулы. После этого
амфибластула оседает оборальным полюсом на дно, ее эктодермальные
жгутиковые клетки выпячиваются внутрь, а энтодермальные остаются с
наружи. Это явление называются извращением зародышевых листков. Однако
наступает и в другом случае, когда личинка – паренхимула оседает на
субстрат. Тогда ее эктодермальные клетки заползают внутрь, где и
образуют воротничково-жгутиковые камеры. Энтодерма ложится поверх
эктодермы. Устье формируется на вегетативном полюсе, который обращен
вверх.
Чаще губки живут колониями, получающимися в результате недовведенного до
конца почкования. Только немного губки являются одиночными, встречаются
также вторично-одиночные организмы. Значение из в жизни водоемов очень
велико. Фильтруя через свое тело огромное количество воды, они
способствуют ее очищению от загрязнения твердыми частицами.
Сравнительная характеристика основных классов губок.
Литература
· Биология. Т.Л.Богданова, Е.А.Солодова. 2001 год.
· Биология для поступающих в вузы. С.Г.Мамонтов. 1992 год.
· Биология. Справочник школьника. 1995 год.
· Энциклопедия для детей. Биология. 1994 год.
· Биология. Н.Грин, У.Стаут, Д.Тейлор. 1990 год.
Нашли опечатку? Выделите и нажмите CTRL+Enter
