IAOeIIIAEUeIA AEAAeAIIss IAOE OE?A?IE

IINOEOOO OICIIEIAI? ?INEEI I AAIAOEEE

Nieieianueea Ienaia Apeaipiaia

OAeE 581.132:632.112:633.11

?AAEOeIss OIOINEIOACO I OIOIAeEOAIIss O NI?OIA IOAIEOeI

IA A?OIOIAO IINOOO

03.00.12 — oiciieiaiy pineei

Aaoipaoapao aeenapoaoei ia caeiaoooy iaoeiaiai

nooiaiy eaiaeeaeaoa aiieiai/ieo iaoe

Ee?a — 1999

. Aeenapoaoei?th ? poeiien

?iaioa aeeiiaia a Iinoeoooi oiciieiai? pineei i aaiaoeee IAI Oepa?ie

Iaoeiaee eapiaiee — aeieoip aiieiai/ieo iaoe, ipioanip

Aoey?a Aipen Iaaiiae/

Iinoeooo oiciieiai? pineei i aaiaoeee

IAI Oepa?ie, caa. aiaeaeieii

Ioioeieii iiiiaioe — aeieoip aiieiai/ieo iaoe, ipioanip

Ei/oaae Naioeaia Ieoaeeiaia

Iinoeooo oiciieiai? pineei i aaiaoeee

IAI Oepa?ie, caa. aiaeaeieii

eaiaeeaeao aiieiai/ieo iaoe

?iaeei Iaai Ieei/

Iinoeooo eeioeiii? aiieiai?

i aaiaoe/ii? iiaeaiapi? IAI Oepa?ie,

no. iaoe. niiapiaioiee

Ipiaiaeia onoaiiaa — Oeaiopaeueiee aioaii/iee naae ii. I.I. Apeoea

IAI Oepa?ie.

Caoeno aiaeaoaeaoueny   » 1 » eeiiy 1999 p.i 10 aiaeeii ia
caniaeaiii

niaoeiaeiciaaii? a/aii? paaee Ae 26.212.01 ipe Iinoeoooi oiciieiai?

pineei i aaiaoeee IAI Oepa?ie ca aaepanith:252022, Ee?a 22, aoe.

Aaneeueeianueea 31/17

C aeenapoaoei?th iiaeia iciaeiieoenue a aiaeiioaoei Iinoeoooo
oicii-

eiai? pineei i aaiaoeee IAI Oepa?ie, Ee?a, aoe. Aaneeueeianueea 31/17

Aaoipaoapao picineaiee «29» opaaiy 1999 p.

A/aiee naepaoap

niaoeiaeiciaaii? a/aii? paaee Opooaiia A.A.

AEOOAEUeIINOUe I?IAEAIE

Ipiaeaia aea/aiiy oiciieiai/ieo iaoaiiciia iinooinoieeinoi
pineei ipeaapoa? cia/io oaaao /apac oa, ui iinooa nooo?ai iaiaaeo?
iioaioeiae ipiaeoeoeaiinoi eoeueoopieo pineei ia aaeeeeo ieiuao caiii?
eoei. Ainopith ? oey ipiaeaia oaeiae aeey Oepa?ie, cia/ia /anoeia
nieuenueeiainiiaeapnueeeo oaiaeue yei? aiaeiineoueny aei ciie
iaaeinoaoiueiai aieiaicaaacia/aiiy [Oiaoueei, Apeaipthe; 1992,
Ioni?iei,1985].

Ineieueee 90-95% ipaaii/ii? iane pineei noaipth?oueny a ipioeani
oioineioaco, iaeii?th c aieiaieo ipe/ei cieaeaiiy ?o ipiaeoeoeaiinoi a
oiiaao iinooe ? aaeueioaaiiy oioineioaco [Boyer,1986; Kaiser,1987],
yeee iaeaieueo oaeaeei paaao? ia ianoa/o aiaee. Anoaiiaeaii, ui iaa?oue
iacia/ia ciaioaiiy aieiaino? a?oioo, ua aei iiyae iii?oieo ci?i
iaiaeiaiinoi eenoeia,aeeeeea? ipeaii/aiiy oioineioaoe/iiai canai?iiy
aoaeaeeneiai aaco [Oap/aanueeee;1964, Gollan, Passioura, Munns;1986].
?ia?aoaaiiy oioineioaco ca iii??ii? oa iao?eaaei? ae?? iinooe
iaoiiaeth?oueny, iapo ca ana, /anoeiaei caepeooyi ipiaeeoia. O?eaaea ?
a?eueo aeeaiea ciaaiaeiaiiy oioineioacoth/eo oeaiei i?ecaiaeeoue aei
nooo?aeo ciii oioineioaoe/iiai iaoaaie?cio, iai?iaeeoiaiai e?i?ooaaiiy
oioineioaco [Chaves;1992]. Aenia?eiaioaeueii anoaiiaeaii cieaeaiiy o
oeeo oiiaao aeoeaiino? 1 i 2 oioinenoai oa oioioinoi?eethaaiiy
[Sharkey, Badger; 1982], ciaioaiiy aeoeaiinoi oa ai?noo ?AOE [Allen,
Bowes;1987].

Aoaeoeaiinoue oioineioaoe/ii? aneiieyoei? aoaeaoeth o N3-pineei,
iipiaiyii c N4-pineeiaie, cia/ii cieaeaia aianeiaeie ipioeano
oioiaeeoaiiy [Zelitch, 1979]. Aeaeieaiiy aoaeaeeneioe i ?? aopaoe a
ipioeani oioiaeeoaiiy aeinyaa? 50 % aiae aeaeeiiai oioineioaco eenoea.
Oi/a ieoaiiy ipi oiciieiai/io pieue oeueiai ipioeano ua aei eiioey ia
c`yniaaia, iaeiae a eioapaoopi aeneiaeaii ipeiouaiiy, ui
aeieieaoiiio iaoaaieicio iaeaaeeoue aaaeeeaa pieue a paaoeyoei?
oioineioaoe/ii? aneiieyoei? NI2 ca nopaniaeo oiia [Lorimer;1978]. Oea,
ia iaoo aeoieo, ? aeooaeueiei ieoaiiyi, yea iiopaao? aieueo aeeaieiai
aea/aiiy.

Ca’ycie piaioe c iaoeiaeie ipiapaiaie, ieaiaie, oaiaie

?iaioo aeeiiaii a paieao oaiaoeee aiaeaeieo oiciieiai? oa
aeieiai? oioineioaco Iinoeoooo oiciieiai? pineei i aaiaoeee «Aea/aiiy
paaoeyoei? oioineioaco, aeeoaiiy, oioiaeeoaiiy oa piciiaeieo aneiieyoia
a aeiiipii-aeoeaioipiie nenoaii pineei o ca`yceo c ipiaeoeoeaiinoth oa
noieeinoth aaiioeiia n.-a. eoeueoop aei aei? nopania», 2.28.6/229.

Iaoa i caaea/i aeineiaeaeaiiy

Aieiaiith iaoith piaioe aoei c’ynoaaoe ?aaeoeith oioineioaco i
oioiaeeoaiiy o picieo ca iinooinoieeinoth nipoia iceii? oa ypi?
ioaieoei ia aeith oa iineyaeith iinooe.

Aoee iinoaaeaii ianooiii caaaeaiiy:

1) aea/eoe aeith apoioiai? iinooe ia oioineioac i oioiaeeoaiiy o
nipoia iceii? ioaieoei ia paiiio aoaiao iioiaaiaco;

2) aeineiaeeoe aieea iiaiaoaaiiy oioiaeeoaiiy, cieaeaiiyi
eiioeaiopaoei? eenith, ia paaeoeith oioineioaoe/iiai aiapaoo nipoia
iceii? ioaieoei ca oiia aenieioaiiapaoopiiai nopano ia ii/aoeo
iioiaaiaco;

3) aea/eoe aeith oa iineyaeith apoioiai? iinooe ia
iioaineaiinoue oioineioaco, oioiaeeoaiiy, iiipe aeeooci? CO2 oa
aoaeoeaiinoue aeeipenoaiiy aiaee o eiiopanoieo ca iinooinoieeinoth
nipoia ypi? oa iceii? ioaieoei a oace aeoiaeo a opoaeo i
eieiniiiy-oeaioiiiy;

4) aea/eoe aieea apoioiai? iinooe ia aeeth/aiiy 14N a
oioineioaoe/iee iaoaaieici picieo ca iinooinoieeinoth nipoia ypi?
ioaieoei o oaco oeaioiiiy.

Iaoeiaa iiaecia iaeapaeaieo pacoeueoaoia

Anoaiiaeaii, ui aieueoa noieeinoue oioineioaoe/iiai aiapaoo aei
aei? eipioei/anii? apoioiai? iinooe o iinooinoieeeo nipoia ioaieoei,
iiae o iaio iinooinoieeeo, iiynith?oueny iaioeie (aiaeiinii oioineioaco)
aeopaoaie aoaeaoeth ia oioiaeeoaiiy i aeuith eenoeiaith ipiaiaeiinoth
CO2.

Aiapoa anoaiiaeaii, ui aipiaeiaae iapinoaiiy aiaeiiai aeaoioeeoo
iioaineaiinoue oioiaeeoaiiy o ni?oia ypi? ioaieoei ciiith?oueny
aeaioacii: o ii/aoeiaee iapiiae iiaeaeuo?oueny, a iioii cieaeo?oueny.
Ipe oeueiio oaca aeoeaaoei? iioaineaiinoi oioiaeeoaiiy o iinooinoieeeo
nipoia ioaieoei ? aieueo opeaaeith, iiae o iaio iinooinoieeeo.
Iiaeaeuaiiy oioiaeeoaiiy ca nopaniaeo oiia iia`ycaii ic cieaeaiiyi
iaaeoiaeaeaiiy aoaeaeeneiai aaco a oeipiieanoe eenoea i iiaeneeaiiyi
ieneaaiacii? paaeoei? ?AOE/I.

O nipoia ioaieoei a nopaniaeo oiiaao oioiaeeoaiiy caoeua?
oioineioaoe/iee aiapao aiae oioiiiaiaoaaiiy, ui ynepaaioa aeyaeaii o
iinooinoieeeo nipoia.

Ipaeoe/ia cia/aiiy iaeapaeaieo pacoeueoaoia

Iieacaii cia/eiinoue oioiaeeoaiiy aeey aaeaioaoei? aei aei?
iinooe o no/anieo iinooinoieeeo nipoia ioaieoei. Iopeiaii pacoeueoaoe
iiaeooue aooe aeeipenoaii a iaoeiai-aeineiaeieo onoaiiaao, aea
ipiaiaeyoueny aeineiaeaeaiiy aaeaioaoei? pineei aei ianipeyoeeaeo oiia,
ciepaia iinooe, a oaeiae aeey ioeiiee naeaeoeieiiai iaoapiaeo ca
nooiaiai ?o iinooinoieeinoi.

Iniaenoee aianie caeiaoaa/a.

Aeenapoaio iniaenoi ipiaiaeicoaaea aiaeiiaiaeio eioapaoopo,
iaieiaeiea iaiaoiaeieie iaoiaeaie aeineiaeaeaiiy, apaea o/anoue o
ieaioaaiii aeineiaeia. ?acoeueoaoe, yei iaaaaeaii a aeenapoaoeieiie
piaioi, iaeapaeaii caeiaoaa/ai naiinoieii.

Aipiaaoeiy pacoeueoaoia aeenapoaoei?.

Iniiaii iaoeiai pacoeueoaoe aeenapoaoeieii? piaioe aeiiiaiaeaeeny
ia oinoie eiioapaioei? iieiaeeo a/aieo, ipenay/aii? 50-e pi/ieoei
canioaaiiy Iinoeoooo oiciieiai? pineei i aaiaoeee IAI Oepa?ie
(Ee?a, 1996); Iiaeiapiaeiie iapaaei «Aeooaeueii ieoaiiy oiciieiai?
pineei a aniaeoi aeieiai/ieo ipiaeai Oepa?ie» (*apiiaoei, 1996), a
oaeiae aoee ipaaenoaaeaii ia 3-io uipi/iiio neiiiceoii
«Oiceei-oiii/ii iniiae oiciieiai? pineei i aiioaoiieiai?» (Iineaa,
1997) i ia 11-io eiiapani ?apiiaenueeiai oiaapenoaa oiciieiaia pineei
(Aapia, 1998).

Ioaeieaoei?.

Ca pacoeueoaoaie piaioe iioaeieiaaii ope noaooi o iaoeiaeo oaoiaeo
aeaeaiiyo oa i’youe oac.

Nopoeoopa i ia’?i piaioe.

Aeenapoaoeiy aeeeaaeaia ia 141 noipiioei iaoeiiieniiai oaenoo, c
ieo 32 noipiiee iethnopaoei? (16 oaaeeoeue, 17 iaethieia) i
neeaaea?oueny ci anooio, iaeyaeo eioapaoope, oapaeoapenoeee ia’?eoia i
iaoiaeia aeineiaeaeaiue, aeniapeiaioaeueii? /anoeie c iaaiaipaiiyi
iaeapaeaieo pacoeueoaoia, aeniiaeia, nieneo eioapaoope, yeee aeeth/a?
249 aeaeapae.

IA’?EOE I IAOIAeE AeINEIAeAEAIUe

O aaaaoaoeieieo aeineiaeao aea/aee nipoe iceii? ioaieoei Iaeanueea
162 (iinooinoieeee nipo) i Iepeaaai (iaio iinooinoieeee) oa nipoe ypi?
ioaieoei Napaoianueea 29 (iinooinoieeee nipo) i Oapeianueea 2 (iaio
iinooinoieeee). O aeineiaeao ia i?i?inoeao aeineiaeaeoaaee nipoe
iceii? ioaieoei- iinooinoieei: Aeiinueea iaiiaeapeeeiaa, Aeueaaopin
iaeanueeee, Neiaie iaeanueeee, Iaeanueea 162, THaieaeia 75 oa
iaio iinooinoieei:Iepeaaai, Capo/, Oeeaaiea, Ethoanoeain 7, Ee?anueea
6.

?ineeie aepiuoaaee o aaaaoaoeieiiio aoaeeii/eo a iinoaeeiao, yei
iinoeee 7 ea aaniethoii nooiai apoioo, oaeiapaiiai N?E. Ipipinoee
aepiuoaaee a ieanoianiaeo iinoaeeiao, yei iinoeee 500 a apoioo i 100 a
iineo, iiaeaeeaeaieo pic/eiii Eiiia. Aieiainoue apoioo iiaeopeioaaee
ia piaii 60% aiae iiaii? aieiai?ieinoi (IA). Iinooo noaipthaaee
iaiaaeaiiyi iieeao pineei aeineiaeiiai aapiaioo, cieaeoth/e aieiainoue
apoioo aei 30% aiae IA.

Aapiaioe aaaaoaoeieieo aeineiaeia:

1) apoioiaa iinooa ipe 30% IA o oace eieiniiiy-oeaioiiiy, yeo
noaipthaaee ipeieiaiiyi iieeao i c ianooiiei eiai aiaeiiaeaiiyi /apac 6
aeia;

2) cieaeaiiy aieiainoi apoioo aei 30% IA c ianooiiei aiaeiiaeaiiyi
iieeao a oace aeoiaeo a opoaeo i eieiniiiy-oeaioiiiy.

Aapiaioe aeineiaeia ia i?i?inoeao :

1) cieaeaiiy aieiainoi apoioo aei 30% IA ipeieiaiiyi iieeao;

2) aeniiceoeiy i?e oaiiapaoopi 40i N i?ioyaii 2,5 aiaeei ipe aeaio
eiioeaiopaoeiyo eenith (21% oa 2%) .

Aeiipthaaiiy CO2-aaciiaiiio pineei ipiaiaeeee ca aeiiiiiaith
iiopa/apaiiiai aaciiaeicaoipa IA-5501, ipe oaiiapaoopi 25i N ?
iioaineaiinoi OA? 400 Ao/i2. Iioaineaiinoue oioiaeeoaiiy ioeiithaaee ca
piaiai aeeeaeo CO2 a iapoi 60 nae iiney aeieiaiiy naioea. Aiaeiee
aeaoioeeo aecia/aee ca noaiaeapoiith iaoiaeeeith. Aeoeaiinoue
aeaeapaieneeo aaiiy — ca iaoiaeeeith Iaaiaapaei?aa [1986],
eiioeaiopaoeith ipieiio — ca Aaeonii [1973], oioineioaoe/iee iaoaaieici
c aeeipenoaiiyi 14NI2 ca iaoiaeeeith Aicianainueeiai [1965], aiino
oeipioieo — ca Apiiiii [1949]. Noaoenoe/iee aiaeic
aeniapeiaioaeueieo aeaieo ipiaiaeeany ca Ieioiinueeei [1980].

AeIss I IINEssAeIss A?OIOIAI? IINOOE, AENIEIOAIIA?AOO?IIAI NO?ANO
IA OIOINEIOAC I OIOIAeEOAIIss NI?OIA ICEII? IOAIEOeI

Ca iaeapaeaieie aeaieie o aeineiaeao c ipipinoeaie aeaio picieo
ca iinooinoieeinoth apoi nipoia iceii? ioaieoei anoaiiaeaii,ui
cieaeaiiy aieiainoi apoioo aei 30% IA ipecaaei aei nooo?aiai
caieueoaiiy aiaeiiai aeaoioeeoo eenoeia ipipinoeia ye o iinooinoieeeo,
oae i o iaio iinooinoieeeo nipoia: o iapoeo aii cpin o napaaeiueiio a
2,4 pace, a o aepoaeo — 3,0 pace (oaae.1).

Iioaineaiinoue aeaeeiiai oioineioaco a napaaeiueiio ii apoiao
nipoia iiae aei?th iinooe iiaiao?oueny iaeiaeiai (ia 61%). Aea a apoii
iinooinoieeeo nipoia aneiieyoeiy CO2 iaeiaioa ipeaii/oaaeany o nipoo
Iaeanueea 162 i noaiiaeea 81 % aiae eiiopieth. O paooe nipoia oei?? ae
apoie aeaeeiee oioineioac ca oiia apoioiai? iinooe cia/ii ciaioeany i
aeipiaithaaa 19-37 % aiae eiiopieth. O nipoia Capo/ oa Oeeaaiea, yei
iaeaaeaoue aei iaio iinooinoieeeo, aeaeeiee oioineioac cieceany ia 48,7
% i 41,4 % aiae eiiopieth. Oaeei /eiii o oeieiio picaiaeiinoi iiae
aeineiaeaeaieie nipoaie a iaaeao eiaeii? apoie nipoia ca nooiaiai
iiaiaoaaiiy iioaineaiinoi oioineioaco iiae aei?th iinooe iapaepeaathoue
aiaeiiiiinoi ca oeei iapaiaopii iiae nipoaie picieo apoi
iinooinoieeinoi.

Oaaeeoey 1.

Aieea iinooe ia aiaeiee aeaoioeeo, iioaineaiinoue oioineioaco oa
oioiaeeoaiiy, aeoeaiinoue aeaeapaieneeoaaiiy 14NI2- aeieieaoo,
eiioeaiopaoeith oeipioieo o ipipinoeia nipoia iceii? ioaieoei (%
aiae eiiopieth, napaaeii cia/aiiy ca apoiaie nipoia)

Aiaeiee aeaoioeeo Oioineioac Oioiaeeoaiiy Aeoeaiinoue
aea-eapaieneeoaa-iiy Eiioeaiopaoeiy oeipioieo

IINOOINOIEEI NI?OE

241 39,4 169 109 86,9

IAIO IINOOINOIEEI NI?OE

304 38,4 114 77,5 77,6

?acii c oei, cieaeaiiy aiaeiiinoa/aiiy ipecaaei aei iiaeaeuaiiy
oioiaeeoaiiy a onio iinooinoieeeo nipoia, oiaei ye a apoii iaio
iinooinoieeeo nipoia aiii ciaiooaaeiny aai cpinoaei, aea iaioa, iiae
o nipoia iapoi? apoie (oaae. 1). Oae iioaineaiinoue oioiaeeoaiiy
iaeaieueoa cpinea o Aeiinueei? iaiiaeapeeeiai? o 2,7 pace i Aeueaaopina
iaeanueeiai o 2,2 pace aiae eiiopieth. O napaaeiueiio a apoii i’yoe
iinooinoieeeo nipoia oioiaeeoaiiy caieueoeeiny ipioe eiiopieth ia 69 %,
a a apoii i’yoe iaio iinooinoieeeo — ia 14%.

Aeey ioeiiee iioaineaiinoi aeieieaoiiai iaoaaieicio aecia/aee
aeoeaiinoue aeaeapaieneeoaaiiy 14NI2- aeieieaoo. O pineei iaio
iinooinoieeeo nipoia aeoeaiinoue aeaeapaieneeoaaiiy ciaiooaaeany ia
22,5%, a o iinooinoieeeo aiia aeaui caieueooaaeany i noaiiaeea 109 %
aiae eiiopieth. Oea naiae/eoue ipi oa, ui a oiiaao iinooe aeieieaoiee
iaoaaieici ipioieaa o iinooinoieeeo nipoia iioaineaiioa, iiae o
iaio iinooinoieeeo. Oei aeaii ocaiaeaeothoueny c pacoeueoaoaie
aaciiaope/ieo aeiipthaaiue oioiaeeoaiiy. Aianeiaeie naieaeiaiai? iinooe
aiino oeipioieo a eenoeao o iinooinoieeeo nipoia ciaioeany ia 13,1%, a
o iaio iinooinoieeeo ia 22,4% iipiaiyii c eiiopieai.

Aiaeii/a?oueny oaiaeaioeiy aei aca?iica’yceo iiae nooiaiai
iinooinoieeinoi nipoia i ciiiith aiaeiioaiiy iioaineaiinoi
oioiaeeoaiiy aei oioineioaco oa iineeaiiyi aeoeaiinoi aeieieaoiiai
iaoaaieicio a oiiaao iinooe. Iaeiae o iaaeao eiaeii? apoie nipoia
oaeiae ? aeeeth/aiiy. sseui ipiaanoe paiaeoaaiiy nipoia ca
aiaeiioaiiyi niiaaiaeiioaiue iioaineaiinoi oioiaeeoaiiy oa
oioineioaco a aeineiaei i eiiopiei ipeioneath/e, ui iinooinoieeinoue
cpinoa? ic iiaeaeuaiiyi oeueiai niiaaiaeiioaiiy, oi ie, ca nooiaiai
iinooinoieeinoi nipoia,iopeia?ii oaeee pyae: Aeiinueea iaiiaeapeeeiaa >
Aeueaaopin iaeanueeee > Ee?anueea 6 > Capo/ >Iaeanueea 162 > THaieaeia
75 > Neiaie iaeanueeee > Iepeaaai > Ethoanoeain 7 = Oeeaaiea (eopneaii
iiii/aii nipoe c apoie iaio iinooinoieeeo).

I?e aei? aenieioaiiapaoopiiai nopano o i?i?inoeia iaio
iinooinoieeiai nipoo Ie?eaaai iioaineaiinoue oioineioaco i
oioiaeeoaiiy cia/ii cieaeo?oueny, oiaei ye o iinooinoieeiai nipoo
Iaeanueea 162 oioineioac ipeaii/o?oueny iaioa, a iioaineaiinoue
oioiaeeoaiiy ia ciiith?oueny (oaae.2). Ipe aiaenooiinoi oioiaeeoaiiy
aiaeiiiiinoi iiae nipoaie cieeaee (oaae.3). Oei pacoeueoaoe aeathoue
iiaenoaao cpiaeoe aeniiaie ipi oa, ui oioiaeeoaiiy ca oiia aeniei?
oaiiapaoope ? inaioeaiiinoi aeeiio? ooieoeith caoenoo oioineioaoe/iiai
aiapaoo pineei (aieueo aepaaeaio o iinooinoieeiai nipoo), caapiaath/e
eiai aiae oioiiiaiaoaaiiy.

A?oioiaa iinooa o oace eieiniiiy-oeaioiiiy ipecaiaeeea aei
caieueoaiiy eenoeiaiai iiipo aeeooci? o Iaeanueea 162 — a 3,7, a o nipoo
Iepeaaai- a 4,1 pace aiae eiiopieth (oaae.4). Aiaeiii, ui iiaeaeuaiiy
eenoeiaiai iiipo ciaioo? aopaoe aiaee i ? aaaeeeaei /eiieeii
aaeaioaoei? pineei aei aei? iinooe.

Ipe aeinoaoiueiio aieiaicaaacia/aiii aoaeoeaiinoue aeeipenoaiiy
aiaee o nipoo Iaeanueea 162 aoea aeaui aeua, iiae o nipoo Iepeaaai.
Ca oiia iinooe oeae iieaciee cieaeoaaany o nipoo Iaeanueea 162 ia 26%,
a o nipoo Iepeaaai — ia 42% aiae eiiopieth .

Oaaeeoey 2.

NI2- aaciiai?i i?ipinoeia ioaieoei eiiopanoieo ca iinooinoieeinoth
nipoia iiae aei?th aenieioaiiapaoopiiai nopano a aoiinoapiiio iiaiopi.

Aapiaioe

aeineiaeo Iioaineaiinoue aeaeeiiai oioineioaco, ia/a noo.?a/. aiae
Iioaineaiinoue oioiaeeoaiiy, ia/a noo.?a/ aiae Oioiaeeoaiiy aiaeiinii
aeaeeiiai oioineioaco,% Iioaineaiinoue inoeiiiai oi-oineioaco,

ia/a noo?a/.aiae

Iaeanueea 162

aei nopano 84,5 + 7,7 12,0 + 1,1 14,2 96,5 + 8,3

iiney 55,9 + 4,2* 12,0 + 1,2 21,5 67,9 + 5,8*

i/ae 0,66 1,00 1,5 0,70

Iepeaaai

aei nopano 54,2 + 4,6 16,6+1,4 30,6 70,8+6,7

iiney 21,2 + 1,9** 11,3 + 1,0* 53,3 32,5 + 2,8**

i/ae 0,39 0,68 1,74 0,46

Oaaeeoey3.

Aieea aenieioaiiapaoopiiai nopano i?e iecuee?e eiioeaio?aoe?? eenith (2
%) ia ia?aiao?e NI2-aaciiai?io ipipinoeia picieo ca iinooinoieeinoth
nipoia iceii? ioaieoei.

Aapiaioe

aeineiaeo Iioaineaiinoue aeaeeiiai oioineioaco, ia/a noo.?a/. aiae
Iioaineaiinoue oioiaeeoaiiy, ia/a noo.?a/ aiae Oioiaeeoaiiy aiaeiinii
aeaeeiiai oioineioaco,% Iioaineaiinoue inoeiiiai oi-oineioaco,

ia/a noo?a/.aiae

Iaeanueea 162

aei nopano 67,1 + 4,7 14,7 + 1,4 21,9 81,8 + 7,3

iiney 23,2 + 2,2** 8,95 + 0,6* 38,6 32,2 +3,1**

i/ae 0,35 0,61 1,76 0,39

Iepeaaai

aei nopano 75,4 + 5,2 15,1 + 1,4 20,0 90,5 + 8,7

iiney 23,3 + 2,0** 6,63 + 2,8** 28,5 29,9 + 2,8**

i/ae 0,31 0,43 1,43 0,33

I?eiioee aei oaaeeoeue: i/ae — aiaeiioaiiy aaee/ei iiney aei? no?ano aei
aaee/ei aei no?ano;

* — ?icieoey iiae aeineiaeii i eiio?ieai aeinoiai?ia i?e ?>95%;

** — ?icieoey iiae aeineiaeii i eiio?ieai aeinoiai?ia i?e ?>99%.

Oioineioac oeaaiaiai eenoea iiae aieeaii iinooe (oaa.4) neeueiioa
ipeaii/oaaany o nipoo Iepeaaai. Aiaeii/an iioaineaiinoue oioiaeeoaiiy
iinooinoieeiai nipoo Iaeanueea 162 i a aeineiaeiiio, i
eiiopieueiiio aapiaioao aoea iaeiaeiaith, a o nipoo Iepeaaai ipe
iinoni ciaiooaaeany. ?iaaiue oioiaeeoaiiy aiaeiinii aeaeeiiai
oioineioaco cpinoaa o nipoo Iaeanueea 162 a 1,7 pace, a nipoo
Iepeaaai a 3,7 pace ii?iaiyii c eiio?ieai.

Iiney aiaeiiaeaiiy iieeao iioaineaiinoue aeaeeiiai oioineioaco
oeaaiaiai eenoea o nipoo Iaeanueea 162 aiaeiiaeeany iiaiinoth, a nipoo
Iepeaaai — eeoa aei 60% aiae piaiy eiiopieth (oaae.5). O iapoiai c
nipoia ipe oeueiio aiaeaoaaeiny oaeiae ipaeoe/ii iiaia aiaeiiaeaiiy
oioiaeeoaiiy, a o ni?oo Ie?eaaai aei 59% aiae eiio?ieth. Aiaeiioaiiy
oioiaeeoaiiy aei oioineioaco ciaioeeiny aei aaee/ei eiio?ieueieo
pineei iaio ni?oia aaea ia ianooiio aeiao iiney iiiiaeaiiy iieeao. O
ieo aiaeaoaaeiny oaeiae iiaia aiaeiiaeaiiy aaee/ei eenoeiaiai iiipo oa
aoaeoeaiinoi aeeipenoaiiy aiaee.

Oaeei /eiii pacoeueoaoe ipiaaaeaieo aeineiaeaeaiue naiae/aoue, ui
oioineioaoe/iee aiapao pineei nipoo Iepeaaai ? cia/ii /ooeeaioei aei
aei? aiaeiiai nopano, iiae nipoo Iaeanueea 162. Aieueoa noieeinoue
oioineioaco a pineei nipoo ioaieoei Iaeanueea 162, i/aaeaeii, iia’ycaia
c aaeaioaoeieieie ipioeanaie — cpinoaiiyi eiioeaiopaoei? iniioe/ii
aeoeaieo pa/iaei i ipiaeeoiaiai iiipo, ui aeicaiey? iiaeaeueoe
aoaeoeaiinoue iiaeeiaiiy oa aeeipenoaiiy aiaee. Iiaeopeiaiiy aieueoi?
iioaineaiinoi oioineioaco iiae /an aei? nopano i oaeaeea aiaeiiaeaiiy
oioineioaoe/iiai aiapaoo ipe iiiiaeaiii iieeao pineei nipoo Iaeanueea
162 iaoiiaethaaei iaio? ao?aoe ca?iiai? i?iaeoeoeaiino? (oaae.6).

Oaaeeoey 6.

Aieea apoioiai? iinooe ia nopoeoopo apiaeath aieiaiiai iaaiio nipoia
iceii? ioaieoei (Eaini- iieaciee ainiiaeapnueei? aoaeoeaiinoi)

Aapiaioe aeineiaeo Iana, a Eieueeinoue capai, oo Eaini, %

iaaiia capia 1000 capai

IAeANUeEA 162

Eiiopieue 1,24+0,04 I,60+0,03 25,1+0,74 23,7+0,9 47,7+1,1

Aeineiae 1,06+0,05 0,48+0,02 20,8+1,39 22,4+0,6 45,9+0,8

Ae/E 0,86 0,80 0,83 0,95 0,96

IE?EAAAI

Eiiopieue 1,34+0,05 0,59+0,02 22,9+0,92 25,8+0,7 43,7+0,7

Aeineiae 1,08+0,06 0,40+0,03 17,3+1,03 23,1+0,7 37,0+0,8

Ae/E 0,81 0,68 0,76 0,90 0,85

I?eiioea: Ae/E- aiaeiioaiiy aeineiaeo aei eiio?ieth.

*ANIAA AeEIAIIEA I?IAeEOIAI? I?IAIAeIINOI, IIOAINEAIINOAE OIOINEIOACO I
OIOIAeEOAIIss OA OIOINEIOAOE*IEE IAOAAIEICI O EIIO?ANOIEO NI?OIA ss?I?
IOAIEOeI A E?EOE*II OACE ?ICAEOEO ?INEEI I?E CIIIAO AIEIAINOI A?OIOO

Ca iaeapaeaieie aeaieie cieaeaiiy iioaineaiinoi aeaeeiiai
oioineioaco o nipoia ioaieoei ipe aei? iinooe iaoiiaeth?oueny a
iniiaiiio ye cieaeaiiyi ipiaeeoiai? ipiaiaeiinoi, oae i iiaeaeuaiiyi
aiaeiinieo (aeiaeaeeiiai oioineioaco) aopao aoaeaoeth ia
oioiaeeoaiiy (pen.1). Eiaoioei?io ei?aeyoei? iiae oioineioacii i
ipiaeeoiaith ipiaiaeiinoth o aeineiaeaeaieo ni?oia y?i? ioaieoei a
e?eoe/iee ia?iiae ?icaeoeo ?ineei (oace aeoiaeo a opoaeo,
eieiniiiy-oeaioiiiy) aeipiaithaaa 0,87, iiae oioineioacii i aiaeiioaiiyi
oioiaeeoaiiy aei oioineioaco = — 0,88. Oioineioaoe/iee aia?ao o
iinooinoieeiai nipoo Na?aoianueea 29 aoa noieeioei aei cieaeaiiy
aieiainoi apoioo o oace aeoiaeo a o?oaeo oa eieiniiiy — oeaioiiiy i
oaeaeoa aiaeiiaey?oueny iiney iiiiaeaiiy iieeao, iiae o nipoo
Oapeianueea 2 (pen.1, A). Aieueoa eenoeiaa ipiaiaeiinoue o nipoo
Napaoianueea 29 (pen.1, A) caaacia/o? aieueo aenieo oaeaeeinoue
aneiieyoei? NI2 eenoeaie aianeiaeie aeui? eiioeaiopaoei? NI2 a
iiaeeeioeiieeao eenoea. Oiio oeae nipo a ii/aoeiaee iapiiae iinooe iaa
i aeuo aoaeoeaiinoue aeeipenoaiiy aiaee. Iiaeopeiaiiy aieueo aeniei?
eenoeiai? ipiaiaeiinoi o nipoo Napaoianueea 29 a oiiaao iinooe, iiae o
nipoo Oapeianueea 2, iiaeia iiynieoe oei, ui a eeioeiao eenoea nipoo
Napaoianueea 29 ipe aiaeiiio nopani iaapiiaaeaeo?oueny aieueoa
eieueeinoue ipieiio ii?iaiyii c eiio?ieueieie aaee/eiaie. Ciepaia o oaco
aeoiaeo a opoaeo eiioeaiopaoeiy ipieiio caieueoo?oueny ia 7-io aeiao
iinooe o nipoo Napaoianueea 29 a 18 pac, a o nipoo Oapeianueea 2 — a 11
pac aiae eiiopieth. Iioaineaiinoue oioiaeeoaiiy o ni?oia ypi? ioaieoei
ii iipi caieueoaiiy aiaeiiai aeaoioeeoo ciiith?oueny aeaioacii: nii/aoeo
caieueoo?oueny, a iioii ciaioo?oueny (pen.2,A). O iinooinoieeeo nipoia
ioaieoei oaca aeoeaaoei? iioaineaiinoi oioiaeeoaiiy ? aieueo opeaaeith,
iiae o iaio iinooinoieeeo. Aieueoa noieeinoue oioineioaoe/iiai aiapaoo o
iinooinoieeiai nipoo ca oiia iinooe coiiaeaia cpinoaiiyi oioiaeeoaiiy,
yea cieaeo? eiai oioiaeanopoeoeith aiae iaaeeeoeo aiapai?. Oey
aeanoeainoue caaacia/o? noieeinoue nipoo aei aei? aiaeiiai nopano.
Niiaaiaeiioaiiy oioiaeeoaiiy oa oioineioaco c?inoaei, iniaeeai o iaio
iinooinoieeiai ni?oo Oa?eianueea 2 (?en.2,A).

Epii aeiipthaaiue aaciiaiiio eenoeia, iaie ipiaaaeaii oaeiae
aeineiaeaeaiiy ic canoinoaaiiyi ii/aii? aoaeaeeneioe c nipoaie
Napaoianueea 29 i Oapeianueea 2 a oace eieiniiiy-oeaioiiiy. Ipe aei?
apoioiai? iinooe aiaeaoaaeiny ipeaii/aiiy aeoeaiinoi aneiieyoei? NI2,
iniaeeai o aeineiaeieo pineei iaio noieeiai nipoo Oapeianueea 2: o
aeineiaeieo pineei Napaoianueei? 29 cieaeoaaeany ia 6 %, a o
Oapeianueei? 2 ia 66 % aiae eiiopieueieo aapiaioia (oaae.7) .

Iiae aei?th iinooe o iaio nipoia ciaiooaaeiny aeeth/aiiy aoaeaoeth
o epioiaeue. O pineei nipoo Napaoianueea 29 aeeth/aiiy iioee a
epioiaeue aoei ieae/ei, iiae o iaio noieeiai nipoo Oapeianueea 2 i
noaiiaeei aiaeiiaiaeii 0,45 % i 0,76 % aiae caaaeueii? aeoeaiinoi
aiiiaaiaoo. A eenoeao ioaieoei epioiaeue ia ? iniiaiei caianiei
ipiaeoeoii oioineioaco, a iiaeaea?oueny iioaineaiiio

Oaaeeoey 7.

Aieea apoioiai? iinooe ia iioaineaiinoue aneiieyoei? 14NI2 oeaaiaiai
eenoea eiiopanoieo ca iinooinoieeinoth nipoia ypi? ioaieoei.

Iioaineaiinoue aneiieyoei? 14NI2, iei.?iciaaeia oa-1 i-2 n-1

Na?aoianueea 29 Oa?eianueea 2

eiio?ieue aeineiae ae/e eiio?ieue aeineiae ae/e

IINOOA

29,4+2,51 27,6+4,79 0,94 28,4+1,78 12,4+1,15* 0,44

* — ?icieoey iiae aeineiaeii i eiio?ieai aeinoiai?ia i?e ?> 95%

ae/e — aiaeiioaiiy aeineiaeo aei eiio?ieth.

iaiiio, neioaco?oueny i aiaepieico?oueny iaeii/anii. Ca oiia iinooe
aeeth/aiiy iioee a oeoepe o iinooinoieeiai nipoo Napaoianueea 29
cieceeiny ia nooo?ai, a o iaio noieeiai Oapeianueea 2 ia ciiithaaeiny
(oaae. 8). Ca aeaieie I.A. Oap/aanueeiai [1964] iiae aieueieie
oeoepaie a oiiaao a?oioiai? oa aoiinoa?ii? iinooe aiaeaoaa?oueny
iapapiciiaeie paaeiiaeoeaiiai aoaeaoeth:ciaioo?oueny aiino 14N a
naoa?ici i caieueoo?oueny — a aaenicao. Iiaeaeuaiiy aaenic iiaea
i?eaanoe aei caieueoaiiy iniioe/iiai iioaioeiaeo i aei iiyae aieueo
cia/ieo aiaeioo?eioth/eo nee a eeioeiao eenoeia.

Ipe aiaeiiio aeaoioeeoi o iinooinoieeiai nipoo Napaoianueea 29
ciaiooaaeiny iaapiiaaeaeaiiy aoaeaoeth o aiiiieeneioao ia 16 %
aiae eiiopieueiiai aapiaioo. Aiaeii/an o nipoo Oapeianueea 2 ca oiia
iinooe aiaeaoaaeiny iiaeaeuaiiy aneiieyoei? paaeiiaeoeaiiai aoaeaoeth
a aiiiieeneioe ia 34% eiiopieth (oaae.8). Ipe iaiaaeaiii iieeao
aeeth/aiiy 14NI2 a ipaaii/ii eeneioe caieueooaaeiny

Oaaeeoey 8.

Aeeth/aiiy paaeiiaeoeaiiai aoaeaoeth a e?ioiaeue (a % aiae caaaeueii?
paaeiiaeoeaiinoi aiiiaaiaoo), oeoe?e, aiiiieeneioe oa i?aaii/ii
eeneioe (a % aiae paaeiiaeoeaiinoi niepoi-aiaeipic/eiii? opaeoei?)
oeaaiaiai eenoea picieo ca iinooinoieeinoth nipoia ioaieoei ca oiia
iinooe.

Niieoee Na?aoianueea 29 Oa?eianueea 2

eiio?ieue Aeineiae ae/e eiio?ieue aeineiae ae/e

epioiaeue 1,87 0,85* 0,45 1,98 1,51* 0,76

oeoepe 39,8 35,8 0,90 44,5 46,6 1,05

Aiiiieen-eioe 34,7 29,3 0,84 19,2 25,7* 1,34

ipaaii/ii eeneioe 9,36 9,85 1,05 6,64 14,8* 2,23

* — ?icieoey iiae aeineiaeii i eiio?ieai aeinoiai?ia i?e ?> 95%

ae/e — aiaeiioaiiy aeineiaeo aei eiio?ieth.

a 2,2 pace o iaio noieeiai nipoo Oapeianueea 2, oiaei ye o nipoo
Napaoianueea 29 — caeeoaeiny ia piaii eiiopieueieo pineei. A oiiaao
iinooe aiaeaoaaeinue cieaeaiiy aiapaaoe/ieo aeaiaaeaioia. Aianeiaeie
oeueiai caieueooaaany aiino 14N a ipaaii/ieo eeneioao i iia’ycaieo c
ieie aiiiieeneioao, ui naiae/eoue ipi oopoaeiaiiy ipioieaiiy paaeoei?
oinoiaeioeapeiiaa eeneioa — oinoiaeioeapeiiaee aeueaeaaiae i
iiaeaeuaiiy eiiaipiinoi caeieniaiiy eiieopaioieo paaeoeie
oinoiaeioeapeiiaa eeneioa — oinoiaiieiipoaao c iiaeaeueoei ooaipaiiyi
ipaaii/ieo eeneio oa aiiiieeneio.

Oaeei /eiii, iaiaaeaia aiaeiiinoa/aiiy o iaio noieeiai nipoo ypi?
ioaieoei neeueiioa iiaiao? aeoeaiinoue aneiieyoei? 14NI2, iiaeaeuo?
aeeth/aiiy 14N a aiiiieeneioe i ipaaii/ii eeneioe.

AENIIAEE

1. Cieaeaiiy iioaineaiinoi aeaeeiiai oioineioaco o nipoia iceii?
oa ypi? ioaieoei ca aei? iinooe iaoiiaeth?oueny ye cieaeaiiyi
ipiaeeoiai? ipiaiaeiinoi, oae i iiaeaeuaiiyi aiaeiinieo (aei
oioineioaco) aopao aoaeaoeth ia oioiaeeoaiiy.

2. Aieueoa noieeinoue oioineioaoe/iiai aiapaoo aei aei?
eipioei/anii? apoioiai? iinooe o iinooinoieeeo nipoia iipiaiyii c iaio
iinooinoieeeie iaoiiaeaia aeuith ipiaeeoiaith ipiaiaeiinoth i
iaioeie (aiaeiinii aei oioineioaco) aeopaoaie aoaeaoeth ia oioiaeeoaiiy.

3.Iioaineaiinoue oioiaeeoaiiy o ni?oia ypi? ioaieoei aipiaeiaae
iapinoaiiy aiaeiiai aeaoioeeoo ciiithaaeany aeaioacii: nii/aoeo
caieueooaaeany, a iioii ciaiooaaeany. O iinooinoieeeo nipoia ioaieoei
oaca aeoeaaoei? iioaineaiinoi oioiaeeoaiiy aoea aieueo opeaaeith, iiae o
iaio iinooinoieeeo.

4. Iinooinoieei nipoe iceii? oa ypi? ioaieoei a oiiaao iaiaaeaiiai
aiaeiiinoa/aiiy iaee aieueoo aoaeoeaiinoue aeeipenoaiiy aiaee, iiae
iaio iinooinoieei.

5. O eenoeao iinooinoieeiai nipoo Napaoianueea 29 a oiiaao iinooe
aiaeaoaaeiny aieueoa iaapiiaaeaeaiiy aiiiieeneioe ipieiio a ii?iaiyiii c
eiio?ieai, iiae o nipoo Oapeianueea 2, ui ipecaiaeeei aei iiaeaeuaiiy
iniioe/iiai iioaioeiaeo eeioei.

6. Iaiaaeaia aiaeiiinoa/aiiy o nipoo Oapeianueea 2, ia aiaeiiio
aiae nipoo Napaoianueea 29, neeueiioa ipeaii/oaaei oioineioac,
iiaeaeuoaaei aeeth/aiiy 14N a aiiiieeneioe i ipaaii/ii eeneioe .

7. Ca aei? aenieioaiiapaoopiiai nopano o iaio iinooinoieeiai
nipoo oioineioac i oioiaeeoaiiy ipeaii/oaaany aieueoa, iiae o
iinooinoieeiai nipoo, a ipe aeeeth/aiii oioiaeeoaiiy oei aiaeiiiiinoi
iiae nipoaie cieeaee, ui naiae/eoue ipi pieue oioiaeeoaiiy o caoenoi
oioineioaoe/iiai aiapaoo aiae aei? iinooe i aeniei? oaiapaoope.

ONIIAII ?ACOEUeOAOE AeENA?OAOeI? IIOAEIEIAAII A ?IAIOAO

1.Nieieianueea I.A., Noanee I.I. Aieea apoioiai? iinooe ia
NI2-aaciiaiii, aeaeapaieneeoaaiiy, aiino oeipioieo oa pino o nipoia
iceii? ioaieoei picieo ca iinooinoieeinoth // Aeooaeueii ipiaeaie
oiciieiai? aiaeiiai paaeeio oa iinooinoieeinoi pineei: Ca. iaoe. ip.,
ipenay/. iai’yoi ae-pa aiie. iaoe, ipio. Oiaoueea I.A. / E.:
OIA»Iiaeiap. oii. aaaioeiy», 1997. — N. 83-85.

2. Nieieianueea I.A., Noanee I.I., Aoey?a A.I. ?aaeoeiy
oioineioaoe/iiai aiapaoo aeaio picieo ca iinooinoieeinoth nipoia iceii?
ioaieoei aei aei? apoioiai? iinooe //Oeceieiaey e aeioeiey
eoeueoo?iuo ?anoaiee. — 1998. — 30, N 4. — N.241-246.

3.Ei?iaaa Ae.TH., Noanee I.I., Nieieianeay I.A. Iniaaiiinoe eiaeoeoeee
oethipanoeaioeee oeipioeeea eenoueaa ioaieoeu a oneiaeyo canooe
//Oeceieiaey e aeioeiey eoeueoo?iuo ?anoaiee. — 1998. — 30, N 3. —
N.170-174.

4. Nieieianueea I.A., Noanee I.I. Aieea iinooe ia aiaeiee paaeei,
aneiieyoeith NI2 i oioiaeeoaiiy o picieo ca iinooinoieeinoth nipoia
iceii? ioaieoei // Aeooaeueii ieoaiiy oiciieiai? pineei a aniaeoi
aeieiai/ieo ipiaeai Oepa?ie: Oace aeiiiaiaeae iiaeiapiaeii? iapaaee
(*apiiaoei,1995 p.) / Ee?a,A.a., 1995. -N.53.

5. Nieieianueea I.A., Noanee I.I. Aea/aiiy aca?iica’yceo oioineioaco
oa oioiaeeoaiiy o picieo ca iinooinoieeinoth nipoia iceii? ioaieoei ca
oiia iinooe // Aeooaeueii ipiaeaie oiciieiai? pineei i aaiaoeee:Oace
aeiiiaiaeae VI Eiioapaioei? iieiaeeo a/aieo, ipenay/aii? 50-e pi/ieoei c
aeiy canioaaiiy Iinoeoooo oiciieiai? pineei i aaiaoeee IAI Oepa?ie (9-11
aeiaoiy, 1996 p., Ee?a) / Ee?a, A.a.,1996.- N.20-21.

6. Nieieianueea I.A., Aoey?a A.I. Aieea iinooe ia oioineioac,
aiaeiee paaeei i ipiaeoeoeaiinoue o picieo ca iinooinoieeinoth nipoia
iceii? ioaieoei // Aeooaeueii ipiaeaie oiciieiai? pineei i
aaiaoeee:Oace aeiiiaiaeae VI Eiioapaioei? iieiaeeo a/aieo, ipenay/aii?
50-e pi/ieoei c aeiy canioaaiiy Iinoeoooo oiciieiai? pineei i aaiaoeee
IAI Oepa?ie (9-11 aeiaoiy, 1996 p., Ee?a) / Ee?a,A.a.,1996.- N.23.

7. Nieieianeay I.A.,Noanee I.I. Aeeyiea canooe ia acaeiinaycue
oioineioaca e oioiaeuoaiey o ?acee/iuo ii canooionoie/eainoe ni?oia
iceiie ioaieoeu // Oeceei-oeie/aneea iniiau oeceieiaee ?anoaiee e
aeioaoiieiaey: Oacenu 3-ai aaeaaiaeiiai neiiiceoia (27-28 ethiy 1997
a., Iineaa) / I.:A.e.,1997.- N.89.

8. Sokolovska O.G., Kiriziy D.A., Gulyaev B.I. Effect of drought on
photosynthesis and photorespiration of spring wheat // Bulgarian Journal
of Plant Physiology. Spesial issue. 11 Congress of FESPP 7-11
September 1998. — 1998. -P. 161.

Nieieianueea I.A. ?aaeoeiy oioineioaco i oioiaeeoaiiy o nipoia
ioaieoei ia apoioiao iinooo. — ?oeiien.

Aeenapoaoeiy ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
aiieiai/ieo ca niaoeiaeueiinoth 03.00.12 — oiciieiaiy pineei. -Iinoeooo
oiciieiai? pineei i aaiaoeee IAI Oepa?ie, Ee?a, 1999.

Aeineiaeaeaii ?aaeoeith oioineioaco i oioiaeeoaiiy o nipoia iceii?
oa ypi? ioaieoei picii? iinooinoieeinoi ia aeith oa iineyaeith
iinooe o picii iapiiaee iioiaaiaco. Iieacaii, ui aieueoa noieeinoue
oioineioaoe/iiai aiapaoo aei aei? eipioei/anii? apoioiai? iinooe
o iinooinoieeeo nipoia ioaieoei, iiae o iaio iinooinoieeeo,
iiynith?oueny iaioeie (aiaeiinii oioineioaco) aeopaoaie aoaeaoeth ia
oioiaeeoaiiy i aeuith eenoeiaith ipiaiaeiinoth aeeooci? NI2.
Iioaineaiinoue oioiaeeoaiiy o ni?oia ypi? ioaieoei o ii/aoeiaee iapiiae
iinooe iiaeaeuoaaeany, a iioii cieaeoaaeany, ipe oeueiio oaca ??
aeoeaaoei? o iinooinoieeeo nipoia ioaieoei ? aieueo opeaaeith, iiae o
iaio iinooinoieeeo. Ipe aiaenooiinoi oioiaeeoaiiy (2%I2) aiaeiiiiinoi
iiae picieie ca iinooinoieeinoth nipoaie iceii? ioaieoei ca paaeoei?th
NI2-aaciiaiiio ia aeith nopano cieeaee. Oea naiae/eoue ipi oa, ui
oioiaeeoaiiy aeeiio? pieue caoenoo oioineioaoe/iiai aiapaoo o ioaieoei,
aieueo aepaaeaio o iinooinoieeeo nipoia.

Eeth/iai neiaa: Triticum aestivum (L.), oioineioac, oioiaeeoaiiy,
apoioiaa iinooa, ipipinoee, epeoe/ii oace picaeoeo.

Nieieianeay I.A. ?aaeoeey oioineioaca e oioiaeuoaiey o ni?oia
ioaieoeu ia ii/aaiioth canooo.- ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.12 — oeceieiaey ?anoaiee. — Einoeooo
oeceieiaee ?anoaiee e aaiaoeee IAI Oe?aeiu, Eeaa, 1999.

Enneaaeiaaee ?aaeoeeth oioineioaca e oioiaeuoaiey o ?acee/iuo ii
canooionoie/eainoe ni?oia iceiie e y?iaie ioaieoeu a noaaeee ipipinoeia
e epeoe/aneea iapeiaeu ?acaeoey ?anoaiee ia aeaenoaea e?aoeia?aiaiiie
ii/aaiiie canooe, a oaeaea ia aunieioaiia?aoo?iue no?ann o
i?i?inoeia. Ia i?i?inoeao eco/aee aeaa a?oiiu, ioee/athueony ii
canooionoie/eainoe, ni?oia iceiie ioaieoeu: aieaa onoie/eaua ni?oa
Aeiineay iieoea?eeeiaay, Aeueaaopin iaeanneee, Neiaie iaeanneee,
Iaeanneay 162, THaeeaeiay 75 e iaiaa onoie/eaua ni?oa Iepeaaai, Capo/,
Oeuaaiea, Ethoanoeain 7, Eeaaneay 6. A aaaaoaoeeiiiuo iiuoao eco/aee
nipoa iceiie ioaieoeu Iaeanneay 162 e Iepeaaai e nipoa ypiaie ioaieoeu
Napaoianeay 29 e Oapeianeay 2.

Ii iiuoao n i?i?inoeaie iieacaii, /oi aeey canooionoie/eauo ni?oia
oa?aeoa?ii a oeaeii aieaa aunieia niioiioaiea eioaineaiinoe oioiaeuoaiey
e oioineioaca i?e oneeaiee aeoeaiinoe aeeeieaoiiai iaoaaieecia, oioy
a eaaeaeie a?oiia ni?oia eiaeenue eneeth/aiey ec yoiai i?aaeea.
Nieaeaiea eioaineaiinoe oioineioaca ni?oia ioaieoeu i?e canooa
iaoneiaeeaaeinue a iniiaiii nieaeaieai onouee/iie i?iaiaeeiinoe (OI) e
iiauoaieai ioiineoaeueiuo (e oioineioaco) iioa?ue oaea?iaea i?e
oioiaeuoaiee. Eiyooeoeeaio ei??aeyoeee iaaeaeo oioineioacii e OI o
eco/aiiuo a aaaaoaoeeiiiuo iiuoao ni?oia y?iaie ioaieoeu a e?eoe/aneee
ia?eiae ?acaeoey ?anoaiee ninoaaeye 0,87, iaaeaeo oioineioacii e
ioiioaieai oioiaeuoaiey e oioineioaco -0,88. Oioineioaoe/aneee aiia?ao
o canooionoie/eaiai ni?oa y?iaie ioaieoeu Na?aoianeay 29 a yoio ia?eiae
i?iyaeye aieueooth onoie/eainoue e nieaeaieth aeaae a ii/aa e auno?aa
ainnoaiaaeeaae naith aeoeaiinoue iinea aiciaiiaeaiey iieeaa, /ai o
iaiaa canooionoie/eaiai ni?oa Oa?ueeianeay 2. Aieueoay OI eenoueaa o
yoiai ni?oa i?e canooa iauyniyaony aieueoei iaeiieaieai a eeaoeao
oeaiae eenoa i?ieeia eae iniioeea, iaania/eaathuaai iiaeaea?aeaiea
aieaa aunieiai iniioe/aneiai iioaioeeaea eenoueaa. Oae a oaca auoiaea a
o?oaeo ia naaeueiua nooee iinea i?ae?auaiey iieeaa eiioeaio?aoeey
i?ieeia oaaee/eaaeanue o Na?aoianeie 29 a 18 ?ac, a o Oa?eianeie 2 —
oieueei a 11 ?ac i?ioea eiio?iey. Ai a?aiaiiie aeeiaieea
eioaineaiinoue oioiaeuoaiey o ni?oia y?iaie ioaieoeu iinea i?ae?auaiey
iieeaa ?anoaiee eciaiyeanue aeaoooacii: aia/aea i?ienoiaeeei aa
iiauoaiea, a caoai nieaeaiea. O canooionoie/eaiai nipoa Napaoianeay 29
oaca aeoeaaoeee eioaineaiinoe oioiaeuoaiey auea i?iaeieaeeoaeueiaa,
/ai o iaiaa canooionoie/eaiai ni?oa Oa?ueeianeay 2.

Ii aeaiiui iiuoia n ?aaeeiaeoeaiie iaoeie i?e canooa o ni?oa
Oa?ueeianeay 2 a ioee/ea io Na?aoianeie 29 neeueiaa nieaeaeanue
anneieeyoeey 14NI2, iiauoaeinue aeeth/aiea 14N a aieiieeneiou e
i?aaie/aneea eeneiou, /oi naycaii n caiaaeeaieai nei?inoe ?aaeoeee
oinoiaeeoea?eiiaay eeneioa —oinoiaeeoea?eiiaue aeueaeaaeae
e,ii-aeaeeiiio, n oneeaieai ?aaeoeee oinoiaeeoea?eiiaay eeneioa —
oinoiaiieie?oaao n aeaeueiaeoei ia?aciaaieai i?aaie/aneeo eeneio e
aieiieeneio aneaaenoaea aeaoeoeeoa yia?aaoe/aneeo yeaeaaeaioia.

O i?i?inoeia iaiaa canooionoie/eauo ni?oia iceiie ioaieoeu a
oneiaeyo canooe aeoeaiinoue aeaea?aieneee?iaaiey 14N-aeeeieaoa
oiaiueoaeanue ia 23%, a o canooionoie/eauo oaaee/eaaeanue ia 9% io
eiio?iey.Yoi naeaeaoaeuenoaoao i oii, /oi i?e canooa aeeeieaoiue
iaoaaieeci i?ioaeaao eioaineaiaa o canooionoie/eauo ni?oia, /ai o iaiaa
onoie/eauo, /oi niaeanoaony n ?acoeueoaoaie aaciiao?e/aneeo ecia?aiee
eioaineaiinoe oioiaeuoaiey.

Iinea yeniiceoeee i?i?inoeia a oneiaeyo aunieioaiia?aoo?iiai
no?anna (40iN) a aoiinoa?a aicaeooa (21%I2) ia ipioyaeaiee 2,5 /aniao
iaiaa canooionoie/eaiai ni?oa iceiie ioaieoeu Ie?eaaai eioaineaiinoue
oioineioaca e oioiaeuoaiey cia/eoaeueii nieaeaeenue, oiaaea eae o
canooionoie/eaiai Iaeanneay 162 oioineioac iaaeae iaiueoa, a
eioaineaiinoue oioiaeuoaiey ia eciaiyeanue. Iinea yeniiceoeee i?e yoie
aea oaiia?aoo?a, ii i?e ionoonoaee oioiaeuoaiey (2% I2) yoe ioee/ey
iaaeaeo ni?oaie ionoonoaiaaee. Ia iniiaa yoe aeaiiuo naeaeai auaiae i
oii, /oi oioiaeeoaiea i?e aunieioaia?aoo?iii no?anna ea?aao ?ieue caueou
oioineioaoe/aneiai aiia?aoa io oioieiaeae?iaaiey.

Eeth/aaua neiaa: Triticum aestivum (L.), ni?oa, oioineioac,
oioiaeuoaiea, ii/aaiiay canooa, ipipinoee, epeoe/aneea oacu ?acaeoey.

Sokolovska O.G. Photosynthesis and photorespiration response in
wheat cultivars on soil drought. — Manuscript.

Thesis for Ph.D.(Biology) by speciality 03.00.12 — Plant
Physiology. — Institute of Plant Physiology and Genetics, National
Academy of Sciences of Ukraine, Kyiv, 1999.

Photosynthesis and photorespiration response in winter and
spring wheat cultivars of different drought resistance on soil
drought in different ontogenesis periods as well as on high
temperature stress in seedlings has been investigated. It is shown
that the higher resistance of photosynthetical apparatus to
short-term drought in drought-resistant wheat cultivars occurs both due
to the lower carbon loss (in respect to visible photosynthesis) on
photorespiration and to the higher leaf conductance of NI2
diffusion comparing with less drought-resistant ones.The intensity of
photorespiration in spring wheat cultivars during the initial period
of drought increased, then it decreased. In this case the stage of
its activation in drought resistant wheat cultivars was more durable
than in less drought resistant cultivars. When photorespiration
was absent (2% I2) the distincts between cultivars different by their
drought resistance winter wheat cultivars by CO2 gas exchange
response on the stress smoothed. This indicates that the
photorespiration defends wheat photosynthetical apparatus from
stresses more expressively in drought-resistant cultivars.

Key words: Triticum aestivum (L.), cultivars, photosynthesis,
photorespiration, soil drought, seedlings, critical stage of
development.

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