iaoe?iiaeueiee aa?a?iee oi?aa?neoao

AeAEAI IEAENAIAe? AIEIAeEIE?IAE*

OAeE 632.7:633.256+92

?ICAEOIE EI?AIAAEO AIEEAE ss?IAI ss*IAITH I?NEss CANOINOAAIIss
AA?A?OeEAe?A O I?AI?*IIIO E?NINOAIO OE?A?IE

06.01.11 — o?oiiaoieia?y

A a o i ? a o a ? a o

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

Ee?a 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia o a?aeae?e? caoenoo ca?iiaeo eoeueoo? a?ae oai?ia oa
oe?aeiee?a ?inoeoooo caoenoo ?ineei Oe?a?inueei? aeaaeai?? aa?a?ieo
iaoe.

Iaoeiaee ea??aiee – eaiaeeaeao a?ieia?/ieo iaoe

EENAIEI Na?a?e Aieiaeeie?iae/,

?inoeooo caoenoo ?ineei OAAI, caa?aeoaa/ a?aeae?eii caoenoo ca?iiaeo
eoeueoo? a?ae oai?ia oa oe?aeiee?a

Io?oe?ei? iiiiaioe – aeieoi? n?euenueeiainiiaea?nueeeo iaoe, aeioeaio

OAA*OE Aaeaioeia Einoyioei?aia,

Eai’yiaoeue-Iiae?euenueeee aea?aeaaiee iaaeaaia?/iee oi?aa?neoao,

aeioeaio eaoaae?e a?ieia??

eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe

AEEI’ssCIEE Aieiaeeie? Aiaoie?eiae/,

Iaoe?iiaeueiee aa?a?iee oi?aa?neoao,

anenoaio eaoaae?e o?oiiaoieia??.

I?ia?aeia onoaiiaa – ?inoeooo caiea?ianoaa OAAI.

Caoeno a?aeaoaeaoueny “26” ethoiai 1999 ?. i 10 aiae. ia can?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae 26.004.02 a Iaoe?iiaeueiiio aa?a?iiio
oi?aa?neoao? ca aae?anith: 252041, i.Ee?a-41, aoe. Aa?i?a Iai?iie 15,
iaa/aeueiee ei?ion ? 3, aoaeeoi??y 65.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? Iaoe?iiaeueiiai
aa?a?iiai oi?aa?neoaoo ca aae?anith: 252041, i.Ee?a-41, aoe. Aa?i?a
Iai?iie 11, iaa/aeueiee ei?ion ? 10.

Aaoi?aoa?ao ?ic?neaiee “26” n?/iy 1999 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee Iaiaeaeoe A.?.

1

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. A aa?ii?iieneiaiio eiiieaeni aio?iiiaaiia
aeiyeueiinoue iai?aaeaia ia oi?ioaaiiy cai?ia?aiiaaii?
aenieii?iaeoeoeaii? aa?iaeinenoaie. I?ioa ia no/aniiio aoai? ?icaeoeo
caiea?ianoaa iaiaaeoth/eie oaeoi?aie iioaioe?eieo iiaeeeainoae
iaea?aeaiiy aenieiai o?iaeath oa yeino? ca?ia y?iai y/iaith ?
o?aaeai?noue iin?a?a ei?aiaaeie aieeyie oa ?o caao?’yiai?noue. I?i
oeiaei/eii?noue ei?aiaaeo aieeae na?ae/aoue oae? aeai?: na?aaei? ao?aoe
ca?ia y/iaith a?ae ei?aiaaeo aieeae noaiiaeyoue 15,4%, iaaeia?? o?iaeath
– a?ae 0,7 aei 3,0 oe/aa (Ei?ooiiaa A.?., *oiaeia A.A., Uaei/eoeia ?.E.,
1976; Aeia?aoeia A.I.,1962; Oaiiiooae I.I., 1979). O naith /a?ao ao?’yie
iaoiiaeththoue 10-11% caaaeueieo ao?ao oa?/iaeo i?iaeoeoia o anueiio
naioi.

O niaoe?aeuei?e e?oa?aoo?? ? iia?aeiieaiiy i?i aieea aa?a?oeeae?a ia
?icaeoie oai?ia ?ineei (Enaaaa E.E., 1974, 1984; Aiaaiaeei A.A., 1986,
1988). sse o?i?/ii aeoeai? niieoee aiie iiaeooue aooe oiene/ieie aeey
caoaeiee?a oai?ia ?ineei (Eiiae?aoueaa TH.A., 1980). O?i?/i? caniae
caoenoo ?ineei a?ae??ciythoueny ??ciiiai?oiith ae??th ia ?ineeie ?
aeeeeeathoue o ieo o?c?ieia?/i? oa ii?oieia?/i? ci?ie, ye?, a naith
/a?ao, a?aeia?aaeathoueny ia ?icaeoeo iaoiaai?a. O ca’yceo c oeei
i?iaaaeaiiy aeine?aeaeaiue uiaei aea/aiiy ?aaeoe?? iaoiaaiii? i?e?ioei?e
a?oioo i?ney canoinoaaiiy aa?a?oeeae?a ? aeooaeueiei.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie. ?iaioa aeeiiaia a ?aieao
aaeocaaeo ? aea?aeaaieo i?ia?ai:

“?ic?iaeoe ciiaeuei? nenoaie ?ioaa?iaaiiai caoenoo ?ineei a?ae
oe?aeeeaeo i?aai?ci?a, ui caaacia/othoue iioei?caoe?th o?oinai?oa?iiai
noaio aa?ioeaiic?a ? cieaeaiiy ao?ao o?iaeath” (aea?aeaaia ?a?no?aoe?y
ca ? 01970012341).

“Noi?ioaaoe ani?oeiaio iiaeo ianoeoeeae?a, ?ic?iaeoe oa aaeineiiaeeoe
oaoiieia?? ?o canoinoaaiiy” (aea?aeaaii? ?a?no?aoe?y ca ? 01970012336).

Iaoa ? caaaeaiiy aeine?aeaeaiue. Aieiaiith iaoith aeineiaeaeaiue aoei
aea/aiiy aieeao aa?a?oeeae?a ia ?icaeoie ei?aiaaeo aieeae y?iai y/iaith
a oiiaao i?ai?/iiai Eininoaio oa ia eiiiiiaioe aa?iaeinenoaie, ye?
eiio?ieththoue ?icaeoie oei?? oai?iae.

Aeinyaiaiiy aieiaii? iaoe caeienithaaeinue oeyoii:

– ooi/iaiiy neiioiiia o?aaeaiiy ?ineei caoaeieeaie ei?aiaaeo aieeae
iiney canoinoaaiiy aa?aioeeaeia;

– aecia/aiiy ?icaeoeo ei?aiaaeo aieeae y?iai y/iaith iiney canoinoaaiiy
aa?aioeeaeia;

– anoaiiaeaiiy aeaeiaiai neeaaeo caoaeieeia cae/aeii? ei?aiaai? aieei
y?iai y/iaith;

– anoaiiaeaiiy aeeiaiiee /enaeueiinoi aaeoa?ie o ?ecinoa?i y?iai
y/iaith;

– aecia/aiiy aieeao aa?aioeeaeia ia aeaeiaee neeaae iie?ioei?e y?iai
y/iaith;

– ioeiiee ooiainoaoe/ii? aei? aa?aioeeaeia ia ie?ai? aeaee a?eaia a
oiiaao in vitro;

– aea/aiiy aei? aa?aioeeaeia ia oice/ii oa oiciieiai/ii yeinoi ca?ia
y?iai y/iaith.

Iaoeiaa iiaecia iaea?aeaieo aeaieo. Aia?oa a oiiaao i?ai?/iiai Eininoaio
Oe?a?ie aea/aii ?icaeoie ei?aiaaeo aieeae y?iai y/iaith i?ney
canoinoaaiiy aa?a?oeeae?a. Na?aae ?ecinoa?ii? i?e?ioei?e aeae?eaii 36
aeae?a a?ea?a, a oiio /ene? caoaeieee ei?aiaaeo aieeae: a?eae ?iaeo
Fusarium, ye? a?aeiinyoueny aei naeoe?e Roseum, Discolor, Elegans,
Martiella oa Bipolaris sorokiniana Shoem.

Anoaiiaeaii, ui i?ae ae??th aa?a?oeeae?a o a?oio? ciaioo?oueny
/enaeuei?noue a?ea?a ?iaeo Fusarium, a oaeiae aaeoa??e ?iae?a
Pseudomonas, Xanthomonas i Bacillus, i?aeaeuo?oueny e?euee?noue a?ea?a
?iae?a Penicillium i Trichoderma.

Aecia/aii a?ieia?/io aoaeoeai?noue aa?a?oeeae?a i?ioe aeaiaeieueieo
ao?’yi?a o iin?aao y?iai y/iaith. Oae, i?e canoinoaaii? o?aci?a, 60%
c.i. ? eaio?io?a, 70% n.e. c ?ic?aooieo 2 e/aa aiia noaiiaeea 96%, a
ae?eiio?a O, 72% a.?., eaio?io?a, 70% n.e. (1,5 e/aa) – 93%.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. Ia iniia? iaea?aeaieo
?acoeueoao?a aeine?aeaeaiue ca?a?no?iaai? ? aianai? a «Ia?aeie
ianoeoeeaeia i aa?ioiiieaoia, aeicaieaieo aei aeei?enoaiiy a Oe?a?ii»
aa?a?oeeaee o?aci?, ae?eiio? O aeey canoinoaaiiy ia iin?aao y?iai
y/iaith. ?acoeueoaoe aeine?aeaeaiue i?ieoee ae?iaie/o ia?aa??eo o ENI
“Aa?eoianueea” Ee?anueei? iaeano? ia ieiu? 50 aa.

Iniaenoee aianie caeiaoaa/a. Aaoi? aaciina?aaeiuei a?aa o/anoue o
ieaioaaii? oa i?iaaaeaii? aeine?ae?a ? niinoa?aaeaiue, ia?iaoe? aeaieo,
aiae?oe/i?e ?iaio?.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iniiaii ?acoeueoaoe aeineiaeaeaiue
aeiiiaiaeaeenue ia Iiaeia?iaeiie iaoeiai-i?aeoe/iie eiioa?aioei?
iieiaeeo a/aieo i niaoeiaeinoia «Oeyoe ?aoeiiiaeueiiai aeei?enoaiiy
caiaeueieo ?ano?nia Oe?a?ie» (n.i.o. *aaaie, ?inoeooo caiea?ianoaa OAAI,
16-17 aa?aciy 1995 ?ieo); ia eiioa?aioei? iieiaeeo a/aieo i
niaoeiaeinoia, i?enay/aiie 50-?i//th Iinoeoooo caoenoo ?ineei OAAI
“I?iaeaie caoenoo ?ineei a?ae oe?aeeeaeo i?aai?ci?a o no/anieo
aeiiii?/ieo oa aeieia?/ieo oiiaao” (i.Ee?a, 13-14 aa?aciy 1996 ?ieo), ia
1-io c’?caei aa?aieiaia Oe?a?ie (i.Ee?a, 1996 ?.). ?acoeueoaoe
aeine?aeaeaiue ii aeena?oaoe?ei?e ?iaio? iaaiai?thaaeenue o a?aeae?e?
caoenoo ca?iiaeo eoeueoo? a?ae oai?ia oa oe?aeiee?a, ia can?aeaii?
a/aii? ?aaee ?inoeoooo caoenoo ?ineei OAAI (1995, 1996, 1997 ??.).
Aeena?oaoe?y caneooaia ? iaaiai?aia ia ?icoe?aiiio can?aeaii? a?aeae?eo
caoenoo ca?iiaeo eoeueoo? a?ae oai?ia oa oe?aeiee?a ?C? OAAI, ia
can?aeaii? a/aii? ?aaee ?inoeoooo caoenoo ?ineei OAAI o 1998 ?ioe?.

Ioae?eaoe??. Ca iaoa??aeaie aeena?oaoe?? iioae?eiaaii 7 iaoeiaeo
i?aoeue.

No?oeoo?a oa ianya aeena?oaoe??. Aeena?oaoe?eia ?iaioa aeeeaaeaia ia 129
noi??ieao iaoeiiieniiai oaenoo. Aiia neeaaea?oueny ?c anooio, caaaeueii?
oa?aeoa?enoeee ?iaioe, iaeyaeo e?oa?aoo?e, iieno iaoa??ae?a ? iaoiae?a
aeine?aeaeaiue, 3 ?icae?e?a aenia?eiaioaeueii? /anoeie, aeniiae?a,
ia?ae?eo iineeaiue, 5 aeiaeaoe?a. O aeena?oaoe?? i?noeoueny 25
oaaeeoeue, 18 iaethie?a. Ia?ae?e e?oa?aoo?ieo aeaea?ae ioiieth? 253
iaeiaioaaiiy, a oiio /ene? 52 ?ioiiiaieo.

Aeneiaethth ue?o iiaeyeo eaiaeeaeaoia? n?euenueeiainiiaea?nueeeo iaoe
*aaaio Aaneeth Neaei?iae/o ca aeiiiiiao a caaa?oaii? ?iaioe oa
i?aeaioiaoe? ?? aei caoenoo.

1. IAEssAe E?OA?AOO?E

Ia iniia? aiae?co ocaaaeueiaiiy aeaieo e?oa?aoo?e aena?oeaia ?noi??y
aea/aiiy, ?iciianthaeaeaiiy ? oeiaei/eii?noue ei?aiaaeo aieeae, iienaia
aeieiai-a?ieia?/ia oa?aeoa?enoeea caoaeiee?a caoai?thaaiiy.

2. IAOIAeEea oa oiiae I?IAAAeAIIss AeINE?AeAEAIUe

Aeena?oaoe?eia ?iaioa aeeiioaaeanue i?ioyaii 1994-1997 ??. ia
aeine?aeieo iieyo ?C? OAAI o Aa?eo?anueeiio ?-i? Ee?anueei? iae., o
a?aeae?e? caoenoo ca?iiaeo eoeueoo? a?ae oai?ia oa oe?aeiee?a ?C? OAAI,
eaoaae?? i?e?ia?ieia?? ? a??onieia?? IAO, ?inoeooo? i?e?ia?ieia?? ?
a??onieia?? ?i. Ae.E.Caaieioiiai.

Ca aa?iee?iaoe/iei ?aeiioaaiiyi ?aeii i?iaaaeaiiy aeine?aeaeaiue
ciaoiaeeoueny a i?ai?/iiio E?ninoaio Oe?a?ie. Ee?iao iii??ii
eiioeiaioaeueiee. A?oio ? na?aaeiuei?iaeth/ei, na?aaeiueicaaacia/aiei ca
ai?noii i?o?aoiiai oa ai?a/iiai acioo, oinoi?o ? eae?th. A on? ?iee
i?e?iaeiee ?ioaeoe?eiee oii aoa aeinoaoi?i aeey ae??oaiiy iinoaaeaieo
caaaeaiue uiaei aea/aiiy oai?iae a iieueiaeo oiiaao.

I?e i?iaaaeaii? eaai?aoi?ii-iieueiaeo aeine?ae?a aeei?enoai?
caaaeueiii?eeiyo? iaoiaeeee i?iaaaeaiiy aeine?aeaeaiue (?iaeanoaaineee
A.I., 1958; Iaiieeia A.Ae., 1966; Aeiniaoia A.A., 1968, 1972).

Iae?e ei?aiaaeo aieeae y?iai y/iaith i?iaiaeeee a oace o?oaeoaaiiy ?
iiei/ii? noeaeino? ca?ia ca iaoiaeeeith A?C? (1984). I?iae ?ineei
a?aeae?aee ia aeaio noi?aeieo ?yaeeao caaaeiaaeee 0,5 i a /ioe?e?acia?e
iiaoi?iino?. Aeaeiaee neeaae caoaeiee?a ei?aiaaeo aieeae y?iai y/iaith
oa niinoa?aaeaiiy ca ?inoii a?ea?a i?iaiaeeee ca iaoiaeeeith E?oa?iiaa
I.A. (1969) ? A?eae A.E. (1982) c aeei?enoaiiyi i?e?ineii?a IAAe-6 oa
“Nikon”.

I?iae aeey aeine?aeaeaiiy ?ecinoa?ii? i?e?ioei?e a?aeae?aee ia 5-oo,
30-oo oa 60-oo aeiao i?ney canoinoaaiiy aa?a?oeeae?a ca iaoiaeeeith
Ca’ya?ioeaaa Ae.A. (1989). I?e i?aeaioiaoe? a?oioo aeey
i?e?ia?ieia?/iiai iin?ao aeei?enoiaoaaee iaoiae ?icaaaeaiue (Cayaeioeaa
Ae.A., Anaaaa E.A., Aaaueaaa E.I., Ie?/eie O.A., 1980). Iin?a a?oioiai?
noniaic?? cae?enithaaee ia aeeaeeuei? na?aaeiaeua: nonei-aaa?,
na?aaeiaeua *aiaea ? ea?oiieyiee aaa? o noa?eeueieo /aoeao Iao??.
Iiaoi?i?noue – /ioe?e?aciaa (Io?iioeaa A.N., Ii?aeoiiaa A.I., *a?ieeiaa
I.E., 1976).

?aeaioeo?eaoe?th aeine?aeaeoaaiiai i?aai?cio i?iaiaeeee ca aeiiiiiaith
aecia/iee?a (Ieeueei A.Ae., 1947; Eeoaeiia I.A., 1967; Ee?eeaiei O.N.,
1977; Ieaeiiee/ei I.I., Ieeueei A.Ae., 1977; Aaeeeaiia E.E., Neaei?iaa
E.E., 1982).

E?euee?noue a?eaieo ca?iaee?a ?ic?aoiaoaaee ia 1 a aaniethoii nooiai
a?oioo ca iaoiaeeeith Ca’ya?ioeaaa Ae.A. oa ?i. (1991). Aeey aecia/aiiy
e?eueeino? aaeoa??e ? iiaeaeueoiai aeae?eaiiy /enoeo eoeueoo?
ei?enooaaeeny iaoiaeii eoeueoeaoaaiiy i?e?ii?aai?ci?a ia iiaeeaieo
na?aaeiaeuao (IIA ? ea?oiieyieo aaa?ao) (Oaiia? A.C., Oeeueieeiaa A.E.,
1987).

Ioe?ieo ooia?noaoe/ieo aeanoeainoae aa?a?oeeae?a ia ie?ai? aeaee a?ea?a
a oiiaao in vitro i?iaiaeeee ca iaoiaeeeith Aieeoeia I.I. (1970).

Ianoaaeaiiy caao?’yiaiino? iin?a?a y?iai y/iaith cae?enithaaee ca
iaoiaeeeith Nooiaeiaa A.A. (1984). Iae?ee i?iaiaeeee ia ieiuaaeeao 0,25
i2 (50o50 ni), iiaoi?i?noue /ioe?e?aciaa.

Ioe?ieo aieeao aa?a?oeeae?a ia iin?ai? yeino? ian?iiy y?iai y/iaith
i?iaiaeeee ca iaoiaeeeith (Iaoiaee/aneea oeacaiey …, 1981;
Iaoiaee/aneea oeacaiey …, 1988).

Noaoenoe/io ia?iaeo aenia?eiaioaeueieo aeaieo cae?enithaaee iaoiaeii
aeenia?n?eiiai aiae?co (Aeiniaoia A.A., 1972) c aeei?enoaiiyi IAII.

?ACOEUeOAOE AeINE?AeAEAIUe

3. ?ICAEOIE EI?AIAAEO AIEEAE O IIN?AAO ss?IAI ss*IAITH I?NEss AIANAIIss
AA?A?OeEAe?A

3.1. Neiioiiiaaiac ei?aiaaeo aieeae y?iai y/iaith i?e canoinoaaii?
aa?a?oeeae?a

Aena?oeth?oueny neiioiiiaaiac ei?aiaaeo aieeae y?iai y/iaith i?ney
canoinoaaiiy aa?a?oeeae?a o i?ai?/iiio E?ninoaio Oe?a?ie.

A?coaeueieie niinoa?aaeaiiyie ca o?aaeaieie ?ineeiaie ? ianooiiei
aeae?eaiiyi caoaeiee?a ei?aiaaeo aieeae o /eno? eoeueoo?e anoaiiaeaii,
ui a oiiaao i?ai?/iiai E?ninoaio y?ee y/i?iue o?aaeo?oueny
ooca??icii-aaeuei?ioinii??iciith oa oea?einii?aeueiciith ei?aiaaeie
aieeyie. Canoinoaaiiy aa?a?oeeae?a ia aieeaa? ia neiioiie oai?iae oe???
eoeueoo?e.

3.2. Aieea aa?a?oeeae?a ia ?iciianthaeaeai?noue oa ?icaeoie ei?aiaaeo
aieeae y?iai y/iaith

Iieacaii ?iciianthaeaeai?noue ? ?icaeoie ei?aiaaeo aieeae y?iai y/iaith
i?ney canoinoaaiiy aa?a?oeeae?a, ae?th/? ?a/iaeie yeeo iaeaaeaoue aei
??cieo eean?a o?i?/ieo niieoe. Canoinoaaiiy aa?a?oeeae?a i?ioyaii
i?iaaaeaiiy aeine?aeaeaiue (1994-1997 ??.) ia aieeaaei ia
?iciianthaeaeai?noue oa ?icaeoie ei?aiaaeo aieeae y?iai y/iaith ca
aeiyoeii aa??aioa, aea aeei?enoiaoaaany o?aci?, 60% c.i. c ii?iith
aeo?aoe 1,2 ea/aa. I?e aeei?enoaii? oeueiai aa?a?oeeaea a na?aaeiueiio
ca 4 ?iee i?iaaaeaiiy aeine?aeaeaiue o oaco o?oaeoaaiiy a?aei?/aii
ciaioaiiy ia 27% ?icaeoeo ooca??icii-aaeuei?ioinii??icii? ? ia 44,8%
oea?einii?aeueicii? ei?aiaaeo aieeae. O oaco iiei/ii? noeaeino? oaeiae
a?aei?/aii nooo?aa ciaioaiiy ?icaeoeo ei?aiaaeo aieeae y?iai y/iaith:
ooca??icii-aaeuei?ioinii??icii? – ia 18,8% ? oea?einii?aeueicii? – ia
32,8% a?ae eiio?ieth (oaae. 1).

Oaaeeoey 1 – ?icaeoie ei?aiaaeo aieeae y?iai y/iaith i?ney canoinoaaiiy
aa?a?oeeae?a (ni?o ?ieaiae, 1994-1997 ??.)

Aa??aio aeine?aeoOaca o?oaeoaaiiyOaca iiei/ii?
noeaeino?ooca??icii-aaeuei?ioinii??iciaoea?einii?aeueiciaooca??icii-aaeu
ei?ioinii??iciaoea?einii?aeueiciaiioe?aiiy, %?icaeoie, %iioe?aiiy,
%?icaeoie, %iioe?aiiy, %?icaeoie, %iioe?aiiy, %?icaeoie,
%Eiio?ieue45,18,926,72,960,319,236,57,0Ae?eiio? O, 72% a.?., 1,0 e/aa
45,28,026,92,260,217,237,66,2O?aci?, 60% c.i.,

1,2 ea/aa45,36,527,21,660,615,

637,84,7Eaio?io?, 70% n.e., 1,5
e/aa44,38,726,72,160,917,637,06,7Eaio?io?, 70% n.e., 2,0
e/aa45,27,727,01,860,117,737,66,2I??051,71,03,12,0

3.3. Aieea aa?a?oeeae?a ia aeaeiaee neeaae caoaeiee?a cae/aeii?
ei?aiaai? aiee? y?iai y/iaith

A?ieia?/iei aiae?cii o?aaeaieo ?ineei anoaiiaeaii, ui a oiiaao
i?ai?/iiai E?ninoaio Oe?a?ie caaeaaeue noiae?a ? iiao??iiy ei?ai?a y?iai
y/iaith aeeeeeathoue a?eae ?c eeano Deuteromycetes ii?yaeeo
Hyphomycetales, ?iaeeie Tuberculariaceae. Iaea?eueoa e?euee?noue
iaoiaai?a i?aaenoaaeaia aeaeaie ?iaeo Fusarium, ye? a?aeiinyoueny aei
naeoe?? Roseum, Discolor, Elegans, Martiella. Noiooi?ie a?eaaie ?iaeo
Fusarium aoee a?eae ?iae?a Bipolaris, Rhizopus, Alternaria.

Aeine?aeaeaiiyie anoaiiaeaii, ui i?ney canoinoaaiiy aa?a?oeeae?a cia/ii
ci?ith?oueny /anoioa ?cieyoe?? ie?aieo aeae?a a?ea?a caoaeiee?a
ooca??icii-aaeuei?ioinii??icii? ei?aiaai? aiee? y?iai y/iaith. Iaea?eueo
yne?aai oe? ci?ie i?iyaeythoueny o aa??aio?, aea canoiniaoaaee
aa?a?oeeae o?aci?, 60% c.i. (1,2 ea/aa) (oaae. 2). Ii??aiyii c
eiio?ieai, c ei?ai?a y?iai y/iaith ia aeae?eai? F.javanicum, F.javanicum
var. radicicola. Ciaioeeanue /anoioa ?cieyoe?? aeae?a F.avenaceum – ia
50%, F.solani – ia 51,3%, F.avenaceum. var. herbarum – ia 46,9%,
F.solani var. redolense – ia 72,3%. Ii?yae c oeei ia 25,7% ca?eueoeeanue
/anoioa ?cieyoe?? Alternaria alternata; ia 80,9% — Rhizopus spp.; 31,6%
— Bipolaris sorokiniana. Oaeei /eiii, ia iaoo aeoieo, iaei??th c i?e/ei
cieaeaiiy ?icaeoeo cae/aeii? ei?aiaai? aiee? o aa??aio?, aea
canoiniaoaaee aa?a?oeeae o?aci?, 60% c.i. (1,2 ea/aa) ? aeiaaeaiiy
ie?aieo aeae?a a?ea?a. E??i oiai, ci?ia ieoiii? aaae ie?aieo aeae?a
oaeiae iiaea aooe i?e/eiith ciaioaiiy ?icaeoeo ei?aiaai? aiee? y?iai
y/iaith.

4. AIEEA AA?A?OeEAe?A IA I?E?IOEI?O ?ECINOA?E ss?IAI ss*IAITH, UI
EIIO?IETH? ?ICAEOIE CAOAeIEE?A

EI?AIAAI? AIEE?

4.1. Aeeiai?ea /enaeueiino? aaeoa??e o ?ecinoa?? y?iai y/iaith i?e
canoinoaaii? aa?a?oeeae?a

A?aeiii, ui na?aae aaeoa??e ? aeineoue aaaaoi aeae?a c aioaaii?noe/ieie
aeanoeainoyie ii a?aeiioaiith aei caoaeiee?a ei?aiaaeo aieeae (Oai?ae
N.I., 1993). I?ioa a?aeiiinoae i?i oa, ye? ci?ie a?aeaoaathoueny a
?ecinoa?? y?iai y/iaith i?ae aieeaii iiaeo aa?a?oeeae?a a oiiaao
i?ai?/iiai E?ninoaio Oe?a?ie na?aae iaoeiaeo ioae?eaoe?e ia aeyaeaii.
Oiio aoee i?iaaaeai? aeine?aeaeaiiy uiaei anoaiiaeaiiy /enaeueiino? oa
aeeiai?ee ?icaeoeo aaeoa??e o ?ecinoa?? y?iai y/iaith, ia iin?aao yeiai
canoiniaoaaee aa?a?oeeaee. I?ae ?o aieeaii a?aeaoaa?oueny ciaioaiiy a
iaaeao 10-20% /enaeueiino? aaeoa??e ?iaeo Pseudomonas, Xanthomonas,
Bacillus, i?o?eo?eaoi??a o ?ecinoa?? y?iai y/iaith i?ioyaii 30 ae?a
i?ney canoinoaaiiy aa?a?oeeae?a. A?aeiiaeaiiy /enaeueiino? aaeoa??e
a?aei?/aii ia 60-oo aeiao i?ney canoinoaaiiy ianoeoeeae?a (?en. 1). Oiio
ci?ie o /enaeueiino? aaeoa??e o ?ecinoa?? y?iai y/iaith ia iathoue
aieeao ia nooi?iue ?icaeoeo ei?aiaai? aiee? oe??? eoeueoo?e.

4.2. Aeeiai?ea /enaeueiino? a?eaieo ca/aoe?a o ?ecinoa?? y?iai y/iaith
i?ney canoinoaaiiy aa?a?oeeae?a

A oiiaao c?inoath/iai ae?iaieoeoaa n?euenueeiainiiaea?nueei? i?iaeoeoe??
iinoa? caaaeaiiy io?eiaiiy aeaoaeueii? aeioieneeieia?/ii? ?ioi?iaoe??
i?i aieea aa?a?oeeae?a ia i?eia?ioo a?oioo, yea a?ae?a?a? aaaeeeao ?ieue
o aeeoo?ae?yeueiino?9

Oaaeeoey 2 – Neeaae iiioeyoe?? caoaeiee?a ei?aiaaeo aieeae y?iai y/iaith
oa noiooi?o a?ea?a i?ney canoinoaaiiy aa?a?oeeae?a (ni?o ?ieaiae,
1994-1996 ??.)

Aa??aio aeine?aeo*anoioa ?cieyoe??, %Alternaria alternataRhizopus
spp.F.avenaceumF.avenaceum var.herbarumF.oxysporumF.oxysporum
var.orthocerasF.solaniF.solani var.coeruleumF.solani
v.redolenseF.sambucinumF.javanicumF.javanicum
var.radicicolaF.gibbosumF.graminearumF.culmorumBipolaris
sorokinianaEiio?ieue10,58,42,48,110,96,73,76,17,67,02,33,15,36,23,87,9Ae
?eiio? O, 72% a.?.,

1,0 e/aa8,27,32,410,06,76,36,38,75,66,31,20,05,615,32,57,7O?aci?, 60%
c.i., 1,2
ea/aa13,215,21,24,316,96,81,82,52,19,20,00,06,04,64,110,4Eaio?io?, 70%
n.e.,

1,5 e/aa7,79,93,62,56,86,41,56,47,69,56,31,57,713,34,49,0Eaio?io?, 70%
n.e.,

2,0 e/aa10,28,52,63,214,96,85,72,84,36,40,32,03,316,54,29,5

15

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?en. 1. *enaeuei?noue aaeoa??e a ?ecinoa?? y?iai y/iaith i?ney
canoinoaaiiy aa?a?oeeae?a, % aei eiio?ieth (1994-1996 ??., ni?o ?ieaiae)

?ineei. Aeeiai?ea oiene/ii? ae?? aa?a?oeeae?a o iaoeo aeine?aeao ia
a?oioia? a?eae ?ecinoa?e y?iai y/iaith ia? aeaeyae V-iiae?aii? e?eai?,
i?ney canoinoaaiiy aa?a?oeeae?a /enaeuei?noue /ooeeaeo aeae?a a?ea?a
??cei ciaiooaaeanue ? i?ioyaii 60-oe aei?a iinooiiai a?aeiiaethaaeanue.
Iai?eeeaae, /a?ac 5 aei?a o aa??aio?, aea canoiniaoaaee aa?a?oeeae
o?aci?, 60% c.i. (1,2 ea/aa) ia 40% ciaioeeanue /enaeuei?noue a?ea?a
?iaeo Fusarium, ia 30-oo aeiao /enaeuei?noue a?ea?a oeueiai ?iaeo
iinooiaeany eiio?ieuei?e o na?aaeiueiio ia 15%, ia 60-oo aeiao – aoea ia
??ai? eiio?ieueii? (?en. 2). Noiniaii aaaeeeaiai c aa?iiii?/ii? oi/ee
ci?o ?iaeo Trichoderma, oi canoinoaaiiy aa?a?oeeae?a o oe?eiio ni?eyei
eiai ?icaeoeo. Iaea?eueoa (aei 90%) c?inoaiiy a?ea?a ?iaeo Trichoderma
a?aei?/aii o aa??aio?, aea canoiniaoaaee o?aci?, 60% c.i. *a?ac 30 aei?a
i?ney canoinoaaiiy aa?a?oeeae?a /enaeuei?noue a?ea?a ?iaeo Trichoderma a
na?aaeiueiio c?inea a 3-6 ?ac?a a?aeiinii eiio?ieth. Ia 60-oo aeiao
aieea aa?a?oeeae?a ia a?eae oeueiai ?iaeo iineaaethaaany ? o?eueee o
aa??aio? c o?aci?ii /enaeuei?noue ia?aaeuoaaea eiio?ieueio o 5 ?ac?a.
I?ney canoinoaaiiy aa?a?oeeae?a oaeiae a?aei?/aeinue c?inoaiiy
/enaeueiino? a?ea?a ?iaeo Penicillium aei e?ioey aaaaoaoe?? y?iai
y/iaith, i?e/iio iaeneioi ?inoo ca?a?no?iaaii ia 30-oo aeiao i?ney
iai?eneoaaiiy. Oaeei /eiii, ciaioaiiy ?icaeoeo
ooca??icii-aaeuei?ioinii??icii? ei?aiaai? aiee? o aa??aio?, aea
canoiniaoaaee o?aci?, iiaeeeai, iiynith?oueny cia/iei niaaeii
/enaeueiino? ooca???a ? ??ceei c?inoaiiyi /enaeueiino? a?ea?a ?iaeo
Trichoderma i Penicillium.

4.3. Ci?ie o aeaeiaiio neeaae? a?ea?a o ?ecinoa?? y?iai y/iaith i?ney
canoinoaaiiy aa?a?oeeae?a

Iaaiaeeoueny ia?ae?e 36 aeae?a a?ea?a, ye? aoee aeae?eai? oa
?aeaioeo?eiaai? c ?ecinoa?iiai a?oioo i?ney canoinoaaiiy aa?a?oeeae?a.
?acoeueoaoe niinoa?aaeaiue ca ci?iaie o aeaeia?e no?oeoo?? anioe?aoe??
a?oioiaeo a?ea?a na?ae/aoue, ui iane?aeee canoinoaaiiy ianoeoeeae?a
oa?aeoa?ecothoueny aieiaiei /eiii aeiaaeaiiyi /ooeeaeo aeae?a oa
ciaioaiiyi /anoioe ?cieyoe?? ?ioeo. O ?ecinoa?? y?iai y/iaith,
ia?iaeaiiai aa?a?oeeaeaie, ooai?th?oueny ye?nii iiaa anioe?aoe?y
i?eioei?e c iecueeei ?iaeaenii aeaeiaiai ??ciiiai?ooy. Aeii?ioth/a
iieiaeaiiy caeiaee 1-2 aeaee, i?e/iio c ieeiii /ano a?aeaoaa?oueny ci?ia
aeii?ioth/eo aeae?a. Iai?eeeaae, canoinoaaiiy o?aci?o, 60% c.i.
i?ecaiaeeoue aei iiaiiai aeiaaeaiiy ?c i?eioei?e Absidia glauca,
Acremonium strictum, Cladosporium macrocarpum, Cuninghamella echinulata,
Fusarium gibbosum, Penicillium purpurogenum, P.expansum, P.stipitatum,
P. spinolosum, P.canescens, P.viridicatum. A?aei?/aii oaeiae ciaioaiiy
iaeaea a 2 ?ace /anoioe ?cieyoe?? Fusarium solani, F.oxysporum.
Aeii?ioth/a iieiaeaiiy caeiaee Alternaria alternata, P.citrinum, P.
nigricans, Trichoderma album, Tr. viride.

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0800000008000000010001000000000020000000c40e0000c40e00000200000002000000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?en. 2. *enaeuei?noue a?eaieo ca/aoe?a a ?ecinoa?? y?iai y/iaith i?ney
canoinoaaiiy aa?a?oeeae?a, % aei eiio?ieth (1994-1996 ??., ni?o ?ieaiae)

4.4. Ioe?iea ooia?noaoe/ii? ae?? aa?a?oeeae?a ia ie?ai? aeaee a?ea?a

a oiiaao in vitro

Iaaiaeyoueny ?acoeueoaoe aea/aiiy ooia?noaoe/ii? ae?? aa?a?oeeae?a ia
ie?ai? aeaee a?ea?a a oiiaao in vitro. Iiaeaeueieie eoeueoo?aie aoee
aeaee a?ea?a ?iae?a Fusarium, Penicillium, Trichoderma. Iieacaii, ui a
oiiaao in vitro aa?a?oeeaee a eiioeaio?aoe?yo 0,0001-1,0% ia
i?eai?/oaaee i?i?inoaiiy nii? a?ea?a ?iaeo Fusarium – iniiaieo
caoaeiee?a ei?aiaai? aiee? y?iai y/iaith. Iaa?oue i?e eiioeaio?aoe?? 10
ie/e i?i?inoaiiy nii? a?ea?a Fusarium avenaceum, F. oxysporum, F.
solani, F. sambucinum, F. graminearum aoei iaeaea iaeiaeiaei c
eiio?ieueiei aa??aioii. Aeeeth/aiiy anoaiiaeaii i?e aeei?enoaii?
o?aci?o, 60% c.i. i?e eiioeaio?aoe?? 0,0001%. Niinoa??aaeanue
ooia?noaoe/ia ae?y i?aia?aoo a?aeiinii F. javanicum. Ca?eueoaiiy
eiioeaio?aoe?? aa?a?oeeaeo i?ecaiaeeei aei i?aeaeuaiiy ooia?noaoe/ii?
ae?? i?aia?aoo (18,4-74,6%), a eiioeaio?aoe?? 1,0% niinoa??aaeanue iiaia
a?aenooi?noue i?i?ineeo nii? F. javanicum (oaae. 3). A oie aea /an, i?ae
ae??th aa?a?oeeae?a niinoa??aaeinue noeioethaaiiy i?i?inoaiiy nii?
a?ea?a Penicillium oa Trichoderma.

Oaaeeoey 3 – Aieea ??cieo eiioeaio?aoe?e aa?a?oeeae?a ia eoeueoo?e
a?ea?a a oiiaao in vitro

Iacaa i?aia?aooEiioeaio?aoe?y ae.?., %E?euee?noue i?i?ineeo nii?,
%F.avenaceumF.oxysporumF.solaniF.sambucinumF.graminearumF.javanicumPenic
illium spp.Trichoderma
spp.Eiio?ieue—69,080,488,766,453,285,182,167,2Ae?eiio? O, 72% a.?.
1,065,478,987,165,052,983,283,470,40,168,179,287,765,653,084,086,170,20,
0169,079,588,065,953,184,987,975,10,00168,780,188,366,053,185,187,375,20
,000169,280,688,566,053,385,092,270,2O?aci?, 60%
c.i.1,067,778,786,965,952,2087,286,90,167,472,987,266,052,712,590,287,10
,0168,479,287,766,053,023,095,485,10,00169,179,788,066,353,166,796,480,3
0,000169,080,088,466,753,080,297,878,2Eaio?io?, 70%
n.e.1,067,978,887,166,052,083,083,272,10,168,179,087,566,252,383,585,875
,30,0168,479,487,966,352,784,288,975,60,00168,680,188,268,053,085,090,17
3,10,000169,480,283,666,353,085,090,969,45. AOAEOEAI?NOUe IIAEO
AA?A?OeEAe?A O IIN?AAO ss?IAI ss*IAITH

5.1. *ooeea?noue ie?aieo aeae?a ao?’yi?a aei ianoeoeeae?a

O i?aeaeine?aeiiio aa?ioeaiic? y/i?iiiai iiey ia?aaaaeaee aeaiaeieuei?
ao?’yie. *ooeea?noue ie?aieo aeae?a ao?’yi?a aei aa?a?oeeae?a aoea
??ciith. O?aci?, 60% c.i. (1,2 ea/aa) ? eaio?io?, 70% n.e. (2,0 e/aa)
iaee ?aeaioe/io aoaeoeai?noue ii a?aeiioaiith aei eiaiaee a?ei?, ue?eoe?
cae/aeii?, ?aaeueee aeeei?, aa?acee iieueiai?, i?ioa eaio?io?, 70% n.e.
(2,0 e/aa) iinooiaany o?aci?o ii ae?? ia inio iieueiaee oa ?iiaoeo
iaiaoo/o. Noiniaii eaio?io?o, 70% n.e. (1,5 e/aa), oi eiai a?ieia?/ia
aoaeoeai?noue ia ie?ai? aeaee aeaiaeieueieo ao?’yi?a aoea iaeaea ia
iaeiiio ??ai? c ae?eiio?ii O, 72% a.?. (1,0 e/aa).

5.2. Aieea aa?a?oeeae?a ia caaaeueio caao?’yiai?noue iin?a?a y?iai
y/iaith

O ?acoeueoao? nenoaiaoe/ieo niinoa?aaeaiue ca ?inoii ? ?icaeoeii
eoeueoo?ieo ?ineei ? ao?’yi?a aeyaeaii, ui ia aeine?aeieo ae?eyieao,
ia?iaeaieo aa?a?oeeaeaie, ao?’yie aoee neaaeeie ? a?aenoaaaee a
?icaeoeo, a oie /an ye ia eiio?ieueieo ae?eyieao aiie ?inee a oae?e
e?eueeino?, ui caaeoooaaee y?ee y/i?iue. ?acoeueoaoe aeine?aeaeaiue
na?ae/aoue, ui a?ieia?/ia aoaeoeai?noue aa?a?oeeae?a a?ae??ciy?oueny ii
?ieao. O na?aaeiueiio iaeaeuo aoaeoeai?noue (96%) i?ioe eiiieaeno
aeaiaeieueieo ao?’yi?a o iin?aao y?iai y/iaith i?iyaeee o?aci?, 60% c.i.
(1,2 ea/aa) ? eaio?io?, 70% n.e. (2,0 e/aa). Ae?eiio? O, 72% a.?. (1,0
e/aa) ? eaio?io?, 70% n.e. (1,5 e/aa) i?iyaeee a?ieia?/io aoaeoeai?noue
i?ioe ao?’yi?a ia ??ai? 92,6 ? 93,0% a?aeiia?aeii. Canoinoaaiiy
aa?a?oeeae?a caaacia/eei nooo?ao i?eaaaeo a?iaeath. Iaeaeu? iieacieee
o?iaeaeiino? a?aei?/aii o aa??aioao, aea canoiniaoaaee o?aci?, 60% c.i.
(1,2 ea/aa) ? eaio?io?, 70% n.e. (2,0 e/aa). Na?aaeiy i?eaaaea noaiiaeea
6,4 oe/aa ? 5,7 oe/aa a?aeiia?aeii.

5.3. Aieea aa?a?oeeae?a ia no?oeoo?o a?iaeath oa iin?ai? yeino? y?iai
y/iaith

Canoinoaaiiy aa?a?oeeaeieo i?aia?ao?a i?ecaiaeeoue aei i?aeaeuaiiy iane
eieino ia 18,3%. Ca ?aooiie ca?eueoaiiy iane ca?ai o eiein? a
na?aaeiueiio ia 0,124 a, aeiiaiai?noue eieino c?inea a na?aaeiueiio ia
10%. Ca?ii c aa??aio?a, aea canoiniaoaaee o?i?/iee caoeno a?ae ao?’yi?a,
oa?aeoa?ecoaaeinue aeuith iaoo?ith, a?eueoith ieoiiith aaaith ca?ai
e?oiii? o?aeoe??, a?aeiia?aeaei aeiiaai noaiaea?oo aeey ieaiaa?iiai
y/iaith ca ai?noii a?eea.

?acoeueoaoe ae?iaie/i? ia?aa??ee a Ee?anuee?e iaeano?, ENI
“Aa?eo?anueea” na?ae/aoue, ui canoinoaaiiy aa?a?oeeaeo o?aci?, 60% c.i.
(1,2 ea/aa) i?ecaiaeeei aei ciaioaiiy e?eueeino? aeaiaeieueieo ao?’yi?a,
i?e oeueiio a?iaeae ca?ia noaiiaea – 37 oe/aa, cao?aoe – 24,42 a?i./aa,
aa?o?noue caa?aaeaiiai a?iaeath – 124,8 a?i./aa, /enoee aeioiae – 99,7
a?i./aa.

AENIIAEE

1. A?coaeueieie niinoa?aaeaiiyie ca o?aaeaieie ?ineeiaie ? iiaeaeueoei
aeae?eaiiyi caoaeiee?a o /eno? eoeueoo?e anoaiiaeaii, ui a oiiaao
i?ai?/iiai E?ninoaio Oe?a?ie iin?ae y?iai y/iaith ui??/ii o?aaeothoueny
ooca??icii-aaeuei?ioinii??iciith oa oea?einii?aeueiciith ei?aiaaeie
aieeyie.

2. Canoinoaaiiy aa?a?oeeae?a a iniiaiiio ia aieeaaei ia neiioiie,
?iciianthaeaeai?noue oa ?icaeoie ei?aiaaeo aieeae y?iai y/iaith, ca
aeeeth/aiiyi o?aci?o, 60% c.i. I?ney aeei?enoaiiy oeueiai aa?a?oeeaeo
niinoa??aaeinue nooo?aa ciaioaiiy ?icaeoeo
ooca??icii-aaeuei?ioinii??icii? oa oea?einii?aeueicii? ei?aiaaeo aieeae
ii??aiyii c eiio?ieai: o oaco o?oaeoaaiiy – ia 27% ? 44,8%, o oaco
iiei/ii? noeaeino? – ia 18,8% oa 32,8% a?aeiia?aeii.

3. Niaae ?icaeoeo ei?aiaaeo aieeae i?ney canoinoaaiiy o?aci?o, 60% c.i.,
iaoiiaeth?oueny oaeeie oaeoi?aie:

a) ciaioaiiy ieoiii? aaae ie?aieo aeae?a caoaeiee?a
ooca??icii-aaeuei?ioinii??icii? ei?aiaai? aiee?, cie?aia Fusarium
avenaceum ia 50,0%, F. avenaceum var. herbarum – ia 46,9%, F.solani – ia
51,3%, F. solani var. redolense – ia 72,3%, F. javanicum oa F. javanicum
var. radicicola – ia 100%;

a) cia/ia ciaioaiiy a ?ecinoa?? y?iai y/iaith e?eueeino? a?ea?a ?iaeo
Fusarium: ia 40% /a?ac 5 aei?a oa ia 15% /a?ac 30 aei?a;

a) ??cea c?inoaiiy /enaeueiino? a?ea?a ?iaeo Trichoderma: iaeaea a 2
?ace /a?ac 5 aei?a, o 6 ?ac?a /a?ac 30 aei?a, o 5 ?ac?a /a?ac 60 aei?a
i?ney iai?eneoaaiiy aa?a?oeeaeii;

a) iiaia aai /anoeiaa aeiaaeaiiy ?c i?eioei?e ?ecinoa?e y?iai y/iaith
aeayeeo aeae?a a?ea?a, iniaeeai oaeeo ye Fusarium gibbosum, Absidia
glauca, Cladosporium macrocarpum, Cuninghamella echinulata, Penicillium
purpurogenum, P. expansum, P. spinolosum, P. canescens, P. viridicatum,
Fusarium solani, F. oxysporum;

ae) nooo?aa ca?eueoaiiy /anoioe ?cieyoe?? a?ea?a Penicillium citrinum,
P. nigricans, Trichoderma album, Tr. viride.

4. Aa?a?oeeae o?aci?, 60% c.i. oa?aeoa?eco?oueny iayai?noth
ooia?noaoe/ii? ae??. O aeine?aeao in vitro a?i i?eai?/oaaa ??no nii?
Fusarium javanicum ia 18,4-100% caeaaeii a?ae eiioeaio?aoe??. A oie aea
/an i?ae ae??th aa?a?oeeae?a niinoa??aaeinue noeioethaaiiy i?i?inoaiiy
a?ea?a Penicillium oa Trichoderma.

5. Aa?a?oeeaee coiiaethaaee iacia/ia ciaioaiiy /enaeueiino? (ia 10-20%)
aaeoa??e ?iae?a Pseudomonas, Xanthomonas, Bacillus oa i?o?eo?eaoi??a,
yea a?aeiiaethaaeanue aei eiio?ieueiiai ??aiy /a?ac 60 aei?a ? ia iaea
nooo?aiai aieeao ia ?icaeoie ei?aiaai? aiee? y?iai y/iaith.

6. Aeine?aeaeai? aa?a?oeeaee iathoue aenieo aoaeoeai?noue i?ioe
eiiieaeno aeaiaeieueieo ao?’yi?a o iin?aao y?iai y/iaith. ?o a?ieia?/ia
aoaeoeai?noue neeaaea?: o?aci?, 60% c.i. (1,2 ea/aa) ? eaio?io?, 70%
n.e. (2,0 e/aa) – 96%, ae?eiio? O, 72% a.?. (1,0 e/aa) – 92,6%,
eaio?io?, 70% n.e. (1,5 e/aa) – 93,0%.

7. Canoinoaaiiy aa?a?oeeae?a i?ecaaei aei i?aeaeuaiiy i?iaeoeoeaii?
eouenoino? y?iai y/iaith ia 1,1 oo./i2, iane eieina – ia 18,3%, iane
ca?ai o eiein? – ia 0,124 a, aeiiaiaiino? eieino – ia 10%, ui
caaacia/eei nooo?ao i?eaaaeo a?iaeath. Iaeaeu? iieacieee o?iaeaeiino?
io?eiaii i?e canoinoaaii? o?aci?o, 60% c.i. (1,2 ea/aa) ? eaio?io?o, 70%
n.e. (2,0 e/aa). Na?aaeiy i?eaaaea neeaaeaea 6,4 oe/aa ? 5,7 oe/aa
a?aeiia?aeii.

8. Ca?ii, a?ae?a?aia c aa??aio?a, aea cae?enithaaee o?i?/iee caoeno a?ae
ao?’yi?a, oa?aeoa?eco?oueny aeuith iaoo?ith (a na?aaeiueiio ia 10 a/e),
a?eueoith ieoiiith aaaith ca?ia e?oiii? o?aeoe??, a?aeiia?aea? aeiiaai
noaiaea?oo ca ai?noii a?eea (8,8-9,2%).

Nienie ?ia?o, iioae?eiaaieo ca iaoa??aeaie aeena?oaoe??

1. Eenaiei N.A., Aeaeai I.A. Aa?a?oeeaee a iin?aao y?iai y/iaith
//Caoeno ?ineei. – 1996. — ? 2. – N. 6-7. (Iniaenoee aianie caeiaoaa/a
80%, i?iaaaeaiiy aenia?eiaio?a oa iae?e?a).

2. Aeaeai I.A. Aa?a?oeeaee ? i?e?ioei?a //Caoeno ?ineei. – 1998. – ? 2.
– N. 7-8.

3. Aeaeai I.A. O?aci? – oea ye?noue //Caoeno ?ineei. – 1998. — ? 7. – N.
7.

4. ?aoueiai N.A., Aeaeai I.A., Ai?aa/iaa I.I. Ia?aaeiin?aia ia?iaea
ian?iiy //Caoeno ?ineei. – 1999. — ? 1. – N. 4-5. (Iniaenoee aianie
caeiaoaa/a 50%, i?iaaaeaiiy aenia?eiaio?a oa iae?e?a).

5. Aeaeai I.A. Aieea aa?a?oeeae?a ia aeaeiaee neeaae i?e?ioei?e a
?ecinoa?? y?iai y/iaith //Oace aeiiia?aeae i?aeia?iaeii?
iaoeiai-i?aeoe/ii? eiioa?aioe?? iieiaeeo a/aieo ? niaoe?ae?no?a “Oeyoe
?aoe?iiaeueiiai aeei?enoaiiy caiaeueieo ?ano?n?a Oe?a?ie”. – 1995. – N.
106.

6. Noi?/ion ?.I., Aeaeai I.A., Aeaeai I.A. Aoaeoeai?noue iiaeo
ia?niaeoeaieo aa?a?oeeae?a o iin?aao y?iai y/iaith //Oace aeiiia?aeae
i?aeia?iaeii? iaoeiai-i?aeoe/ii? eiioa?aioe?? iieiaeeo a/aieo ?
niaoe?ae?no?a “Oeyoe ?aoe?iiaeueiiai aeei?enoaiiy caiaeueieo ?ano?n?a
Oe?a?ie”. – 1995. – N. 143. (Iniaenoee aianie caeiaoaa/a – 65%,
i?iaaaeaiiy aenia?eiaio?a oa iae?e?a).

7. Aeaeai I.A. Aieea aa?a?oeeae?a ia aeaeiaee neeaae caoaeiee?a
cae/aeii? ei?aiaai? aiee? y?iai y/iaith //Oace aeiiia?aeae
iaoeiai-i?aeoe/ii? eiioa?aioe?? iieiaeeo a/aieo ? niaoe?ae?no?a,
i?enay/aii? 50-??//th ?inoeoooo caoenoo ?ineei “I?iaeaie caoenoo ?ineei
a?ae oe?aeeeaeo i?aai?ci?a o no/anieo aeiiii?/ieo oa aeieia?/ieo
oiiaao”. – 1996. – N. 123-124.

Aeaeai I.A. ?icaeoie ei?aiaaeo aieeae y?iai y/iaith i?ney canoinoaaiiy
aa?a?oeeae?a o i?ai?/iiio E?ninoaio Oe?a?ie. – ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
n?euenueeiainiiaea?nueeeo iaoe ca niaoe?aeuei?noth 06.01.11 –
o?oiiaoieia?y. – Iaoe?iiaeueiee aa?a?iee oi?aa?neoao, Ee?a, 1999.

Aea/aii ?icaeoie oa aeaeiaee neeaae ei?aiaaeo aieeae y?iai y/iaith i?ney
canoinoaaiiy aa?a?oeeae?a. Aia?oa a oiiaao Oe?a?ie aea/aia i?e?ioei?a
?ecinoa?e y?iai y/iaith oa aeeiai?ea ?? /enaeueiino?, ooi/iaiee aeaeiaee
neeaae i?eioei?e a?oioo i?ney canoinoaaiiy aa?a?oeeae?a. Aecia/aii
a?ieia?/io aoaeoeai?noue iiaeo aa?a?oeeae?a i?ioe aeaiaeieueieo ao?’yi?a
o iin?aao y?iai y/iaith, ui aeaei iiaeeea?noue ca?a?no?oaaoe ?o ? aianoe
a “Ia?ae?e ianoeoeeae?a ? aa?io?i?eao?a, aeicaieaieo aei aeei?enoaiiy a
Oe?a?i?”.

Eeth/ia? neiaa: y?ee y/i?iue, ei?aiaa? aiee?, aa?a?oeeaee, i?e?ioei?a,
aaeoa???, a?eae.

Aeaeai I.A. ?acaeoea ei?iaauo aieeae y?iaiai y/iaiy iinea eniieueciaaiey
aa?aeoeeaeia a naaa?iie Eaninoaie Oe?aeiu. – ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa
naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe 06.01.11 – oeoiiaoieiaey.
— Iaoeeiiaeueiue aa?a?iue oieaa?neoao, Eeaa, 1999.

Eco/aii ?acaeoea, ?ani?ino?aiaiea e neiioiiiaaiac ei?iaauo aieeae
y?iaiai y/iaiy i?e eniieueciaaiee aa?aeoeeaeia. Onoaiiaeaii, /oi
?anoaiey y?iaiai y/iaiy a oneiaeyo naaa?iie Eaninoaie Oe?aeiu
ii?aaeathony ooca?eicii-aaeueieioinii?eiciie, oea?einii?aeeaciie
ei?iaauie aieeyie. Eniieueciaaiea aa?aeoeeaeia ia aeeyao ia neiioiiu
i?iyaeaiey aieacie. A oeaeii aa?aeoeeaeu ia aeeyee ia ?ani?ino?aiaiea e
?acaeoea yoiai caaieaaaiey, ca eneeth/aieai aa?eaioa, aaea eniieueciaaee
aa?aeoeeae o?aci? 60% n.i. A oacu o?oaeiaaiey, iiei/iie niaeinoe
ioia/aii nieaeaiea ?acaeoey ooca?eicii-aaeueieioinii?eiciie e
oea?einii?aeeaciie ei?iaauo aieeae.

Aeieiae/aneei aiaeecii ii?aaeaiiuo ?anoaiee onoaiiaeaii, /oi a oneiaeyo
naaa?iie Eaninoaie Oe?aeiu aeaaeue anoiaeia e iiao?aiea ei?iae y?iaiai
y/iaiy aucuaatho a?eau eeanna ianiaa?oaiiuo ii?yaeea Hyphomycetales
naiaenoaa Tuberculariaceae. Iiaeaaeythuaa aieueoeinoai iaoiaaiia
i?aaenoaaeaii aeaeaie ?iaea Fusarium, eioi?ua ioiinyony e naeoeeyi
Roseum, Discolor, Elegans, Martiella. Niioonoaothueie a?eaaie ?iaea
Fusarium auee Bipolaris, Rhizopus, Alternaria.

Onoaiiaeaii aeeyiea aa?aeoeeaeia ia aeaeiaie ninoaa aicaoaeeoaeae
ooca?eicii-aaeueieioinii?eiciie ei?iaaie aieee y?iaiai y/iaiy. Iiae
aeaenoaeai aa?aeoeeaeia a ?ecinoa?a y?iaiai y/iaiy iaaethaeaeinue
nieaeaiea /eneaiiinoe aaeoa?ee ?iaea Pseudomonas, Xanthomonas, Bacillus,
ieo?eoeeaoi?ia ia i?ioyaeaiee 30-oe aeiae iinea eniieueciaaiey
aa?aeoeeaeia, a iinoaiaiiia ainnoaiiaeaiea i?ienoiaeeo ia i?ioyaeaiee
aeaoo ianyoeaa.

I?e eco/aiee ?ecinoa?iie ieeioei?u y?iaiai y/iaiy onoaiiaeaii, /oi
aeeiaieea oiene/aneiai aeaenoaey aa?aeoeeaeia ia ii/aaiiua a?eau eiaao
aeae V-iiaeiaiie e?eaie. Iinea eniieueciaaiey aa?aeoeeaeia /eneaiiinoue
/oanoaeoaeueiuo aeaeia a?eaia ?acei oiaiueoaeanue e ia i?ioyaeaiee 60-oe
aeiae iinoaiaiii ainnoaiaaeeaaeanue. I?e eniieueciaaiee aa?aeoeeaea
o?aci?, 60% n.i. i?ienoiaeeei nieaeaiea /eneaiiinoe a?eaia ?iaea
Fusarium e ?aceia oaaee/aiea a?eaia ?iaea Trichoderma e Penicillium.

A ?ecinoa?a y?iaiai y/iaiy ia?acoaony ea/anoaaiii iiaia niiauanoai
ieeioei?u n ieceei eiaeaenii aeaeiaiai ?aciiia?acey. Aeiieie?othuaa
iieiaeaiea caieiaee 1-2 aeaea. Ni a?aiaiai aeiieie?othuea aeaeu
eciaiyeenue.

Onoaiiaeaii, /oi a oneiaeyo in vitro aa?aeoeeaeu a eiioeaio?aoeeyo
0,0001-1,0% ia oaiaoaee i?i?anoaiey nii? a?eaia ?iaea Fusarium –
iniiaiuo aicaoaeeoaeae ei?iaauo aieeae y?iaiai y/iaiy. Eneeth/aiea
iaia?oaeaii i?e eniieueciaaiee o?aci?a, 60% n.i. i?e eiioeaio?aoeee
0,0001%. Ioia/aii ooiaenoaoe/aneia aeaenoaea i?aia?aoa ioiineoaeueii
Fusarium javanicum. Oaaee/aiea eiioeaio?aoeee oneeeaaei ooiaenoaoe/aneea
naienoaa o?aci?a, 60% n.i. Iiae aeaenoaeai aa?aeoeeaeia i?ienoiaeeei
noeioee?iaaiea i?i?anoaiey nii? a?eaia Penicillium e Trichoderma.

Eco/aii aeeyiea aa?aeoeeaeia ia iinaaiua e oaoiieiae/aneea ea/anoaa
naiyi y?iaiai y/iaiy, ii?aaeaeaia aeieiae/aneay yooaeoeaiinoue
aa?aeoeeaeia i?ioea aeaoaeieueiuo ni?iyeia a iinaaao y?iaiai y/iaiy.

Eeth/aaua neiaa: y?iaie y/iaiue, ei?iaaua aieee, aa?aeoeeaeu,
iee?ioei?a, aaeoa?ee, a?eau.

Djam O.V. Development of spring barley root rots after herbicide
application in northern Forest-Steppe of Ukraine. – Manuscript.

The thesis for a candidates degree of agriculture sciences, the
speciality 06.01.11 – phytopathology. National Agrarian University,
Kyiv, 1999.

The development and species composition of spring barley root rots at
use of herbicides were investigated. For the first time in Ukraine
mycoflora of rhyzosphere of spring barley and its dynamics were
investigated, species composition of soil microflora after herbicide
application was improved. Biological efficiency of new herbicides
against broad-leaved weeds in spring barley was evaluated, what made it
possible to register them and bring to the “List of pesticides and
agrochemicals, allowed for use in Ukraine”.

Key words: spring barley, root rots, herbicides, microflora, bacteria,
fungi.

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