OA?E?ANUeEEE AeA?AEAAIEE OI?AA?NEOAO

EIIA?ENOA A?EOI??ss IAAE?AIA

OAeE 577.123+577.125+612.35

ACA?IICA’ssCIE ?IOAINEAIINO? I?IE?OA?AOe?? ?

E?I?AeII-EA?IOEII?AeIIAI IAI?IO A EE?OEIAO DUNALIELLA TEOD.

OA AIEEA E?I?AeII-EA?IOEII?AeIEO AENO?AEO?A IA

OOIEOe?IIAEUeIO AEOEAI?NOUe AAIAOIOeEO?A

03.00.04 — a?io?i?y

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Oa?e?a – 1999

Aeena?oaoe??th ? ?oeiien

?iaioo aeeiiaii a iaoeiai-aeine?aeiiio ?inoeooo? a?ieia?? Oa?e?anueeiai
aea?aeaaiiai

oi?aa?neoaoo I?i?noa?noaa ina?oe Oe?a?ie

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, noa?oee iaoeiaee
ni?a?ia?oiee Aiaeeia

Aiaoie?e ?aaiiae/, caa?aeoaa/ a?aeae?eii iieaeoey?ii? a?ieia?? oa
a?ioaoiieia?? iaoeiai-

aeine?aeiiai ?inoeoooo a?ieia?? Oa?e?anueeiai aea?aeaaiiai oi?aa?neoaoo
I?i?noa?noaa

ina?oe Oe?a?ie

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, noa?oee iaoeiaee
ni?a?ia?oiee

Aiiaea?aiei Oaoyia Iao??aia, caa?aeoaa/ a?aeae?eii a?io?i?? ?
oa?iaeieia??

iae?iaoii?aeueieo nenoai ?inoeoooo i?iaeai e??ia?ieia?? ? e??iiaaeeoeeie
IAI Oe?a?ie

aeieoi? n?euenueeiainiiaea?nueeeo iaoe, noa?oee iaoeiaee ni?a?ia?oiee

?iiia ?ai? Aiaoie?eiae/, caa?aeoaa/ a?aeae?eii o?c?ieia??, a?io?i?? oa

oa?/oaaiiy ioao?a ?inoeoooo ioao?aieoeoaa OAAI, i. Ai?ee

I?ia?aeia onoaiiaa: ?inoeooo aa?iioieia?? AII Oe?a?ie (eaai?aoi??y
iieaeoey?ii?

aaiaoeee), i. Ee?a.

Caoeno a?aeaoaeaoueny » 20 » aeiaoiy 1999 ?. i 15-15 aiaeei? ia
can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee

E 64.051.17 i?e Oa?e?anueeiio aea?aeaaiiio oi?aa?neoao? I?i?noa?noaa
ina?oe Oe?a?ie ca aae?anith:

310077, i. Oa?e?a, i. Naiaiaee, 4, aoae. III-15.

C aeena?oaoe??th iiaeia iciaeiieoeny a Oeaio?aeuei?e iaoeia?e
a?ae?ioaoe? Oa?e?anueeiai

aea?aeaaiiai oi?aa?neoaoo I?i?noa?noaa ina?oe Oe?a?ie ca aae?anith:

310077, i. Oa?e?a, i. Naiaiaee, 4.

Aaoi?aoa?ao ?ic?neaiee » 17 » aa?aniy 1999 ?.

A/aiee nae?aoa? niaoe?ae?ciaaii? a/aii? ?aaee

eaiaeeaeao a?ieia?/ieo iaoe,

noa?oee iaoeiaee ni?a?ia?oiee Iaaeaeei A.?.

1

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. Ea?ioeii?aee aeyaeythoueny a ee?oeiao aaeoa??e,
aiaei?inoae, a?ea?a, aeueo ?ineei, a oaeiae oaa?ei (Liaaen-Jensen,
1977). Oi?aa?naeueiee oa?aeoa? ?iciianthaeaeaiiy oeeo niieoe aeicaiey?
i?eionoeoe, ui a ee?oeiao oaeeo ??cieo i?aai?ci?a ea?ioeii?aee
aeeiiothoue noiae? ooieoe??. Caeaoi?noue iiaeeiaoe na?oei iaaii?
aeiaaeeie oaee? coiiaeth? ooieoe?th oeeo niieoe ye i?aiaio?a, ui
iaaeathoue eie?? oeaieiai i?aai?ci?a. Aeinoaoiuei aeia?a aea/aiith ?
ooieoe?y ea?ioeii?ae?a ye oioi?aoeaioi?ieo iieaeoe o oioineioacoth/eo
i?aai?ci?a oa ooieoe?y aiiea?ioeii?ae?a a i?aaiao ci?o oaa?ei (A?eooii,
1986). Ea?ioeii?aee ? e?iio?eueieie ? caeaoi? ca’ycoaaoe iieaeoey?iee
eenaiue ? ia?ieneaei? ?aae?eaee. ?niothoue /eneaii? e?oa?aoo?i? aeai?
i?i aioeieneaeaioio ae?th ea?ioeii?ae?a (Lavy et al., 1993), iaio
a?aeiii i?i aieea ea?ioeii?ae?a ia noai a?ieia?/ieo iaia?ai (Britton,
1995). Caeeoa?oueny iaaeine?aeaeaiei ieoaiiy i?i aca?iica’ycie iai?io
ea?ioeii?ae?a c ?ioeie iaoaaie?/ieie i?ioeanaie a ee?oei? ? i?ioeanii
ee?oeiiiai oeeeeo, oi/a a?aeiii, ui i?e ieenethaaeuei?e aeaa?aaeaoe??
iieaeoe ea?ioeii?ae?a ooai?ththoueny ?a/iaeie, ui aieeaathoue ia
i?ie?oa?aoe?th, aeni?an?th aaiiio oa aeeoa?aioe?aoe?th ee?oei ??cieo
i?aai?ci?a: o?enii?ia? eeneioe — o a?ea?a, aanoeeciaa eeneioa — o
?ineei, a?oai?i A oa eiai aiaeiae — o oaa?ei (Olson, 1993). C oe???
oi/ee ci?o aeine?aeaeaiiy aca?iica’yceo ?ioaineaiino? i?ie?oa?aoe??
ee?oei c iai?iii ea?ioeii?ae?a ? e?i?ae?a ia? caaaeueiia?ieia?/ia
cia/aiiy.

Aeaye? aeaee aiaei?inoae ? a?ea?a iathoue caeaoi?noue aei iaaeneioaco ?
iaeiie/aiiy ea?ioeii?ae?a ca iaaieo oiia eoeueoeaoaaiiy ? iiaeooue aooe
aeei?enoaieie ye iiaeaeuei? ia’?eoe i?e aeine?aeaeaii? iaoaaie?cio ?
ooieoe?e ea?ioeii?ae?a.

Aiaei?ino? ?iaeo Dunaliella Teod. — i?aai?cie, ui eaaei
eoeueoeaothoueny ? iiaeooue aooe aeei?enoaieie ye aeaea?aei
ea?ioeii?ae?a oa ?ioeo a?ieia?/ii aeoeaieo ?a/iaei ia iio?aao
iaaeeoeeie, oa?/iai? ? einiaoe/ii? i?iieneiaino?. Iacaaaeath/e ia
/eneaii? aeine?aeaeaiiy, ui aoee i?enay/ai? ?ic?iaoe? oaoiieia??
i?iieneiaiai eoeueoeaoaaiiy Dunaliella, ia aaea?oueny io?eiaoe
noaa?eueiiai aeoiaeo a?iiane ? b-ea?ioeio. Oea iiaea aooe iia’ycaii c
ooieoe?iiaeueiith aaoa?iaaii?noth ee?oei a eoeueoo?? oa iaaeinoaoiueith
aeine?aeaeai?noth no?oeoo?e iiioeyoe?? ee?oei Dunaliella a i?ioean?
?inoo eoeueoo?e oa ca ??cieo oiia eoeueoeaoaaiiy (Bozhkov, Menzyanova,
1995; Aiaeeia, Iaicyiiaa, 1997). Caeeoa?oueny ia aeine?aeaeaiei
aca?iica’ycie ?ioaineaiino? i?ie?oa?aoe?? ? iai?io ea?ioeii?ae?a ?
e?i?ae?a o ee?oeiao aiaei?inoae ?iaeo Dunaliella. Ciaiiy aca?iica’yceo
i?ae oeeie iaoaaie?/ieie nenoaiaie aeicaieeoue ?ic?iaeoe nenoaio
?ioaineaiiai eoeueoeaoaaiiy aiaei?inoae c aenieei aeoiaeii
ea?ioeii?ae?a. Aeine?aeaeaiiy inoaii?o 5-oe ?ie?a iieacaee
ia?niaeoeai?noue aeei?enoaiiy ea?ioeii?ae?a, io?eiaieo c Dunaliella, ye
e?ea?nueeeo i?aia?ao?a. Iaeiae, i?iaaaeaii eeoa iiiaeeiie? ?iaioe, a
yeeo aeine?aeaeaii oa?iaeie?iaoeeo ea?ioeii?ae?a oa ?oi?
aioeieneaeaioi? aeanoeaino?, a

2

aeine?aeaeaiue ?oiueiai aieeao ia i?ie?oa?aoe?th ? aeeoa?aioe?aoe?th
ee?oei oaa?ei aeoaea iaei.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. Aeine?aeaeaiiy
i?iaiaeeeenue o a?aeae?e? iieaeoey?ii? a?ieia?? ? a?ioaoiieia?? IAe?
a?ieia?? Oa?e?anueeiai aea?aeoi?aa?neoaoo a iaaeao oaie «Aeine?aeaeaiiy
iaoai?ci?a ae?? aiaeiaaiieo ?aaoeyoi??a a?eeiai? oa e?i?aeii? i?e?iaee
ia i?ie?oa?aoe?th ee?oei oaa?eiieo ? ?ineeiieo i?aai?ci?a oa ?ieue oeeo
?aaoeyoi??a a iioiaaiaoe/i?e aaeaioaoe?? aei aeciaaiieo oaeoi??a
aeiae?eey» N Ae? 0198U005803, yeo aeeth/aii aei eii?aeeiaoe?eiiai ieaio
N16 «Iaoai?cie ?icaeoeo ? noa??iiy a?ieia?/ieo nenoai» ca oaoiaei
iai?yieii «A?ieia?y» I?i?noa?noaa ina?oe Oe?a?ie.

Iaoa ? caaea/? aeine?aeaeaiiy. Aecia/eoe aca?iica’ycie iaoaaie?cio
ea?ioeii?ae?a c e?i?aeiei iai?iii ? i?ie?oa?aoeaiith aeoeai?noth ee?oei,
a oaeiae ?ic?iaeoe ?aaeeie eoeueoeaoaaiiy ea?ioeiiiinieo
i?e?iaiaei?inoae, ui caaacia/othoue cae?eniaiiy iaaeneioaco
ea?ioeii?ae?a. Aeey aeinyaiaiiy iaoe ae??ooaaee ianooii? caaea/?:

1. Aecia/eoe ?ioaineai?noue neioaco ioeea?iiaeo eeneio o aeaio aeae?a
i?e?iaiaei?inoae — Dunaliella salina Teod. ? Dunaliella viridis Teod.,
ui ?ic??ciythoueny ca oaeaee?noth i?ie?oa?aoe??, o ia??iaee/i?e
eoeueoo?? — iiaeae? ee?oeiii? iiioeyoe?? c i?ie?oa?aoeaiith aeoeai?noth,
ui ci?ith?oueny c ieeaii /ano;

2. Aecia/eoe neeaae, ai?no o ee?oei? ? ?ioaineai?noue neioaco
ea?ioeii?ae?a ? e?i?ae?a o ia??iaee/i?e eoeueoo?? i?e?iaiaei?ino? D.
salina, ui ia? caeaoi?noue aei iaaeneioaco b-ea?ioeio, ? D. viridis, ui
ia iaeiie/o? b-ea?ioei;

3. Aecia/eoe aca?iica’ycie i?ae eiioeaio?aoe??th ee?oei a eoeueoo??,
a?eii ee?oeiii? iiioeyoe??, ina?oeaiiyi ? oaiia?aoo?ith eoeueoeaoaaiiy ?
iaeiie/aiiyi ea?ioeii?ae?a ? e?i?ae?a o ee?oeiao D. salina ? D. viridis;

4. Aecia/eoe aieea e?i?aeii-ea?ioeii?aeieo aeno?aeo?a i?e?iaiaei?inoae
?iaeo Dunaliella ia aeaye? iieacieee ooieoe?iiaeueii? aeoeaiino? ia/?iee
(?ioaineai?noue neioaco ?IE ? ai?no e?i?ae?a) i?e aaiaoeo? ? o?a?ic?
ia/?iee.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Ia i?aenoaa? iaea?aeaieo
?acoeueoao?a noi?ioeueiaaii ?iai/o a?iioaco i?i ?nioaaiiy aaciaiai oa
?iaeoeiaaiiai ??aiy ea?ioeiiaaiaco a ee?oeiao aiaei?inoae. Aaciaee
??aaiue o a?eueo?e i??? ciaoiaeeoueny i?ae aaiaoe/iei eiio?ieai, aea
ci?ithth/e oiiae eoeueoeaoaaiiy, iiaeia ?iaeoeoaaoe ea?ioeiiaaiac, ui
ciaoiaeeoueny i?ae ai?aaiaoe/iei eiio?ieai. Eiiieaeniee aieea oaeeo
/eiiee?a, ye ii/aoeiaa eiioeaio?aoe?y ee?oei, oaiia?aoo?a ? ina?oeaiiy,
i?eai?/o? i?ie?oa?aoe?th ee?oei ? ?iaeoeo? e?ii- ? ea?ioeiiaaiac, i?e
oeueiio ai?no e?i?ae?a ? ea?ioeii?ae?a ca?eueoo?oueny a 2-4 ?ace a iaio
aeae?a aiaei?inoae.

A i?ioean? eoeueoeaoaaiiy ee?oei Dunaliella ci?ith?oueny ??aaiue
neio?iiiino? i?ioiaeaeaiiy

3

ee?oeiiiai oeeeeo, ? ia 16-20-oo aeiao ?inoo niinoa??aa?oueny iaeaeua
i??a neio?ii?caoe?? ee?oeiieo oeeee?a.

Iieacaii, ui oaeaee?noue iiaeeiaiiy iai?aai?/iiai aoaeaoeth, ui aoa
aaaaeaiee a eoeueoo?aeueia na?aaeiaeua o oi?i? NaH14CO3, ee?oeiaie D.
salina ? D. viridis ciaioo?oueny c 4-i? aei 24-i? aeiae ?inoo eoeueoo?e.
Oea iaoiiaeaii ci?iaiiyi neeaaeo ? o?ceei-o?i?/ieo aeanoeainoae
eoeueoo?aeueiiai na?aaeiaeua.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. ?ic?iaeaii a?i?aaeoi?
cae?eoiai oeio aeey ianiaiai eoeueoeaoaaiiy i?e?iaiaei?inoae ?iaeo
Dunaliella (na?aeioeoai i?aa aaoi?a IA N641).

?ic?iaeaii noaio ?iaeoeoe?? iaaeneioaco ea?ioeii?ae?a a ee?oeiao
Dunaliella, ui aeicaiey? ca?eueoeoe aeo?ae b-ea?ioeio a 2-4 ?ace a
caeaaeiino? a?ae aeaeo aiaei?ino?. Iaea?aeai? ?acoeueoaoe aeicaieythoue
cai?iiiioaaoe D. viridis ii?yae c D. salina ye ia?niaeoeaiiai
i?iaeooeaioa ea?ioeii?ae?a ? iaeo?aeueieo e?i?ae?a.

Iaea?aeaii c?acee e?i?aeii-ea?ioeii?aeieo aeno?aeo?a Dunaliella, ye?
?aeiiaiaeiaaii aeey ?ic?iaee aaiaoio?iiieo e?ea?nueeeo i?aia?ao?a.

Iniaenoee aianie caeiaoaa/a. Iaienaiiy iaeyaeo e?oa?aoo?e, aenia?eiaioe
c aeine?aeaeaiiy ai?noo ? oaeaeeino? neioaco ioeea?iiaeo eeneio,
ea?ioeii?ae?a ? e?i?ae?a a i?ioean? ?inoo eoeueoo?e, aieeao
eiioeaio?aoe?? ee?oei ? a?eo eoeueoo?e ia ?ioaineai?noue i?ie?oa?aoe??
ee?oei, eiiieaeniiai aieeao eiioeaio?aoe?? ee?oei, ina?oeaiiy ?
oaiia?aoo?e ia ai?no ea?ioeii?ae?a ? e?i?ae?a a ee?oeiao D. salina ? D.
viridis aoei i?iaaaeaii naiino?eii. Aeine?aeaeaiiy aieeao c?ace?a
e?i?aeii-ea?ioeii?aeieo aeno?aeo?a c D. viridis ia ooieoe?iiaeueiee noai
ee?oei ia/?iee uo??a aoei i?iaaaeaii ?acii c iaoeiaei ea??aieeii. Aiae?c
oa ?ioa?i?aoaoe?th iaea?aeaieo ?acoeueoao?a aoei i?iaaaeaii naiino?eii.

Aea?? ia’?eoa aeine?aeaeaiiy oa oiia eoeueoeaoaaiiy i?e?iaiaei?inoae
aoei cae?eniaii i?e ni?aea??aieoeoa? aeieoi?a a?ieia?/ieo iaoe,
i?ioani?a O.A. Aeiaaaeeiie.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. ?acoeueoaoe ? iniiai? iieiaeaiiy
?iaioe aeiiia?aeaeeny ia I?aeia?iaeiiio neiiic?oi? «A?ieia?/i? iaoai?cie
noa??iiy» (Oa?e?a, 1996) ? iaoeiaeo eiioa?aioe?yo iieiaeeo o/aieo
a?ieia?/iiai oaeoeueoaoo ? iaoeiai-aeine?aeiiai ?inoeoooo a?ieia??
Oa?e?anueeiai aea?aeaaiiai oi?aa?neoaoo a 1995 ? 1996 ?ieao.

Ioaeeeaoe??. Ca ?acoeueoaoaie aeena?oaoe?eii? ?iaioe aoei iioae?eiaaii
9 i?aoeue, c ieo — 4 noaoo? ? 1 na?aeioeoai i?aa aaoi?a.

No?oeoo?a ? ianya aeena?oaoe??. Aeena?oaoe?eia ?iaioa neeaaea?oueny c
anooio ? ?icae?e?a: iaeyae e?oa?aoo?e, iaoa??aee ? iaoiaee
aeine?aeaeaiiy, ?acoeueoaoe aeanieo aeine?aeaeaiue ? ?oi? iaaiai?aiiy,
caeeth/aiiy ? aeniiaee, a oaeiae ia?ae?eo oeeoiaaii? e?oa?aoo?e c 204
iaeiaioaaiue. ?iaioo aeeeaaeaii ia 150 noi??ieao ae?oeiaaiiai oaenoo,
aiia i?noeoue 2 oaaeeoe? ? 25 ?enoie?a.

4

IAOA??AEE ? IAOIAeE AeINE?AeAEAIIss

Aeueaieia?/ii /eno? eoeueoo?e Dunaliella salina ssp. salina f. magna
IBASU-A N11 ? D. viridis var. viridis f. euchlora IBASU-A N29 (Index of
cultures…, 1991) aaee ia ??aeeiio na?aaeiaeu? A?oa?? a iiaeeo?eaoe??
Ianthe (1973) a neeyieo eii?/ieo eieaao ia 500 ie (oa? noniaic?? ia
a?eueo 1,5 ni caaoiaoee). Ia?an?a eoeueoo? cae?enithaaee eiaei? 3 oeaeii
(noaoe?iia?ia oaca ?inoo), ?icaiaey/e ?o na?aeei eoeueoo?aeueiei
na?aaeiaeuai aei eiioeaio?aoe?? ee?oei aeecueei 1,5 iei./ie. Aeey
i?iaaaeaiiy aeine?aeaeaiue eoeueoo?e ia?an?thaaee a eii?/i? eieae,
?ii?noth 100 ie ii 25 ie noniaic?? a eieao (oa? 1 ni caaoiaoee).
E?euee?noue ee?oei a iaeeieoe? ia’?io na?aaeiaeua e?/eee i?ae
i?e?ineiiii a eaia?? Ai?y?aa aai aecia/aee ca iiaeeiaiiyi noniaic?? i?e
550 ii ia niaeo?ioioiiao?? NO-46 ? ?ic?aoiaoaaee ca eae?a?iai/iith
e?eaith.

Ia?aae aianaiiyi i?/aieo iiia?aaeiee?a eiioeaio?aoe?th ee?oei
ae??aithaaee aei 12,5 iei. ee?oei a 1 ie. Iayai?noue ia?oaeo ee?oei
eiio?iethaaee ca caaa?aeaiiyi o?eiaiiaei nei?i. A 19 ie oaei? eoeueoo?e
aiineee 1,5 IAe NaH14CO3 aai 1 IAe [3I]oei?aeeia. Eoeueoo?e ?ieoaoaaee
ca ina?oeaiiy 6,5 eee a?ae 4-o eaii EA-40 ? oaiia?aoo?e 26-28 C i?ioyaii
1 aiaeeie. Iio?i eoeueoo?e ioieiaeaeoaaee ? a?aeieaaee ee?oeie a?ae
?aae?iaeoeaiiai iiia?aaeieea ioieiaeaeaiei eoeueoo?aeueiei na?aaeiaeuai.

Ee?oeie a?aeae?eyee a?ae eoeueoo?aeueii? ??aeeie oeaio?eooaoaaiiyi ca
3000 g i?ioyaii 10 oa. I?aiaioe ? e?i?aee aeno?aaoaaee ca Oie/ai (Eaeon,
1975). Aeno?aeoe ?icae?eyee iaoiaeii OOO ia ieanoeiao Silufol
(Czechoslovakia) a nenoaiao aaenai:aoeeiaee ao?? (4:1, ca ia’?iii) ?
aaenai:aaicie:aoeaoii (1:1:1, ca ia’?iii). O?aeoe?? b-ea?ioeio ?
ethoa?io ?aeaioeo?eoaaee ca aeiiiiiaith na?aee?a. I?aiaioe aeththaaee
noi?ooth oei?ioi?i:iaoaiie (2:1, ca ia’?iii) ? aecia/aee e?euee?nii ca
aeiaaeeie oaee?, ui a?aeiia?aea? iaeneioio iiaeeiaiiy i?aiaioo a
iaaeai?e noi?o? ?ic/eiiee?a: b-ea?ioei — ca 440 ii, ethoa?i — ca 460
ii, enaioio?ee a?ieaenaioeiiaiai oeeeeo — ca 420 ii. Ai?no i?aiaio?a
?ic?aoiaoaaee a iea ia ee?oeio aiaei?inoae. O?iiaoia?aie i?iyaeyee a
ia?ao eiaeo, o?aeoe?? iaeo?aeueieo e?i?ae?a ?aeaioeo?eoaaee ca
aeiiiiiaith na?aee?a ? aeththaaee noi?ooth oei?ioi?i:iaoaiie (1:1, ca
ia’?iii). E?i?aee aecia/aee e?euee?nii ye iienaii a ?iaio? (A?eaaiia,
Na?aaaa, 1975). ?ic?aooiie i?iaiaeeee a iea ia ee?oeio aiaei?inoae.

Inaae i?ney aeno?aeoe?? i?aiaio?a ? e?i?ae?a aea?/? i?iieaaee 5% HClO4
? i?iaiaeeee a?ae?ie?c ioeea?iiaeo eeneio (Aoaa?ue e ae?., 1986).
Ioeea?iia? eeneioe aecia/aee ca iiaeeiaiiyi a?ae?ie?cao?a i?e 270 ? 290
ii (Nie?ei, 1958) ? ?ic?aoiaoaaee ?oi?e ai?no a iea ia ee?oeio
aiaei?inoae. Inaae i?ney a?ae?ie?co ioeea?iiaeo eeneio ?ic/eiyee a 2 ie
1N NaOH ? aecia/aee a?eee ca iaoiaeii Eio?? (Lowry et al., 1957).

Aeey aecia/aiiy iiaeeiaiiy i?/aieo iiia?aaeiee?a ee?oeiaie ?o ?oeioaaee
a ?ic/ei? oaeiai

5

neeaaeo: 0,5% o?eoii O-100, 0,02I Tris-HCl ?I 7,0, 0,0044I AAeOA
i?ioyaii 30 oa ca e?iiaoii? oaiia?aoo?e. I?iae i?aeeenethaaee HClO4 aei
5%, caeeoaee ia 12 aiaeei ca 0 N, a iio?i oeaio?eooaoaaee ca 3000g.
Iaaeinaaeiao ??aeeio, ui i?noeea iecueeiiieaeoey?i? eiiiiiaioe ee?oei,
i?iieaaee oei?ioi?iii, a inaae, ui i?noea aenieiiieaeoey?i? eiiiiiaioe,
— noi?ooth oei?ioi?i:iaoaiie (1:2, ca ia’?iii). O o?aeoe??
iecueeiiieaeoey?ieo eiiiiiaio?a ee?oei aecia/aee e?euee?noue iaoa??aeo,
iiaeeiath/iai ca 270 ? 290 ii.

Ae?eaioe aethao?a i?aiaio?a ? e?i?ae?a, a?ae?ie?cao?a ioeea?iiaeo
eeneio ? iecueeiiieaeoey?ieo iiia?aaeiee?a aiineee a noeeioeeyoe?ei?
oeaeiie c ae?ienaiiaei noeeioeeyoi?ii (5 a ??I, 250 ia ?I?I?, 100 a
iaooae?io ia 1000 ie ae?ienaio) ? e?/eee ia noeeioeeyoe?eiiio
b-niaeo?iiao?? Aeckman LS-7800 (USA). A?ae?ie?caoe AeIE ia?aae aianaiiyi
o oeaeiie iaeo?ae?coaaee 60% EII. ?ic?aoiaoaaee ieoiio ?aae?iaeoeai?noue
(oen. ?ii. a oa ia ia ?a/iaeie). Aenia?eiaioe c aeine?aeaeaiiy aieeao
a?iiane ? i?aiaio-e?i?aeiiai aeno?aeoo D. viridis i?iaiaeeee ia
o?ei?ny/ieo naioeyo uo??a e?i?? Wistar. Aeey iiaeaethaaiiy aaiaoeoo
iaeio a?oio oaa?ei i?aeaeaaaee eieaeuei?e a?ia?oa?i?? ia/?iee ye iienaii
(Aiaeeia, Neey?, 1993). ?io?e a?oi? oaa?ei aeey iiaeaethaaiiy o?a?ico
ia/?iee aioo??oiuei/a?aaii aaiaeeee n??/aiieeneo i?aeue, yeo aiooaaee ia
o?c?ieia?/iiio ?ic/ei? aaciina?aaeiuei ia?aae aaaaeaiiyi, a aeic? 1 ia
ia 100 a iane oaa?eie 3 ?ace c ?ioa?aaeii 48 aiaeei. Oaa?eie eiaeii?
aenia?eiaioaeueii? a?oie iaea?aeoaaee aioo??oiueioeoieiai ae?oea?ei,
a?iiano aai aeno?aeo D. viridis, ?anoniaiaeiaai? a ae?oea?ei? c
?ic?aooieo 0,2 ia b-ea?ioeio ia 100 a iane oaa?eie. Oaa?eie c
aenia?eiaioaeueiei aaiaoeoii iaea?aeoaaee i?aia?aoe a?ae?aco i?ney
iia?aoe?? ? aeae? i?ioyaii 2 ae?a (3 aaaaeaiiy). Oaa?eie c o?a?icii
ia/?iee ii/eiaee iaea?aeoaaoe i?aia?aoe ianooiii? aeiae i?ney
inoaiiueiai aaaaeaiiy n??/aiieenei? i?ae? ? iaea?aeoaaee ?o i?ioyaii 5
ae?a (5 aaaaeaiue).

*a?ac aeiao i?ney inoaiiueiai aaaaeaiiy uo?ai aaiaeeee
aioo??oiuei/a?aaii ii 2 IAe [14C]i?ioiai? eeneioe ia 100 a iane oaa?eie
? /a?ac 20 oa caaeaaee ?o aeaeai?oaoe??th. I?ae /an caaeaaiiy cae?aee
e?ia ? a?aeae?eyee ne?iaaoeo. Ia/?ieo ia?oocoaaee oieiaeiei 0,9% NaCl ?
aiiiaai?coaaee a ?aaeoea? A (0,25I naoa?ica, 5iI IgCl2, 50iI ENl, 25iI
Tris-INl, ?H 7,5). Noaee?oeii? o?aeoe?? iaea?aeoaaee aeeoa?aioe?aeueiei
oeaio?eooaoaaiiyi aiiiaaiaoo. Iinoi?e?iniiaeueio o?aeoe?th
aeei?enoiaoaaee ye o?aeoe?th oeeoicieth. A inaaeao yaea? ? ae?eaioao
noaee?oeiieo o?aeoe?e aecia/aee ieoiio ?aae?iaeoeai?noue ?IE, a
ae?eaioao oeeoicieth ? i?e?inii ee?oei ia/?iee ? ne?iaaoee e?ia?
aecia/aee ai?no e?i?ae?a ? a?eea, aeei?enoiaoth/e o? ae nai? iaoiaee, ui
? aeey ee?oei aiaei?inoae. Ai?no e?i?ae?a ?ic?aoiaoaaee a iea ia 1 ia
a?eea.

Iaea?aeai? ?acoeueoaoe ia?iaethaaee noaoenoe/ii, aeei?enoiaoth/e
iaia?aiao?e/i? iaoiaee aiae?co (Aoaea?, 1978).

6

INIIAI? ?ACOEUeOAOE AeINE?AeAEAIIss

?ioaineai?noue neioaco ? ai?no ioeea?iiaeo eeneio o aiaei?inoae ?iaeo
Dunaliella Teod. a i?ioean? ia??iaee/iiai eoeueoeaoaaiiy. Noia?a/eea?
a?aeiiino?, ui ? a e?oa?aoo??, i?i ca’ycie i?ie?oa?aoe?? ee?oei
Dunaliella c e?i?aeii-ea?ioeii?aeiei iai?iii aoei iaea?aeaii ia
i?aenoaa? aeaieo uiaei aeeiai?ee ?inoo aac o?aooaaiiy aaoa?iaaiiino?
ee?oeiieo eoeueoo? ca ooieoe?iiaeueiei noaiii ee?oei. Ia?oa i?ae
aeei?enoiaoaaoe ia??iaee/io eoeueoo?o Dunaliella aeey ??oaiiy caaea/, ui
iinoaaeaii a ?iaio?, iaiao?aeii ioe?ieoe ooieoe?iiaeueio aaoa?iaaii?noue
ee?oei o i?e. *ooeeaei oa ?ioi?iaoeaiei iieacieeii ooieoe?iiaeueii?
aaoa?iaaiiino? ? i?ie?oa?aoeaii? aeoeaiino? ee?oei iiaea aooe
oaeaee?noue neioaco ioeea?iiaeo eeneio, ine?eueee ?oi?e neioac ?
a?aeianaiei aei noaae?e ee?oeiiiai oeeeeo, a?aeia?aaea? ni?yiiaai?noue
caaaeueiiai iaoaaie?cio ee?oei ? ia? ae?aaeaio /aniao oa?aeoa?enoeeo.

?ioaineai?noue i?/aiiy AeIE ?aae?iaeoeaiei iiia?aaeieeii NaH14CO3 a
i?ioean? eoeueoeaoaaiiy D. salina ? D. viridis aoea iaea?eueoith a 4-o
aeiaiai? eoeueoo?e, ??cei ciaiooaaeany ia 8-12-oo aeiao ?inoo eoeueoo?,
c iacia/iei ca?eueoaiiyi oeueiai iieacieea ia 16-20-oo aeiao, ? ciia
ciaiooaaeany a ee?oeiao 24-o aeiaiaeo eoeueoo?. Aecia/aiiy ieoiii?
?aae?iaeoeaiino? ?IE a eoeueoo?ao ??ciiai a?eo iieacaei, ui oaeaee?noue
?? neioaco ciaiooaaeany c 4-i? aei 8-12-i? aeiae ?inoo a 2 ?ace.
Iaeiae, iaaeae? oeae iieaciee iaeaea e?i?eii ciaiooaaany aei 24-i?
aeiae eoeueoeaoaaiiy (?en. 1). ?acoeueoaoe ioe?iee oaeaeeino? neioaco
AeIE a i?ioean? eoeueoeaoaaiiy aiaei?inoae ?, cie?aia, ??cea ??
ciaioaiiy ia 8-12-oo aeiao ?inoo, aeyaeeeny ianiiae?aaieie, ine?eueee
aiie ia ca?aaeeny c aeinoaoiuei aaeeeith oaeaee?noth ?inoo eoeueoo?e a
oeae ia??iae. Oea i?ioe??//y aoei iia’ycaii c oei, ui a?ae?iea?aiiao
iao??th c ??ciith oaeaee?noth aeeth/aany a ee?oeie eoeueoo? ??ciiai a?eo
(?en. 1).
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f8000000ffffffffffffffffffffffffffffffffffffffffffc07ffffffffffffff01fff
f8ffffffffffffffff8f1fc7ffffffffffffff01ffffffffffffffc07fffe3ffffffffff
fffffe3c7f1ffffffffffffff8000000ffffffffffffffffffffffffffffffffffffffff
ffc07ffffffffffffff01ffff8ffffffffffffffff8f1fc7ffffffffffffff01ffffffff
ffffffc07fffe3fffffffffffffffe3c7f1ffffffffffffff8000000ffffffffffffffff
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f8000000fffffffffffffffffffffffffffffffffffffffffff07ffffffffffffffc1fe3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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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f8000000ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
f8000000fffffffffffffffffffffffff803ffffffffffffffffffffffffffffffffffff
ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
fffffffffffffffffffffffff8000000ffffffffffffffffffffffffe3f8ffffffffffff
ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
fffffffffffffffffffffffffffffffffffffffffffffffff8000000ffffffffffffffff
ffffffffe3f8ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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f8000000ffffffffffffffffffffffffe3f8ffffffffffffffffffffffffffffffffffff
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fffffcfffffffff9fffffffff8000000ffffffffffffffffffffffffe3f8ffffffffffff
fff9fffffffff3fffffffff3fffffffff3fffffffff3fffffffcfffbfdffff3fffffffff
3fffffffff9fffffffff9ffffffffcfffffffff9fffffffff8000000ffffffffffffffff
ffffffffe3f8fffffffffffffff9fffffffff3fffffffff3fffffffff3fffffffff3ffff
fffcfffbfdffff3fffffffff3fffffffff9fffffffff9ffffffffcfffffffff9ffffffff
f8000000ffffffffffffffffffffffffe3f8fffffffffffffff9fffffffff3fffffffff3
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fff9fffffffff3fffffffff3fffffffff3fffffffff3fffffffcfffbfc00000000000000
000000000000000000000000000000000000000000000000f8000000ffffffffffffffff
ffffffffe3f8fffffffffffffff9fffffffff3fffffffff3fffffffff3fffffffff3ffff
fffcfffbfdffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
f8000000ffffffffffffffffffffffffe3f8fffff8000000000000000000000000000000
00000000000000000000000000000003fdffffffffffffffffffffffffffffffffffffff
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f8000000ffffffffffffffffffffffffe3f8fffffbffffffffffffffffffffffffffffff
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fffffffffffffffffffffffffffff07ffffffffbfffffffff8000000ffffffffffffffff
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ffc07ffbfdfffffffffffffffffffffffffffffffffffffffffffdfffffffffbffffffff
f8000000fffffffffffffffffffffffff803fffffbffffffffffffffffffffffffffffff
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fffffdfffffffffbff8003fff8000000fffffffffffffffffffffffff803fffffbffffff
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fffffffffffffffffffffffffffffdffffffffe0ff8003fff8000000ffffffffffffffff
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fffbfffffffffffffffffffffffffffffffffffffffc1ffffffffdfffffff001ff8fe3ff
f8000000fffffffffffffffffffffffffffffffffbffffffffffffffffffffffffffffff
ffffffffffffffffffc0fffffe0bffffffffffffffffffffffffffffffffffffffff7fff
fffffdfffeff007bff8fe3fff8000000fffffffffffffffffffffffffffffffffbffffff
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fffbfffffffffffffffffffffffffffff01fffffffff7fffffff000007fffffbffe3ffff
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ffc0001ffffffffbfff8fffff8000000fffffffffffffffffffffffffffffffffbffffff
fffffffffffffffffffffffffffffffffbfffffff807fffffffbffffffffffffffffffe0
3ffffffffeffffffffe07c003fc7fdfffffffffbfff8fffff8000000ffffffffffffffff
fffffffffffffffffbfffffffffffffffffffffffffffffffffffffffbffffff87f7ffff
fffbfffffffffffffffffffdfffffffffeffffffbfc0007ffffffdfffffffffbfffe3fff
f8000000fffffffffffffffffffffffffffffffffbffffffffffffffffffffffffffffc0
7ffffffffbffff1fff80fffffffbfffffffffffffffffffdfffffffffeffff803fff7fff
fffffdffffffffc07ffe3ffff8000000fffffffffffffffffffffffffffffffffbffffff
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fffffffffef87f8fffff7ffffffffdffffffffffffff8ffff8000000ffffffffffffffff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ffffffc7ffffffffffffffddffffffffffffffffffffffffffffffffffffffffffffffff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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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f8000000ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
fffffffffffffffffffffffffffffffffffffffffffffffff80000000400000007010100
030000000000

?en. 1. Ieoiia ?aae?iaeoeai?noue AeIE (1), ?IE (2) ?
aioo??oiueiee?oeiiiai ioea iiia?aaeiee?a ioeea?iiaeo eeneio (3) a
ee?oeiao D. salina (a) ? D. viridis (a) /a?ac 60 oa i?ney aianaiiy
NaH14CO3 aei eoeueoo? aac iiia?aaeiueiai ci?iaiiy eoeueoo?aeueiiai
na?aaeiaeua

Ca?aeii aeaieo e?oa?aoo?e ?ic/eiaiee CO2 ? ia?aaaaeiith oi?iith
iai?aai?/iiai aoaeaoeth, yeo iiaeeiathoue ee?oeie Dunaliella
(Ianthe, 1965). ??aiiaaaa i?ae H14CO3 ? i?iaeoeoii eiai

7

a?ae?ie?co 14CO2, yea anoaiiaeth?oueny a eoeueoo?aeuei?e ??aeei? i?ney
aianaiiy NaH14CO3, caeaaeeoue a?ae

pH eoeueoo?aeueii? ??aeeie. Iiaeia /aeaoe, ui aenia?eiaioaeueia ci?iaiiy
eoeueoo?aeueiiai na?aaeiaeua aieeia ia aioo??oiueiee?oeiiee ai?no
H14CO3. Aeaeaeaiiy eoeueoo?aeueiiai na?aaeiaeua ? ia?aianaiiy ee?oei 4,
8, 12, 16, 20, 24-oe aeiaiaeo eoeueoo? ia na?aea na?aaeiaeua i?eaiaeeei
aei i?aeoe/ii iaeiaeiaiai aeeth/aiiy ?ii?a a?ae?iea?aiiaoo a ee?oeie. A
?ac? iaeiaeiaiai ??aiy ioea i?/aieo iiia?aaeiee?a neioac AeIE
cae?enithaaany c iaea?eueoith oaeaee?noth a ee?oeiao 16-oe aeiaiaeo
eoeueoo? D. salina ? D. viridis. Iaaeae?, ?c ca?eueoaiiyi no?ie?a
eoeueoeaoaaiiy, ?ioaineai?noue neioaco AeIE a ee?oeiao D. salina ? D.
viridis aeae?eueea ciaiooaaeany, iaeiae caeeoaeany cia/ii aeuith i?ae a
4-8-ie aeiaiaeo eoeueoo? (?en. 2). Oaeaee?noue neioaco ?IE a i?ioean?
eoeueoeaoaaiiy ca iaci?iiiai ??aiy ioea i?/aieo iiia?aaeiee?a
ni?aiaaeaea ca oa?aeoa?ii c oaeith AeIE (?en. 2). I?e oeueiio
oaeaee?noue neioaco ?IE aoea ye ? a ia?oiio ?ac? cia/ii (a 10 ?ac?a)
iaioa ii??aiyii c oaeaee?noth neioaco AeIE.

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ff8fe38fe38fe3ffffffffffffffffffffffffffffffffffffffffffffffffffffffffff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?en. 2. Ieoiia ?aae?iaeoeai?noue AeIE (1), ?IE (2) ?
aioo??oiueiee?oeiiiai ioea iiia?aaeiee?a ioeea?iiaeo eeneio (3) a
ee?oeiao D. salina (a) ? D. viridis (a) /a?ac 60 oa i?ney aianaiiy
NaH14CO3 aei eoeueoo? ??ciiai a?eo, ui aoee ia?aaaaeaieie ia na?aea
na?aaeiaeua A?oa??

Iaea?aeai? ?acoeueoaoe na?ae/aoue i?i oa, ui a i?ioean? ia??iaee/iiai
eoeueoeaoaaiiy ee?oeie ?ic??ciyeeny ca ?oiueith ooieoe?iiaeueiith
aeoeai?noth ? 16-oe — 20-oe aeiaiaa eoeueoo?a D. salina ? D. viridis ?
iaea?eueo iaeii??aeiith ca i?ie?oa?aoeaiith ? ooieoe?iiaeueiith
aeoeai?noth ee?oei. Oaeei /eiii, a i?ioean? ia??iaee/iiai eoeueoeaoaaiiy
?niothoue aoaie a?aeiinii? neio?ii?caoe?? ooieoe?iiaeueiiai noaio
ee?oei, oiaoi iaeii/aniiai neioaco AeIE ? ?IE a iniiai?e ian? ee?oei.
Iaiao?aeii a?aei?oeoe, ui c a?eii eoeueoo?e aiaei?inoae ?iaeo Dunaliella
??cei iaaea? oaeaee?noue aeeth/aiiy I14NI3 a ee?oeie ? a cia/i?e i???
oea iaoiiaeaii aieeaii eoeueoo?aeueiiai na?aaeiaeua ia oe? i?ioeane.

E?i?aeii-ea?ioeii?aeiee iai?i a ee?oeiao i?e?iaiaei?inoae ?iaeo
Dunaliella Teod. a i?ioean? ia??iaee/iiai eoeueoeaoaaiiy. A ianooii?e
na??? aenia?eiaio?a aeine?aeaeoaaee aeeiai?eo

8

ai?noo ea?ioeii?aeieo i?aiaio?a a ee?oeiao D. salina ? D. viridis a
i?ioean? ia??iaee/iiai eoeueoeaoaaiiy. A iaoeo aenia?eiaioao
ea?ioeii?aee, aeno?aaiaai? c ee?oei D. salina ? D. viridis,
?iciiae?eyeeny ia 6 o?aeoe?e iaoiaeii oiieioa?iai? o?iiaoia?ao?? a
nenoai? aaenai:aaicie:aoeaoii (1:1:1). E?euee?niee aiae?c iaea?aeaieo
o?aeoe?e iieacaa, ui ia ethoa?i i?eiaaea? a?eueo iieiaeie a?ae on?o
ea?ioeii?ae?a (69 ? 57 % o D. salina ? D. viridis, a?aeiia?aeii), a ia
noia?io o?aeoe?th enaioio?e?a a?ieaenaioeiiaiai oeeeeo — a?ey 20 % (21 ?
19 % a?aeiia?aeii o D. salina ? D. viridis). Ai?no b-ea?ioeio o D.
salina ci?ithaaany a?eueo i?ae a 3 ?ace c 4-i? aei 24-i? aeiae ?inoo.
Ai?no b-ea?ioeio a i?ioean? ?inoo eoeueoo?e D. viridis caeeoaany
iaci?iiei ? aoa a 2 ?ace iaioa ii??aiyii c D. salina c 4-i? aei 16-i?
aeiae ?inoo (?en. 3).

picscalex334010009000003d04500000000ae4500000000040000000301080005000000
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fffffffffffffffffffffffffffffffffffffffffffff800040000000701010003000000
0000

?en. 3. Ai?no b-ea?ioeio (1), ethoa?io (2) ? enaioio?e?a
a?ieaenaioeiiaiai oeeeeo (3) a i?ioean? ia??iaee/iiai eoeueoeaoaaiiy
ee?oei D. salina (a) ? D. viridis (a)

Ii??aiyiiy e?eaeo aeeiai?ee neioaco AeIE (?en. 2) ? ai?noo b-ea?ioeio
(?en. 3) a i?ioean? ?inoo ia??iaee/ii? eoeueoo?e iieacaei, ui o D.
salina ia ii/aoeiaeo noaae?yo eoeueoeaoaaiiy (4-oa-8-ia aeiaa ?inoo)
niinoa??aa?oueny i?yia ei?aeyoe?y i?ae oaeaee?noth neioaco AeIE ?
ai?noii ea?ioeii?ae?a. Iaaeae? (12-oa-16-oa aeiaa) oa?aeoa? caeaaeiino?
i?ae oaeaee?noth neioaco AeIE ? iaoaaie?ciii ea?ioeii?ae?a ci?ithaaany
ia i?ioeeaaeiee — ciaioaiiy ?ioaineaiino? i?ie?oa?aoe??
noi?iaiaeaeo?oueny iaeiie/aiiyi b-ea?ioeio, a ca?eueoaiiy ?ioaineaiino?
i?ie?oa?aoe?? — ciaioaiiyi ai?noo b-ea?ioeio. O D. viridis ia oe?
ci?iaiiy ?ioaineaiino? i?ie?oa?aoe?? a i?ioean? eoeueoeaoaaiiy ai?no
b-ea?ioeio caeeoa?oueny iino?eiei.

Iiaeia i?eionoeoe, ui iae?i?eia aeeiai?ea ai?noo b-ea?ioeio a ee?oeiao
D. salina iia’ycaia c aeei?enoaiiyi ioea iiia?aaeiee?a ea?ioeii?ae?a,
cie?aia aoeaoeeKoA, ia neioac ?ioeo i?iaeoeo?a, iai?eeeaae iaeo?aeueieo
e?i?ae?a, aai c iaoaaie?/ieie ia?aoai?aiiyie naiiai b-ea?ioeio,
iai?eeeaae, a ?io? ea?ioeii?aee. Ai?no ethoa?io ? noia?ii? o?aeoe??
enaioio?e?a ci?ithaaany iae?i?eii a i?ioean? ?inoo eoeueoo? iaio aeae?a
aiaei?inoae. Ni?aa?aeiioaiiy o?aeoe?e ea?ioeii?ae?a oaeiae aa??thaaei

9

(?en. 3). Aoei aeyaeaii, ui a ee?oeiao D. salina ? D. viridis aei 55%
a?ae noie iaeo?aeueieo e?i?ae?a (o?eaoeeeae?oea?eaee, noa?i?aee ?
iaaoa?eo?eiaai? aee?i? eeneioe) i?eiaaea? ia iaaoa?eo?eiaai? aee?i?
eeneioe. A ee?oeiao D. salina i?noeeiny a 1,3-1,5 ?ace a?eueo
iaeo?aeueieo e?i?ae?a ii??aiyii c D. viridis. A i?ioean? eoeueoeaoaaiiy
ee?oei aiaei?inoae ai?no aeine?aeaeoaaieo o?aeoe?e e?i?ae?a caeeoaany
iaci?iiei (?en. 4). Oaeei /eiii, ai?no ea?ioeii?ae?a a i?ioean? ?inoo
eoeueoo?e aiaei?inoae ia ei?aethaaa ?c ai?noii iaeo?aeueieo e?i?ae?a a
?oi?o ee?oeiao. Iiaeia aaaaeaoe, ui aeyaeaiee iae?i?eiee oa?aeoa?
ci?iaiiy ai?noo ea?ioeii?ae?a a i?ioean? eoeueoeaoaaiiy ? iia’ycaiei c?
ci?iaiiyi ?ioaineaiino? ?oiueiai neioaco ? ia caeaaeeoue a?ae neioaco
e?i?ae?a.

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ffffffffffffffffff800000ffffffffffffe3ffff8fe3ffffffffffffffffffffffffff
ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ff800000ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffff800000ffffffffffffffffffffffffffffffffffffffffffffffff
ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
ffffffffffffffffffffffffff800000ffffffffffffffffffffffffffffffffffffffff
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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffff80000004000000
07010100030000000000

?en. 4. Ai?no iaaoa?eo?eiaaieo aee?ieo eeneio (1), o?eaoeeeae?oea?eae?a
(2) ? noa?i?ae?a (3) a ee?oeiao D. salina (a) ? D. viridis (a) a
i?ioean? ia??iaee/iiai eoeueoeaoaaiiy

A ianooii?e na??? aenia?eiaio?a aecia/aee oaeaee?noue neioaco o?aeoe?e
ea?ioeii?ae?a, i?i yeo noaeeee ca aeeth/aiiyi ?aae?iaeoeaiiai aoaeaoeth
aei neeaaeo ea?ioeii?ae?a i?e aiaeeii?e aeniiceoe?? c NaH14CO3. C
iaea?eueoith oaeaee?noth ?aae?iaeoeaia i?oea aeeth/aeany aei b-ea?ioeio.
Oae, eiai neioac a 4-o aeiaia?e eoeueoo?? D. salina cae?enithaaany a 6,3
?ace oaeae/a ii??aiyii c ethoa?iii ? a 3,8 ?ace oaeae/a ii??aiyii c
?aooith o?aeoe?e ea?ioeii?ae?a. Aeey D. viridis oey ??cieoey aoea
aeae?eueea iaioith — a 5,0 ? 2,0 ?ace a?aeiia?aeii. E?i?eia ciaioaiiy
ieoiii? ?aae?iaeoeaiino? a i?ioean? eoeueoeaoaaiiy iia’ycaii c?
ciaioaiiyi ioea i?/aieo iiia?aaeiee?a (H14CO3) a ee?oei? a i?ioean?
eoeueoeaoaaiiy. Aeeiai?ea ieoiii? ?aae?iaeoeaiino? ??cieo o?aeoe?e
ea?ioeii?ae?a aoea ??ciith ia iaeiaeiaiio oe? ciaioaiiy
aioo??oiueiee?oeiiiai ioea i?/aieo iiia?aaeiee?a ea?ioeii?ae?a a
i?ioean? ?inoo eoeueoo?e. Oae, ieoiia ?aae?iaeoeai?noue b-ea?ioeio
iaeaea e?i?eii ciaiooaaeany c 4-o aei 24-i? aeiae ?inoo. Ieoiia
?aae?iaeoeai?noue ethoa?io ? noia?ii? o?aeoe?? enaioio?e?a ciaiooaaeany
aei 12-14-i? aeiae ? iaaeae? ia ci?ithaaeany aai ae ca?eueooaaeany, ui
na?ae/eoue i?i ci?iaiiy oaeaeeino? neioaco oeeo o?aeoe?e ea?ioeii?ae?a a
i?ioean? eoeueoeaoaaiiy (?en. 5).

Oaeei /eiii, b-ea?ioei ? iaeaeoeai?oei iai?ithaaiei ea?ioeii?aeiei
i?aiaioii a ee?oei? Dunaliella. Aeey eiaeii? c aeine?aeaeoaaieo o?aeoe?e
ea?ioeii?ae?a aoa aeyaeaiee ?iaeea?aeoaeueiee oa?aeoa? ci?iaiiy
?oiueiai ai?noo ? oaeaeeino? neioaco a i?ioean? eoeueoeaoaaiiy, ?
a?i

10

?ic??ciyany o D. salina ? D. viridis. Ea?ioeii?aeiee iai?i ?
aa??aaaeueiei iieacieeii, ui ci?ith?oueny a i?ioean? eoeueoeaoaaiiy
aiaei?inoae ? eiai ca’ycie c ?ioaineai?noth i?ie?oa?aoe?? ee?oei a
eoeueoo?? ia ? iaeiicia/iei. Oea aeaco? ia oa, ui a?i caeaaeeoue a?ae
??ciiiai?oieo /eiiee?a, ui ci?ithaaeeny a i?ioean? ?inoo eoeueoo?e,
oiaoi eiio?ieth?oueny ia o?eueee aaiaoe/ii, aea ? ai?aaiaoe/ii.

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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
ffffffffffffffffffffffffffff80000400000007010100030000000000

?en. 5. Ieoiia ?aae?iaeoeai?noue b-ea?ioeio (1), ethoa?io (2) ?
enaioio?e?a a?ieaenaioeiiaiai oeeeeo (3) a i?ioean? ia??iaee/iiai
eoeueoeaoaaiiy ee?oei D. salina (a) ? D. viridis (a)

Aieea eiioeaio?aoe?? ee?oei ? a?eo eoeueoo?e ia aeeiai?eo ?inoo ?
e?i?aeii-ea?ioeii?aeiee iai?i a ee?oeiao Dunaliella Teod. A i?ioean?
?inoo eoeueoo?e a ia??iaee/iiio ?aaeei? aacia?a?aii ci?iththoueny oae?
ia?aiao?e ye eiioeaio?aoe?y ee?oei, ooieoe?iiaeueiee noai ee?oeiii?
iiioeyoe??, neeaae eoeueoo?aeueii? ??aeeie. Ci?iaiiy eiiieaeno oeeo
ia?aiao??a a naith /a?ao aoaea aieeaaoe ia iaoaaie?ci ee?oeie, ?
cie?aia, ia e?ii- ? ea?ioeiiaaiac.

Oaeaee?noue iiae?eo ee?oei Dunaliella caeaaeaea a?ae ii/aoeiai?
eiioeaio?aoe?? ee?oei a eoeueoo??. Iaea?eueoee i?e??no /enea ee?oei
niinoa??aaany ca ii/aoeiai? eiioeaio?aoe?? 2,5 iei. ee?oei a 1 ie
eoeueoo?aeueiiai na?aaeiaeua. Iiaeaeueoa ca?eueoaiiy ii/aoeiai?
eiioeaio?aoe?? ee?oei i?ecaiaeeei aei i?eai?/aiiy ?inoo eoeueoo?e. Aeae?
aecia/aee aieea eiioeaio?aoe?? ee?oei a eoeueoo?? ia ai?no b-ea?ioeio.
Aeey oeueiai ee?oeie c 4-o, 8-ie, 12-oe, 16-oe, 20-oe ? 24-o aeiaiaeo
eoeueoo? ia?an?thaaee ia na?aea eoeueoo?aeueia na?aaeiaeua oaeei /eiii,
uia ii/aoeiaa eiioeaio?aoe?y noaiiaeea aeecueei 5, 15 ? 20 iei./ie ?
/a?ac 4 aeiae ?inoo ca ina?oeaiiy 6,5 eee ? oaiia?aoo?e 27-30 C
aecia/aee ai?no b-ea?ioeio a ee?oeiao. Ca oaeeo aenieeo eiioeaio?aoe?e
ee?oei a na?aaeiaeu? ai?no b-ea?ioeio ca?eueooaaany iaeaea a 2 ?ace ye a
ee?oeiao D. salina, oae ? D. viridis ii??aiyii c eiioeaio?aoe??th ee?oei
1,3 iei./ie (?en. 6). Aaaeeeaei ? oa, ui ca oaeeo eiioeaio?aoe?e ee?oei
a eoeueoo?? ia niinoa??aaeiny i?e?inoo e?eueeino? ee?oei ? i?aeaeaeiaeo
??cieoeue, a oaeiae ??cieoeue, iia’ycaiieo c a?eii eoeueoo?e, ye? aoee
oa?aeoa?ieie aeey noaiaea?oieo oiia eoeueoeaoaaiiy (?en. 3). Oea
na?ae/eoue i?i

11

aenieo iaoaaie?/io aa??aaaeuei?noue ea?ioeiiaaiaco a ee?oeiao
aiaei?inoae ? iniaeeai — D. viridis. Iaeii/ania ca?eueoaiiy ina?oeaiiy
aei 8 eee ? oaiia?aoo?e aei 38-40 C ca aenieeo eiioeaio?aoe?e ee?oei a
eoeueoo?? ia i?ecaiaeeei aei caaeaae? eoeueoo?e ? noi?iaiaeaeoaaeiny
i?aeaeuaiiyi ai?noo b-ea?ioeio a ee?oeiao D. salina ? D. viridis, i?e
oeueiio caa??aaany ca’ycie oeueiai iieacieea c eiioeaio?aoe??th ee?oei
i?e ia?an?a? (?en. 6).

?en. 6. Ai?no b-ea?ioeio ia 4-oo aeiao ?inoo ee?oei D. salina ? D.
viridis ca ina?oeaiiy 6,5 eee ? oaiia?aoo?e 25-27 N (), ? ina?oeaiiy
8,0 eee ? oaiia?aoo?e 38-40 N () ca ii/aoeiai? eiioeaio?aoe?? ee?oei 4-6
iei./ie (1), 10-14 iei./ie (2) ? 18-20 iei./ie (3)

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Oaeei /eiii, ca?eueooth/e eiioeaio?aoe?th ee?oei, ina?oeaiiy aei 8 eee
? oaiia?aoo?o eoeueoeaoaaiiy aei 38-40 C aaeaeiny cia/ii ca?eueoeoe
ai?no b-ea?ioeio a ee?oeiao D. salina ? D. viridis, i?e oeueiio
i?aeaeaeiaa ??cieoey e?ea?aeoaaeany.

Ua a?eueo aa??aaaeueiei aeyaeany ai?no iaeo?aeueieo e?i?ae?a a ee?oeiao
aiaei?inoae ca icia/aieo oiia eoeueoeaoaaiiy. Ci?iaiiy eiioeaio?aoe??
ee?oei ia aieeaaei ia ai?no iaeo?aeueieo e?i?ae?a a ee?oeiao D. salina ?
D. viridis. Iaeiae, ca?eueoaiiy ina?oeaiiy ? oaiia?aoo?e eoeueoeaoaaiiy
noi?iaiaeaeoaaeiny cia/iei ca?eueoaiiyi ai?noo noa?i?ae?a — a 2,0-3,0
?ace, iaaoa?eo?eiaaieo aee?ieo eeneio — a 2-4 ?ace, o?eaoeeeae?oea?eae?a
? ao???a noa?i?ae?a — oaeiae a 2-3 ?ace. Iaiao?aeii a?aei?oeoe, ui ca
ca?eueoaiiy ina?oeaiiy ? oaiia?aoo?e eoeueoeaoaaiiy iaea?eueoee ai?no
e?i?ae?a a ee?oeiao aeyaeyany o aeiaaeeo aaeeeeo eiioeaio?aoe?e ee?oei
(?en. 7). Oaeei /eiii, eiiieaenia ca?eueoaiiy eiioeaio?aoe?? ee?oei,
ina?oeaiiy ? oaiia?aoo?e i?ae /an eoeueoeaoaaiiy noi?iaiaeaeoaaeiny
iaea?eueoei i?aeaeuaiiyi ai?noo iaeo?aeueieo e?i?ae?a.

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ffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffffff
fffffffffffffffffffff8000400000007010100030000000000

?en. 7. Ai?no iaaoa?eo?eiaaieo aee?ieo eeneio (I), o?eaoeeeae?oea?eae?a
(II), noa?i?ae?a (III) ? ao???a noa?i?ae?a (IV) ia 4-oo aeiao ?inoo D.
salina (a) ? D. viridis (a) ca ina?oeaiiy 6,5 eee ? oaiia?aoo?e 25-27 N
(), ? ina?oeaiiy 8,0 eee ? oaiia?aoo?e 38-40 N () ca ii/aoeiai?
eiioeaio?aoe?? ee?oei 4-6 iei./ie (1), 10-14 iei./ie (2) ? 18-20 iei./ie
(3)

12

Iaiao?aeii a?aei?oeoe, ui D. salina ? D. viridis ca noaiaea?oieo oiia
eoeueoeaoaaiiy ?ic??ciythoueny i?ae niaith ca ai?noii ea?ioeii?ae?a ?
e?i?ae?a a 1,5-2 ?ace. Eiiieaeniee aieea oaeeo /eiiee?a ye ii/aoeiaa
eiioeaio?aoe?y ee?oei, oaiia?aoo?a ? ina?oeaiiy ?iaeoeothoue e?ii- ?
ea?ioeiiaaiac, i?e oeueiio ai?no e?i?ae?a ? ea?ioeii?ae?a ca?eueoo?oueny
a 2-4 ?ace a iaio aeae?a aiaei?inoae.

Ia i?aenoaa? iaea?aeaieo aeaieo iiaeia noi?ioethaaoe ?iai/o a?iioaco
i?i ?nioaaiiy aaciaiai ? ?iaeoeiaaiiai ??aiy ea?ioeiiaaiaco a ee?oeiao
aiaei?inoae. Aaciaee ??aaiue niinoa??aa?oueny a iioeiaeueieo oiiaao
eoeueoeaoaaiiy. Aeey aaciaiai ??aiy oa?aeoa?iith ? i?yia ei?aeyoe?y
?ioaineaiino? i?ie?oa?aoe?? c ai?noii ea?ioeii?ae?a o ee?oei? ?
a?aenooi?noue ca’yceo c iaeiie/aiiyi iaeo?aeueieo e?i?ae?a. Aeey
?iaeoeiaaiiai ??aiy oa?aeoa?iith ? cai?ioia ei?aeyoe?y c ?ioaineai?noth
i?ie?oa?aoe?? ? i?yia — c iaeiie/aiiyi iaeo?aeueieo e?i?ae?a.
Iaiao?aeii a?aei?oeoe, ui iaeaeaa aeine?aeaeoaaieo aeaee Dunaliella ca
aecia/aieo oiia eoeueoeaoaaiiy iiaeooue aooe ia?niaeoeaieie
i?iaeooeaioaie ea?ioeii?ae?a ? e?i?ae?a.

Aieea a?iiane ? oei?io?e-ea?ioeii?aeiiai aeno?aeoo Dunaliella viridis
Teod. ia neioac ?IE a ee?oeiao ia/?iee uo??a c aenia?eiaioaeueiei
aaiaoeoii. AEe?i?ic/eiiee eiiieaen (oei?io?e-ea?ioeii?aeia ianoa) c oai?
ninie, yeeie ? i?ooe aeaaii aeei?enoiao?oueny a oa?ai?? ?ai ?
caiaethaaeueieo oai?ia (O?ieia e ae?., 1969). Oei?io?e-ea?ioeii?aeia
ianoa c neiueicaeaieo aiaei?inoae i?enei?thaaea caai?iiy
aenia?eiaioaeueieo oa?i?/ieo ii?e?a o e?ie?a (Naeaae/, 1985). A?iiana
Dunaliella ? iaoiene/iith aeey oaa?ei iaa?oue i?e o?eaaeiio aaeeaaii?
per os (Mokady et al., 1989), ? iiaea aeyaeoeny aoaeoeaiith a oa?ai??
caoai?thaaiue aioo??oieo i?aai?a. A ianooii?e na??? aenia?eiaio?a
aeine?aeaeoaaee aieea a?iiane ? oei?io?e-ea?ioeii?aeiiai aeno?aeoo D.
viridis ia oaeaee?noue neioaco ? o?ainii?oo ?IE a ee?oeiao ia/?iee uo??a
ia aeaio iiaeaeyo o?aaeaiiy ia/?iee — aenia?eiaioaeueiiai aaiaoeoo ?
o?a?ico ia/?iee.

Ia 4-oo aeiao i?ney eieaeueii? a?ia?oa?i?? ia/?iee uo??a oaeaee?noue
neioaco ?IE a yae?ao aoea a 1,8 ?ace aeuith ii??aiyii c ?ioaeoieie
oaa?eiaie (?en. 8). Aeoeaaoe?y neioaco ?IE a ee?oeiao ia/?iee
uo??a i?ney eieaeueii? a?ia?oa?i?? ? iane?aeeii
?aaaia?aoeaieo i?ioean?a a

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ffffffffffffffffff7fffefffffe3ffffffffffffffffffffffffffdffffbffffffffff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?en. 8. Ieoiia ?aae?iaeoeai?noue ?IE yaea? (I), oeeoicieth (II) ?
i?e?inii (III) ee?oei ia/?iee ?ioaeoieo uo??a (a), uo??a c
aenia?eiaioaeueiei aaiaoeoii (a), uo??a c aenia?eiaioaeueiei aaiaoeoii,
ui iaea?aeoaaee c?acee a?iiane (a) aai aeno?aeoo (a) Dunaliella
viridis i?ioyaii 3-o ae?a i?ney eieaeueii? a?ia?oa?i?? ia/?iee

ia/?ioe?. Aaaaeaiiy a?iiane D. viridis oaa?eiai i?ney eieaeueii?
a?ia?oa?i?? i?eaaei aei oiai, ui ia

13

4-oo aeiao i?ney iia?aoe?? ieoiia ?aae?iaeoeai?noue ?IE yaea? aoea
anueiai a 1,3 ?ace aeuith, i?ae ieoiia ?aae?iaeoeai?noue ?IE yaea?
ee?oei ia/?iee ?ioaeoieo oaa?ei (?en. 8), oiaoi aaaaeaiiy a?iiane
i?eaaei aei iaaeeaeaiiy aeaiiai iieacieea aei ii?ie. Aaaaeaiiy aeno?aeoo
iacia/ii ciaiooaaei ieoiio ?aae?iaeoeai?noue ?IE yaea? ee?oei ia/?iee
uo??a c aenia?eiaioaeueiei aaiaoeoii (?en. 8).

Ieoiia ?aae?iaeoeai?noue ?IE oeeoicieth ee?oei ia/?iee uo??a c?inea
ii??aiyii c ?ioaeoieie uo?aie ye ? ?IE yaea? a 1,8 ?ace, ui na?ae/eoue
i?i oa, ui oaeaee?noue o?ainii?oo iiaineioaciaaii? ?IE aei oeeoiieacie
c?inea i?iii?oe?eii oaeaeeino? neioaco ?IE i?ney eieaeueii? a?ia?oa?i??.
Ieoiia ?aae?iaeoeai?noue ?IE o?aeoe?? i?e?inii ca?eueoeeany a iaio?e
i??? — a 1,4 ?ace. Iiaeeeai, oey o?aeoe?y ?IE, ui i?aaenoaaeaia a
iniiaiiio ??IE ? aoiaeeoue aei neeaaeo ?iainii, ca’ycaieo c iaia?aiaie
AI?, iai?ith?oueny iia?euei?oa, i?ae ?IE oeeoicieth (?en. 8). Aaaaeaiiy
c?ace?a a?iiane i?eaiaeeei aei ii?iae?caoe?? oaeiae ? oeeo iieaciee?a.
Ieoiia ?aae?iaeoeai?noue ?IE oeeoicieth ? i?e?inii aoea a oeueiio
aa??aio? eeoa a 1,3 ?ace aeuith, i?ae a ?ioaeoieo oaa?ei (?en. 8).
C?acie aeno?aeoo ?ioei /eiii aieeaaa ia aeine?aeaeoaai? iieacieee
ii??aiyii c? c?aceii a?iiane D. viridis. Ieoiia ?aae?iaeoeai?noue ?IE
oeeoicieth ? i?e?inii aoea aeuith a 1,9 ?ace, i?ae a ?ioaeoieo oaa?ei
(?en. 8), oiae? ye ieoiia ?aae?iaeoeai?noue ?IE yaea? — aeuith a 1,6
?ace. Oea na?ae/eoue i?i cia/ia ca?eueoaiiy oaeaeeino? o?ainii?oo
iiaineioaciaaii? ?IE aei oeeoiieacie ? oaeaeeino? neioaco ?/aai
o?ainii?oo ??IE, ca’ycaii? c i?e?iniiaie, a oaeiae i?i ci?iaiiy /aniaiai
oa?aeoa?o i?ioean?a neioaco ? o?ainii?oo ?IE a ee?oeiao ia/?iee. ??ciee
oa?aeoa? ae?? c?ace?a a?iiane ? aeno?aeoo D. viridis ia ieoiio
?aae?iaeoeai?noue ?IE ia/?iee uo??a iaoiiaeaiee ??cieoeath ?oiueiai
neeaaeo. Oe?eueia a?iiana aiaei?inoae e??i eiiieaeno e?iio?eueieo
?a/iaei i?noeoue aiaei?ic/eii? eiiiiiaioe ee?oei, ui oaeiae iiaeooue
iaoe a?ieia?/io aeoeai?noue, ? a?eee, ye?, noai?thth/e eiiieaene c
e?iio?eueieie eiiiiiaioaie, iiaeooue aieeaaoe ia ?oith noaa?euei?noue ?
a?iaeinooii?noue.

Ia 6-oo aeiao i?ney inoaiiueiai aaaaeaiiy n??/aiieenei? i?ae? ieoiia
?aae?iaeoeai?noue ?IE noaee?oeiieo o?aeoe?e ee?oei ia/?iee uo??a, ui
iaea?aeoaaee ye ae?oea?ei, oae ? iaeaeaa c?acee, ia a?ae??ciyeany a?ae
a?aeiia?aeieo iieaciee?a ?ioaeoieo oaa?ei. E?i?aeiee neeaae oeeoicieth
ee?oei ia/?iee ? ne?iaaoee e?ia? o an?o a?oiao oaa?ei ia ?ic??ciyany. Ia
6-oo aeiao i?ney o?e?aciaiai aaaaeaiiy n??/aiieenei? i?ae?
niinoa??aaeiny 20%-a ciaioaiiy ai?noo o?eaoeeeae?oea?eae?a a i?e?iniiao
ee?oei ia/?iee. AEiaeai c aeine?aeaeoaaieo c?ace?a ia i?eaiaeea aei
a?aeiiaeaiiy oeueiai iieacieea.

Oaeei /eiii, c?acee ye a?iiane, oae ? aeno?aeoo D. viridis aieeaaee ia
ieoiio ?aae?iaeoeai?noue ?IE noaee?oeiieo o?aeoe?e ee?oei ia/?iee uo??a
ia 4-oo aeiao i?ney eieaeueii? a?ia?oa?i?? ia/?iee. O?e?aciaa aaaaeaiiy
a?iiane D. viridis i?eaiaeeei aei ciaioaiiy ieoiii? ?aae?iaeoeaiino? ?IE
an?o aeine?aeaeoaaieo noaee?oeiieo o?aeoe?e ii??aiyii c oaa?eiaie, ui
iaea?aeoaaee ae?oea?ei i?ioyaii 3-o ae?a i?ney iia?aoe??.
Aeno?aeo iacia/ii ciaiooaaa ieoiio

14

?aae?iaeoeai?noue ?IE yaea?, aea i?e oeueiio i?eaiaeea aei ca?eueoaiiy
ieoiii? ?aae?iaeoeaiino? ?IE oeeoicieth ? i?e?inii.

AENIIAEE

1. A i?ioean? eoeueoeaoaaiiy ee?oei Dunaliella ci?ith?oueny
?ioaineai?noue neioaco ioeea?iiaeo eeneio, a ia 16-oo-20-oo aeiao
niinoa??aa?oueny a?aeiinia neio?ii?caoe?y i?ioean?a i?ie?oa?aoe?? ee?oei
a eoeueoo??.

2. A i?ioean? eoeueoeaoaaiiy ee?oei Dunaliella ciaioo?oueny oaeaee?noue
iiaeeiaiiy i?/aiiai H14CO3, ui iia’ycaii c? ci?iaiiyi o?ceei-o?i?/ieo
aeanoeainoae eoeueoo?aeueii? ??aeeie.

3. A iioeiaeueieo oiiaao eoeueoeaoaaiiy (25 C, e?oaeiaeiaiaa
ina?oeaiiy, 6 eee) a ee?oeiao D. salina i?noeeiny a 2 ?ace a?eueoa
ea?ioeii?ae?a ? a 1,5 ?ace a?eueoa iaeo?aeueieo e?i?ae?a ii??aiyii c D.
viridis. A ee?oeiao D. salina ? D. viridis aeecueei 55% a?ae on?o
iaeo?aeueieo e?i?ae?a i?eiaaea? ia iaaoa?eo?eiaai? aee?i? eeneioe.
Aeyaeaii ?nioaaiiy aaciaiai ??aiy ai?noo ea?ioeii?ae?a.

4. Ca oaeaee?noth neioaco ea?ioeii?aee D. salina ? D. viridis
?icoaoiaothoueny a oae?e iine?aeiaiino?: b-ea?ioei>enaioio?ee
a?ieaenaioeiiaiai oeeeeo>ethoa?i. Ia aaciaiio ??ai? ai?no b-ea?ioeio a
ee?oeiao D. viridis caeeoa?oueny iaci?iiei a i?ioean? eoeueoeaoaaiiy, a
a ee?oeiao D. salina — ci?ith?oueny ?eoi?/ii.

5. Ia ii/aoeiaeo noaae?yo eoeueoeaoaaiiy D. salina (4-oa-8-ia aeiaa
?inoo) niinoa??aa?oueny i?yia ei?aeyoe?y i?ae oaeaee?noth neioaco AeIE ?
ai?noii ea?ioeii?ae?a. Iaaeae? (12-oa-16-oa aeiaa) oa?aeoa? caeaaeiino?
i?ae oaeaee?noth neioaco AeIE ? iaoaaie?ciii ea?ioeii?ae?a ci?ith?oueny
ia i?ioeeaaeiee.

6. Oaeaee?noue iiae?eo ee?oei Dunaliella caeaaeeoue a?ae ii/aoeiai?
eiioeaio?aoe?? ee?oei a eoeueoo??. Iaea?eueoee i?e??no e?eueeino? ee?oei
niinoa??aa?oueny ca ii/aoeiai? eiioeaio?aoe?? 2,5 iei. ee?oei a 1 ie
eoeueoo?aeueiiai na?aaeiaeua.

7. ?iaeaoaaiiy ?inoo eoeueoo?e Dunaliella ca?eueoaiiyi eiioeaio?aoe??
ee?oei aei 10 iei./ie ? ca?eueoaiiy ina?oeaiiy aei 8 eee ? oaiia?aoo?e
eoeueoeaoaaiiy aei 38-40 C noi?iaiaeaeoaaeiny ca?eueoaiiyi ai?noo
b-ea?ioeio a ee?oeiao D. salina ? D. viridis a 2-3 ?ace, noa?i?ae?a — a
2-3 ?ace, iaaoa?eo?eiaaieo aee?ieo eeneio — a 2-4 ?ace,
o?eaoeeeae?oea?eae?a ? ao???a noa?i?ae?a — a 2-3 ?ace. Oaeei /eiii, aoa
aeyaeaiee ?iaeoeiaaiee ??aaiue neioaco b-ea?ioeio a ee?oeiao Dunaliella,
ca yeiai aeaeiaa ??cieoey i?ae D. salina ? D. viridis ca ai?noii
ea?ioeii?ae?a ? iaeo?aeueieo e?i?ae?a iiai?noth e?ea?aeoaaeany.

8. C?acee a?iiane ? e?i?aeii-ea?ioeii?aeiiai aeno?aeoo D. viridis
aieeaathoue ia i?ioeane neioaco ? o?ainii?oo ?IE a ee?oeiao ia/?iee
uo??a c aenia?eiaioaeueiei aaiaoeoii ? iiaeooue aooe ?aeiiaiaeiaaieie
aeey ?ic?iaee aaiaoio?iiieo e?ea?nueeeo i?aia?ao?a.

15

NIENIE I?AOeUe. UI A?AeIA?AAEATHOUe

INIIAI? IIEIAEAIIss AeENA?OAOe??

1. Oiiyeia A.A., Aeiaaaeeia O.A., Eiia?enoay A.I. Aeaenoaea
noaeaoaeueiuo aeic eiiia iaaee ia eoeueoo?o Dunaliella viridis Teod.
(Chlorophyta) // Aeueaieiaey. — 1994. — O. 4, N4. — N. 30-35.

2. Eiia?enoay A.I., Aiaeeia A.E., Aeiaaaeeia O.A. Eioaineaiinoue
neioaca ioeeaeiiauo eeneio o aiaei?ineae ?iaea Dunaliella Teod. a
i?ioeanna ia?eiaee/aneiai eoeueoeae?iaaiey // Aeueaieiaey. — 1997. — O.
7, N4. — N. 350-357.

3. Eiia?enoay A.I. Aeeyiea enoiaeiie eiioeaio?aoeee eeaoie e aic?anoa
eoeueoo?u ia aeeiaieeo ?inoa e iaeiieaiea b-ea?ioeia a eeaoeao
Dunaliella viridis Teod. // Aeieiae/aneee aanoiee. — 1997. — O. 1, N1.
— N. 100-106.

4. Eiia?enoay A.I. Aeeyiea aeiiannu e oei?ioeee-ea?ioeiieaeiiai
yeno?aeoa Dunaliella viridis Teod. ia neioac ?IE a eeaoeao ia/aie e?un
n yenia?eiaioaeueiui aaiaoeoii e oea?icii ia/aie // Aeieiae/aneee
aanoiee. — 1998. — O.2, N1. — N. 57-60.

5. Aiaeeia A.E., Iaicyiiaa I.A., Eiia?enoay A.I. Ianniaia
eoeueoeae?iaaiea iee?iaiaei?ineae e iieo/aiea aeiiannu ii?aaeaeaiiiai
ninoaaa. // Na?aeioeoai i?i aea?aeaaio ?a?no?aoe?th i?aa aaoi?a ia oa??
IA N641. Ca?a?no?iaaii a Aea?aeaaiiio aaaionoa? Oe?a?ie c aaoi?nueeeo ?
noi?aeieo i?aa 30 eeiiy 1997 ?ieo. — 9 n.

6. Eiia?enoay A.I. Aeeyiea ?aciuo aeic eiiia iaaee ia yene?aoeeth
aaeeia eeaoeaie aiaei?ineae Dunaliella viridis Teod. // Aeooaeuei?
ieoaiiy aioai?ee ? aeieia??. Eiioa?aioe?y iieiaeeo o/aieo ?
niaoe?ae?no?a, (19-21 aeiaoiy 1993, sseoa). Oace aeiiia?aeae. — Ee?a,
1993. — n. 61.

7. Eiia?enoay A.I. Aeeyiea oaiiiaiai ia?eiaea ia auoiae aeiiannu e
b-ea?ioeia o aeaoo aeaeia Dunaliella Teod. // Iaoa?eaeu iao/iie
eiioa?aioeee iieiaeuo o/aiuo aeieiae/aneiai oaeoeueoaoa e
iao/ii-enneaaeiaaoaeueneiai einoeoooa aeieiaee. — Oa?ueeia, 1996. — n.
30.

8. Eiia?enoay A.I. Caaeneiinoue aeeiaieee ?inoa eoeueoo? iaeiieeaoi/iuo
aiaei?ineae io iiioeyoeeiiiiai aic?anoa enoiaeiie eoeueoo?u // Iaoa?eaeu
iao/iie eiioa?aioeee iieiaeuo o/aiuo aeieiae/aneiai oaeoeueoaoa e
iao/ii-enneaaeiaaoaeueneiai einoeoooa aeieiaee. — Oa?ueeia, 1996. — n.
27-28.

9. Eiia?enoay A.I. Caaeneiinoue nei?inoe ?inoa eoeueoo?u iaeiieeaoi/iuo
i?aaieciia iinea ia?anaaa io aic?anoa enoiaeiie eoeueoo?u // A?ieia?/i?
iaoai?cie noa??iiy. I?aeia?iaeiee n?iiic?oi (15-17 o?aaiy 1996 ?ieo),
oace aeiiia?aeae. — Oa?e?a, 1996. — n. 75.

16

AIIOAOe?ss

Eiia?enoa A.I. Aca?iica’ycie ?ioaineaiino? i?ie?oa?aoe?? ?
e?i?aeii-ea?ioeii?aeiiai iai?io a ee?oeiao Dunaliella Teod. oa aieea
e?i?aeii-ea?ioeii?aeieo aeno?aeo?a ia ooieoe?iiaeueio aeoeai?noue
aaiaoioeeo?a. — ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo
iaoe ca niaoe?aeuei?noth 03.00.04 — a?io?i?y. — Oa?e?anueeee aea?aeaaiee
oi?aa?neoao I?i?noa?noaa ina?oe Oe?a?ie, Oa?e?a, 1999.

Aeena?oaoe?th i?enay/aii aeine?aeaeaiith aca?iica’yceo iaoaaie?cio
ea?ioeii?ae?a, e?i?ae?a ? i?ie?oa?aoe?? ee?oei Dunaliella, a oaeiae
aieeao e?i?aeii-ea?ioeii?aeieo aeno?aeo?a oeeo i?e?iaiaei?inoae ia
ooieoe?iiaeueio aeoeai?noue ia/?iee i?e aaiaoeo? ? o?a?ic?.
Noi?ioeueiaaii a?iioaco aaciaiai oa ?iaeoeiaaiiai ??aiy ea?ioeiiaaiaco a
ee?oeiao aiaei?inoae. Aeey aaciaiai ??aiy oa?aeoa?iith ? i?yia
ei?aeyoe?y ?ioaineaiino? i?ie?oa?aoe?? c ai?noii ea?ioeii?ae?a o
ee?oei?, aeey ?iaeoeiaaiiai — cai?ioia ei?aeyoe?y c ?ioaineai?noth
i?ie?oa?aoe?? ? i?yia — c iaeiie/aiiyi iaeo?aeueieo e?i?ae?a. C?acee
a?iiane ? aeno?aeoo D. viridis aieeaathoue ia neioac ? o?ainii?o ?IE a
ee?oeiao ia/?iee uo??a c aaiaoeoii ? ?aeiiaiaeothoueny aeey ?ic?iaee
aaiaoio?iiieo i?aia?ao?a.

Eeth/ia? neiaa: Dunaliella, i?ie?oa?aoe?y, ea?ioeii?aee, iaeo?aeuei?
e?i?aee, aaiaoeo, o?a?ic ia/?iee.

AIIIOAOeEss

Eiia?enoay A.I. Acaeiinaycue eioaineaiinoe i?ieeoa?aoeee e
eeieaeii-ea?ioeiieaeiiai iaiaia a eeaoeao Dunaliella Teod. e aeeyiea
eeieaeii-ea?ioeiieaeiuo yeno?aeoia ia ooieoeeiiaeueioth aeoeaiinoue
aaiaoioeeoia. — ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.04 — aeioeiey. — Oa?ueeianeee
ainoaea?noaaiiue oieaa?neoao Ieienoa?noaa ia?aciaaiey Oe?aeiu, Oa?ueeia,
1999.

Aeenna?oaoeey iinayuaia enneaaeiaaieth acaeiinayce iaoaaieecia
ea?ioeiieaeia, eeieaeia e i?ieeoa?aoeee eeaoie Dunaliella, a oaeaea
aeeyiey eeieaeii-ea?ioeiieaeiuo yeno?aeoia yoeo iee?iaiaei?ineae ia
ooieoeeiiaeueioth aeoeaiinoue ia/aie i?e aaiaoeoa e oea?ica.
Noi?ioee?iaaia aeiioaca aaciaiai e eiaeooee?iaaiiiai o?iaiy
ea?ioeiiaaiaca a eeaoeao aiaei?ineae. Aeey aaciaiai o?iaiy oa?aeoa?ia
i?yiay ei??aeyoeey eioaineaiinoe i?ieeoa?aoeee n niaea?aeaieai
ea?ioeiieaeia a eeaoea, aeey eiaeooee?iaaiiiai — ia?aoiay ei??aeyoeey n
eioaineaiinoueth i?ieeoa?aoeee e i?yiay — n iaeiieaieai iaeo?aeueiuo
eeieaeia. Ia?acoeu aeiiannu e yeno?aeoa D. viridis aeeytho ia

17

neioac e o?ainii?o ?IE a eeaoeao ia/aie e?un n aaiaoeoii e
?aeiiaiaeiaaiu aeey ?ac?aaioee aaiaoio?iiiuo i?aia?aoia.

Eeth/aaua neiaa: Dunaliella, i?ieeoa?aoeey, ea?ioeiieaeu, iaeo?aeueiua
eeieaeu, aaiaoeo, oea?ic ia/aie.

ABSTRACT

Komaristaya V.P. Interconnections between proliferation intensity and
lipid-carotenoid metabolism in Dunaliella Teod. and effect of the
lipid-carotenoid extracts on hepatocytes functional activity. —
Manuscript.

Thesis for a candidate’s degree by speciality 03.00.04 — biochemistry.
— Kharkiv State University of Ministry of education of Ukraine.

The dissertation is devoted to investigation of interconnections
between carotenoid and lipid metabolism and proliferation activity in
the unicellular algae of genus Dunaliella cells as well as to definition
of their lipid-carotenoid extracts effect on several liver functional
activity indexes under hepatitis and liver fibrosis.

While cultivating Dunaliella the intensity of nucleic acids synthesis
in the cells was changing and on the 16-th-20-th days the relative
synchronization of cell proliferation processes occurred in the culture.
This fact was discovered using radioactive labeling of nucleic acids
with NaH14CO3 and confirmed by DNA labeling with [14C]thymidine. The
H14CO3 absorption rate decreased in the cultivation process and that was
connected to the physico-chemical properties of culture medium changing
during the culture growth. After changing culture medium for the fresh
one the incorporation of H14CO3 in the cells of cultures of different
ages became equal.

The b-carotene content in D. salina cells was changing rhythmically in
the process of culture growth under the optimal conditions. On the
initial stages of D. salina culture growth the straight correlation
between b-carotene content and DNA synthesis was observed but on the
further stages of the growth the correlation became reverse. The
b-carotene content in D. viridis cells remained constant during the
cultivation. The neutral lipids content in the cells of the both species
remained unchanged during the cultures growth and this fact made it
possible to conclude that it was independent on carotenoid metabolism
and the rhythmical pattern of b-carotene content in D. salina cells
depended on carotenoids interconvertions themselves.

The investigation of different carotenoid fractions synthesis in the
both species showed the individual patterns of synthesis rate for
b-carotene, lutein and the violaxanthin cycle xanthophylls that were

18

depended on the culture growth stage. b-carotene was the most rapidly
synthesized carotenoid in Dunaliella cells, violaxanthin cycle
xanthophylls came the next, and lutein was the most slowly synthesized.
It was somewhat surprising as the lutein content was the greatest — 60%
of the total carotenoid content in the cell. Using H14CO3 as the
precursor it could not be a success to evaluate the input of the
different fractions synthesis in the b-carotene content fluctuations
during the culture growth because of different H14CO3 absorbtion rates
in the cultures of different ages. Nevertheless, carotenoid metabolism
was under control of different conditions which were changing during the
cultures growth such as the concentration of cells was.

The inhibition of Dunaliella culture growth by cells concentration
increase, high light intensity and superoptimal temperature led to the
b-carotene, triacylglycerides, sterols and their esters content 2-3 time
increase and the non-esterified fatty acids content 2-4 time increase in
the cells of the both species.

It was proposed the hypothesis of basic and induced carotenogenesis
levels in the algae cells. On the basic level the straight correlation
between proliferation intensity and b-carotene content was
characteristic but the correlation with neutral lipids accumulation was
absent. On the induced level of carotenogenesis the reverse correlation
with proliferation intensity and the straight connection with neutral
lipids accumulation were characteristic. On the basic level under the
optimal cultivation conditions D. salina cells contained 2 times more
carotenoids and 1,5 times more neutral lipids than D. viridis cells did.
Our data showed that the both species investigated under the inductive
conditions may be prospective carotenoids and lipids producers.

The samples of D. viridis biomass and lipid-carotenoid extract
influenced RNA synthesis and transport processes in liver cells of the
rats suffering from the experimental hepatitis induced by the local
hyperthermy of liver. Three time administration of D. viridis biomass
led to RNA labeling with [14C]orotic acid decrease in nuclei, cytosol
and microsomes fractions of rat liver cells comparing to the animals
that had obtained placebo. The extract decreased RNA putative
radioactivity in nuclei slightly but increased the one in cytosol and
microsomes. The samples did not influence RNA synthesis in rat liver
cells under the experimental fibrosis induced by copper sulfate
administration and the neutral lipids content in liver cells under both
hepatitis and fibrosis. The samples were recommended for hepatotropic
drugs elaboration.

Key words: Dunaliella, proliferation, carotenoids, neutral lipids,
hepatitis, liver fibrosis.

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