OE?A?INUeEA AEAAeAI?ss AA?A?IEO IAOE

?INOEOOO ?INEEIIEOeOAA ?i. A.ss.TH?(?AA

Eaiioiae/ Aaea??e Iao?iae/

OAeE 577.121:(631.531.01:633.11

EI?AIAA? AECIIAOAAIE?OE I?I?INOATH*IAI

IAN?IIss IOAIEOe?, ?O IAeA?AEAIIss, AeINE?AeAEAIIss OA

AEEI?ENOAIIss A ?INEEIIEOeOA?

06.01.09 — ?ineeiieoeoai

A a o i ? a o a ? a o

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia a Oa?e?anueeiio aea?aeaaiiio aa?a?iiio oi?aa?neoao?
?i. A.A. Aeieo/a?aa

I?i?noa?noaa aa?ii?iieneiaiai eiiieaeno
Oe?a?ie

iaoe aeui? oeiee,

eaiaeeaeao
n?euenueeiainiiaea?nueeeo iaoe, i?ioani?

Iaoiia Aa?iai Oaaei?iae/

— aeieoi? a?ieia?/ieo iaoe,
noa?oee iaoeiaee ni?a?ia?oiee

Aiaeeia Aiaoie?e ?aaiiae/,
IAe? a?ieia?? i?e OAeO,

caa. a?aeae?eii iieaeoey?ii?
a?ieia?? oa a?ioaoiieia??

Io?oe?ei? iiiiaioe: -aeieoi? a?ieia?/ieo iaoe, i?ioani?,
Caneoaeaiee ?ia?oiee n?euenueeiai

ainiiaea?noaa Oe?a?ie
Oeeaoeueei Aieiaeeie? Na?a?eiae/, ?inoeooo

?ineeiieoeoaa ?i. A.ss. TH?(?aa
OAAI, aaaeo/ee iaoeiaee

ni?a?ia?oiee

-eaiaeeaeao
n?euenueeiainiiaea?nueeeo iaoe, noa?oee iaoeiaee

ni?a?ia?oiee Iieeiaiee ?a?na
Ieeei?aia, ?inoeooo

oaa?eiieoeoaa OAAI, a/aiee
nae?aoa?

I?ia?aeia onoaiiaa: Oeaio?aeueiee aioai?/iee naae ?i. I.I. A?eoea
IAI Oe?a?ie, i. Ee?a

Caoeno a?aeaoaeaoueny “ 23 “ aa?aciy 1999 ?. i 13 aiaeei? ia
can?aeaii?

niaoe?ae?ciaaii? a/aii? ?aaee Ae. 64.366.01. a ?inoeooo?
?ineeiieoeoaa ?i. A.ss. TH?(?aa,

310060, i. Oa?e?a, Iineianueeee i?iniaeo, 142, oae. 92-23-78

C aeena?oaoe??th iiaeia iciaeiieoeny a a?ae?ioaoe? ?inoeoooo
?ineeiieoeoaa ?i. A.ss. TH?(?aa,

i. Oa?e?a, Iineianueeee i?iniaeo, 142

Aaoi?aoa?ao ?ic?neaiee “ 17 “ ethoiai 1999 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee
Aiiaea?aiei E.A.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aaaeeeaei caaaeaiiyi no/anii? n?euenueeiainiiaea?nueei? iaoee ?
?ic?iaea niinia?a iaea?aeaiiy a?ieia?/ii aeoeaieo ?a/iaei oa ?o
aeei?enoaiiy a i?aeoeoe? ?ineeiieoeoaa. Ei?aiaa? aeae?eaiiy — aaaeeeaa
aeaea?aei iaea?aeaiiy a?ieia?/ii aeoeaieo ?a/iaei. Aea/aiiy
a?ieia?/ieo aeanoeainoae ei?aiaaeo aeae?eaiue i?i?inoath/iai ian?iiy
iieueiaeo eoeueoo? ? aaaeeeaith oa ia?niaeoeaiith ae?eyieith
aeine?aeaeaiue.

Aeooaeuei?noue oaie. Aeey aeine?aeiee?a iniaeeai oe?eaaeie ?
ieoaiiy niaoeeo?/iino? oa iie?ooieoe?iiaeueiino? ae?? oeeo ?a/iaei.
Aei ?o neeaaeo aoiaeyoue eaeoeie, i?aai?/i? eeneioe, o?iiie, oaiiee,
ai?iieeneioe oa ?io? a?ieia?/ii aeoeai? ?a/iaeie ( Putman A. et al.,
1983; Rice E., 1985; A?iaec?inueeee A.I. oa ?i., 1987 ).

Aaea iaeiie/eeanue aaeeea e?euee?noue ?ia?o, ia i?aenoaa? yeeo
iiaeeeai neeanoe oe?iea oyaeaiiy i?i a?ieia?/ii aeoeai? aeae?eaiiy (
aeciiaoaaie?oe) i?i?inoath/iai ian?iiy. I?ioa, ?acoeueoaoe
aeine?aeaeaiue o?aaiaioa?i? oa /anii iinyoue noia?a/eeaee oa?aeoa?.
Eiiieaeniee i?aeo?ae aei aea/aiiy neeaaeo, aa?aaaoiiai noaio,
a?ieia?/ieo ooieoe?e ei?aiaaeo aeae?eaiue i?i?inoath/iai ian?iiy
aeicaiey? aeyaeoe ?o neeaaei?noue ? iie?ooieoe?iiaeuei?noue ae??,
?ic?iaeoe iniiae i?aeoe/iiai aeei?enoaiiy ?o o ?ineeiieoeoa?.

Ca(ycie ?iaioe c iaoeiaeie i?ia?aiaie. Aeine?aeaeaiiy ii
aeena?oaoe?? aeeiiai? a ?aieao oaiaoe/iiai ieaio Oa?e?anueeiai
aea?aeaaiiai aa?a?iiai oi?aa?neoaoo ?i.A.A. Aeieo/a?aa ca
oaiith:»?ic?iaea oai?aoe/ieo iniia oa i?aeoe/ieo iaoiae?a iaea?aeaiiy
a?ieia?/ii aeoeaieo ?a/iaei noeioethth/i? oa ?aaoethth/i? ae?? ?c
i?i?inoath/iai ian?iiy — aeiii?a», ? aea?aeaaii? ?a?no?aoe??
194U023437, a oaeiae a ?aieao aea?aeaaii? iaoeiai — oaoi?/ii? i?ia?aie
«A?ii?aia?aoe eiiieaenii? ae?? ye aeueoa?iaoeaa o?i?/iei caniaai a
AIE».

Iaoa ? caaea/? aeine?aeaeaiiy. Iaoith aeine?aeaeaiiy aoei aea/aiiy
a?ieia?/ieo aeanoeainoae, neeaaeo oa aeoeaiino? ei?aiaaeo
aeciiaoaaie?o?a i?i?inoath/iai ian?iiy ioaieoe?, ?ic?iaea

niiniao ?o iaea?aeaiiy oa aeei?enoaiiy a ?ineeiieoeoa?.

A caaea/? aeine?aeaeaiiy aoiaeeei:

1.Aeine?aeeoe iniaeeaino? aeae?eueii? nenoaie i?i?inoe?a ioaieoe?;

2.Aecia/eoe e?i?aeiee neeaae ei?aiaaeo aeciiaoaaie?o?a;

3.?ic?iaeoe oaoiieia?th iaea?aeaiiy a?ieia?/ii aeoeaieo ei?aiaaeo

aeciiaoaaie?o?a i?i?anoath/iai ian?iiy ioaieoe?;

4.Aeine?aeeoe aieea aeciiaoaaie?o?a ia ??ci? ?ineeii? ia(?eoe;

5.Aeine?aeeoe o?oiioeeaeio oa aioeaaeoa??aeueio aeoeai?noue ei?aiaaeo
aeciiaoaaie?o?a;

6.Aea/eoe iie?ooieoe?iiaeuei?noue ae?? ei?aiaaeo aeciiaoaaie?o?a;

7.Ioe?ieoe ia?niaeoeai?noue oa aoaeoeai?noue aeei?enoaiiy
ei?aiaaeo aeciiaoaaie?o?a a ?ineeiieoeoa?.

iaeaia oaoiieia?y iaea?aeaiiy nooi? ?a/iaeie Aeaeinoei ?c
aeciiaoaaie?o?a i?i?inoath/iai ian?iiy iceii? ioaieoe? ( iaoaio
18419(UA) Oe?a?ie, iaoaio 2095956(RUS) ?in?? ). Anoaiiaeaii ye?niee oa
e?euee?niee e?i?aeiee neeaae ei?ai?a oa ei?aiaaeo aeciiaoaaie?o?a
i?i?inoe?a ioaieoe?. Anoaiiaeaii aioeaaeoa??aeueiee aoaeo i/euaieo
e?i?ae?a ei?aiaaeo aeciiaoaaie?o?a. Ia ??cieo a?ieia?/ieo ia(?eoao
iieacaia iie?ooieoe?iiaeuei?noue ?o ae??. Ia iniia? aeei?enoaiiy nooi?
?a/iaeie Aeaeinoei noai?aii aoaeoeaia iiaeeaia na?aaeiaeua aeey
i?i?iuoaaiiy ieeeo eoeueoo?ieo ?ineei (iaoaio 9982 (UA) Oe?a?ie ).
?ic?iaeaiee oa ai?iaaaeaeaiee o i?aeoeeo ?ineeiieoeoaa niin?a aeey
ia?aaeiin?aii? ia?iaee ian?iiy ca?iiaiaiaeo eoeueoo?

( iaoaio 8741 (UA) Oe?a?ie ). Iieacaia ia?niaeoeai?noue aeei?enoaiiy
ei?aiaaeo aeciiaoaaie?o?a i?i?inoath/iai ian?iiy a ?ineeiieoeoa? oa
?ioeo aaeocyo.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. Niin?a aeey
ia?aaeiin?aii? ia?iaee ian?iiy aiaiaeo eoeueoo? i?aeaeuo?
aoaeoeai?noue ??cia?aeueiiai neia?ico, ca?eueoo? ??aaiue
acioo?enaoe?? oa o?iaeaei?noue oeeo eoeueoo?. Iiaeeaia na?aaeiaeua
aeey i?i?iuoaaiiy ieeeo noeioeth? i?i?inoaiiy ieeeo oa ??no ieeeiaeo
o?oaie, ui iiaea aeei?enoiaoaaoeny aeey ia?iaee aaaeeicaieethaaieo
?ineei, c iaoith i?aeaeuaiiy ?o o?iaeaeiino?. I/euai? e?i?aee
ei?aiaaeo aeciiaoaaie?o?a iiaeooue

i?aeoe/ii aeei?enoiaoaaoeny a yeino? aioeaaeoa??aeueieo, ia oiene/ieo
cania?a.

A caeaaeiino? a?ae aeice Aeaeinoei aeyaey? ye noeioethth/ee oae ?
?ia?a??oth/ee aoaeo, ui ciaeaea oe?iea canoinoaaiiy ia o?eueee o
n?euenueeiio ainiiaea?noa?, a ? o i?e?ia?ieia?/i?e i?iieneiaino?,
a?ioaoiieia??, iaaeeoeei? oa ?ioeo iai?yieao.

Iniaenoee aianie caeiaoaa/a. Aeena?oaio i?eeiaa iniaenoo o/anoue a
?ic?iaoe? iaoiaeieia?? ? i?ia?aie aeine?aeaeaiue, ?o aeeiiaii?, a
ioi?ieaii? ? iiaea/? cayaie ia aeiaoiaee. Caeiaoaa/ — aaoi?
oaoiieia?? iaea?aeaiiy nooi? ?a/iaeie Aeaeinoei (aeieueiaa o/anoue 70%),
iiaeeaiiai na?aaeiaeua aeey i?i?iuoaaiiy ieeeo (aeieueiaa o/anoue 30%),
niiniaa aeey ia?aaeiin?aii? ia?iaee ian?iiy ca?iiaiaiaeo eoeueoo?
(aeieueiaa o/anoue 40%). ?c ni?eueieo iaoeiaeo aenia?eiaio?a ?
ioae?eaoe?e caeiaoaa/ aeei?enoaa i?e caiae? ni?aaaoi??a ni?eueio
/anoeio ?acoeueoao?a.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iaoa??aee aeena?oaoe?eii?
?iaioe aeiiia?aeaeenue ia I?aeia?iaei?e iaoeia?e eiioa?aioe??
«A?ioao — 95»: “A?ioaoiieia?y iaea?aeaiiy ei?iiaiai a?eea, aeieia?/ii
/enoeo i?aia?ao?a, ui i?aeaeuothoue o?iaeaei?noue, i?ai?en?a, oa?iaio?a
oa a?oai?i?a ei?iiaiai i?ecia/aiiy” (Aei?i?iiao?ianuee, 1995); ia
nai?ia?ao eaai?aoi??? a?ieia?/ii? noeioeyoe?? ian?iiy ? ?ineei OAeAO
?i. A.A.Aeieo/aaaa oa IAe? a?ieia?? OAeO; caiaoaioiaai? ?ic?iaee (
8741, 9982, 18419, 95321061 ) aeei?enoiaothoueny a
n?euenueeiainaea?nueeiio ae?iaieoeoa?.

Ioae?eaoe??. Ca iniiaieie iaoa??aeaie aeena?oaoe?? iioae?eiaaii 4
?iaioe oa iaea?aeaii 4 iaoaioa.

No?oeoo?a oa ianya ?iaioe. Aeena?oaoe?y aeeeaaeaia ia 142
noi??ieao ? neeaaea?oueny ?c anooio, iaeyaeo e?oa?aoo?e, iaoiae?a oa
?acoeueoao?a aeine?aeaeaiue, aeniiae?a, i?iiiceoe?e ae?iaieoeoao oa
nieneo aeei?enoaii? e?oa?aoo?e. ?iaioa i?noeoue 19 oaaeeoeue,
11 iaethie?a, 2 ae?aa?aie oa 9

aeiaeaoe?a. Nienie aeei?enoaii? e?oa?aoo?e aeeth/a? 148 aeaea?ae.

IAOA??AE OA IAOIAeEEA AeINE?AeAEAIUe

Ioaie/i? aeno?aeoe ?c aeae?eaiue i?i?inoath/iai ian?iiy io?eioaaee
a iaeei /an aeiae i?ioyaii 5 aei?a. Aeeiai?eo ia?inoaiiy
aeciiaoaaie?o?a aea/aee ia NO-26 i?e 940 ii. Aei??thaaee ieenethaaeueii
— a?aeiiaiee iioaioe?ae ? ?I aeno?aeo?a c 1 ii 5 aeiao i?i?iuoaaiiy
ia ?I — 121. E?euee?noue

ei?aiaaeo aeae?eaiue oa ii?oi-a?iiao?e/i? ci?ie aniieooth/i? ciie
ei?aiy, ui ?inoa, a caeaaeiino? a?ae niiniao ia?aaeiin?aii? ia?iaee
aea/aee ia ian?ii? ai?ioo Oeaae?anueeee 10 oa ic. ioaieoe? Aeuea?aeoi
114. Ian?iiy ia?iaeyee aiaeith, ioaie/iei aeno?aeoii,
??cioi?o?iii, noniaic??th aeno?aeo-??cioi?o?i a e?eueeino? 3% a?ae ?o
iane oa i?i?iuoaaee i?ioyaii 3 ae?a. I?e?ineii?/ii aiae?coaaee ieiuo
ei?aiaaeo aeae?eaiue, e?euee?noue ei?aiaaeo aieine?a, ?o aeiaaeeio oa
oe?eio. Aeey a?iiao?e/ieo aei??thaaiue aeei?enoiaoaaee IA? — 11, IE — 1,
ieoey?io n?oeo, ieoey?-i?e?iiao? oa ia(?eo-i?e?iiao?. Ei?aiaa?
aeae?eaiiy ia?iaeyee 3,7%-aith iaoeeaiiaith neiith. Iaoaiaoe/io ia?iaeo
i?iaiaeeee ii Aeiniaoiao A.A. ( 1985 ). Oioia?aooaaee c aeei?enoaiiyi
i?e?ioioiianaaeee IOI — 3.

Na?aeee ioaie/iee aeno?aeo io?eioaaee ca oaoiieia??th, ?ic?iaeaiith
i?io. Iaoiiaei A.O. (A.N. 425569 NNN?, 1974 ), eio?ee ioieiaeaeoaaee oa
oeaio?eooaoaaiiyi a?aeie?aiethaaee e?ioiaeue. Io?eiaiee oeaio?eooaao
aenoooaaee i?e iecuee?e oaiia?aoo?? o aaeooi? ia IA — 960 ? ia?aaiaeeee
o ii?ioeoaaoee noai. Aea/aee niaeo?aeuei? oa?aeoa?enoeee na?aeiai
aeno?aeoo oa eiai nooiai aiaeiao ( ?a/iaeie Aeaeinoei ) a ae?aiacii?
oaeeue a?ae 400 aei 750 ii ia ?aa?no?oth/iio niaeo?ioioiiao?? NO-14.

?c ei?ai?a oa e?io?eueii aenooaieo aeciiaoaaie?o?a, io?eiaieo ?c
o?eaeaiieo i?i?inoe?a ioaieoe?, aeno?aaoaaee e?i?aee iine?aeiaii
noi?ooth oei?ioi?i:iaoaiie (2:1) oa oei?ioi?i:iaoaiie:aiaea (1:2:0,8) ?
io?eiaiee aeno?aeo oia?thaaee i?ae aaeooiii. E?i?aee aeno?aeoo
?ic/eiyee a oei?ioi?i? oa o?iiaoia?ao?/ii ?icae?eyee ia ieanoeiao
Silufol. Aecia/aee ai?no eiaeii? ?c o?aeoe?e niaeo?ioioiiao?e/ii (400
ii) ii eae?a?iai/iei e?eaei oa aeneiaethaaee o iea/ia e?io?eueii
aenooaieo aeciiaoaaie?o?a. Aiae?c e?i?ae?a o?eaeaiieo i?i?inoe?a oa ?o
aeciiaoaaie?o?a i?iaiaeeee a iaeiaeiaeo oiiaao. Aenia?eiaioe
iiaoi?thaaee o?ee?aoii a /ioe?ueio aiae?oe/ieo iiaoi?aiiyo.
?acoeueoaoe ia?iaeyee ca e?eoa???i Nouethaeaioa.

Aea/aee aieea ?a/iaeie Aeaeinoei ia ??ciaaiac eaanie? iaoiaeii
a?ioanooaaiiy ca Oo?aoeueeith ?.O. Ca eiio?ieue i?eeiaee oei??iaiiy
eaanie? o aiae?. Aaoa?iaoenei aiooaaee a eiioeaio?aoe?? 40 ia/ie, a
4,62 a. Io?aneaa-Neoaa (Serva) ?ic/eiyee a 1e. aeanoeeueiaaii? aiaee.
Aeaeinoei aeei?enoi-aoaaee a 100, 25 oa 10% — ai? eiioeaio?aoe??.
?iai/o ? caaaeueio aaeni?aoth/o iiaa?oith ei?aiaai?

nenoaie aecia/aee iaoiaeii Naa?i?ia Ae.A.-Eieiniaa I.I., ?o ia(?iiiao?ii
aei??yee ia(?i ei?ai?a. O?e iacaeaaei? aenia?eiaioe o /ioe?ueio
iiaoi?aiiyo ia?iaeyee noaoenoe/ii (Aeiniaoia A.A. 1985).

ian.%). Aea/aee i?i?inoaiiy ieeeo othothio caiaoiiai, a?oo?
cae/aeii?, yaeoi? neeaieenoii?, /oaooieea a?ioeaaiai (oi?ia cieioenoa)
oa Eiaae?? A?ai?on. Ieeie i?i?i- uoaaee o aeny/eo e?aieyo
aeine?aeaeoaaiiai na?aaeiaeua i?e oaiia?aoo?? (23-30(N i?ioyaii 12
aiaeei. Eiio?ieai aoea cae/aeia aeanoeeueiaaia aiaea c naoa?icith ??cii?
eiioeaio?aoe?? (10 — 20 an.%).

Aenia?eiaioe i?iaiaeeee o o?ee?aoi?e iiaoi?iino? c i?ae?aooaaiiyi /a?ac
12 aiaeei i?i?ineeo ieeeiaeo ca?ai o o?ueio iieyo ci?o i?e?ineiio.
Aei??thaaee 20 ieeeiaeo o?oaie ii eiaeiiio aeine?aeaeoaaiiio
na?aaeiaeuo.

Aeine?aeaeoaaee aieea ei?aiaaeo aeciiaoaaie?o?a ia ?iciiiaeaiiy
aoeueai/eiaeo aaeoa??e Rhizobium L., ye? ?icaiaeeee a ioaie/iiio
aeno?aeo? oa aecia/aee ?ioaineai?noue ?o ?iciiiaeaiiy i?yiei
i?e?ineiioaaiiyi o oiieiio oa?? e?/eeueii? eaia?e Oaoene?. Aeey
i?aeaeuaiiy o?iaeaeiino? ca?iiaiaiaeo eoeueoo? iaie cai?iiiiiaaiee
niin?a aeey ia?aaeiin?aii? ia?iaee ian?iiy oeeo eoeueoo?, ui ai?uo?
i?aia?ao aoeueai/eiaeo aaeoa??e (??cioi?o?i) oa ?ic/eiiee (aiaeiee
?ic/ei ei?aiaaeo aeciiaoaaie?o?a) a ni?aa?aeiioaii? 2,8 : 97,2 ian.%
a?aeiia?aeii.

Iieueia? aeine?aeaeaiiy ii aea/aiith aieeao ia?aaeiin?aii? ia?iaee
ian?iiy ai?ioo Oeaae?anueeee 10 aeeiioaaee ia aeine?aeiiio iie?
Oa?e?anueeiai aa?ioi?aa?neoaoo a 1991 — 1993 ?ieao. Ia?aaeiin?aio
ia?iaeo ian?iiy aiaeith, ??cioi?o?iii, aeno?aeoii oa noniaic??th
aeno?aeo — ??cioi?o?i i?iaiaeeee o e?eueeino? 3,0-3,5% a?ae ?o iane.
Iae?eiaa ieiua ae?eyiie 25i . ?ici?uaiiy aa??aio?a ?aiaeii?ciaaia o
/ioe?ee?aoi?e iiaoi?iino?. Aia?a?th i?i?inoaiiy, noiae?noue aecia/aee ca
aea?aenoaiaea?oii 12038-84, aiae?c no?oeoo?e o?iaeath ca iaoiaeeeith
Iena?aaa A.?.(1964). Ai?no a?eeo o ca?i? ai?ioa aiae?coaaee iaoiaeii
?io?a/a?aiii? niaeo?ineii?? ia aiae?caoi?? ?io?ai?ae — 61 (E??uaiei
A.I. oa ?io?, 1986). Aeai? iieueiaeo aeine?ae?a ia?iaeyee iaoiaeii
aeenia?n?eiiai aiae?co ca Aeiniaoiaei A.A. (1985).

A?ae?iiiiiee caeaiee ei?i ae?iuoaaee ia aaaaoaoe?eieo
iiaa?oiyo ?ic?iaeaii? onoaiiaee

i?ioyaii 7 ae?a aei oace ianiaeuii? noeaeino? y/iaith (11-15ni.) a
ooo/ieo oiiaao na?oeiaiai oa ei?aiaaiai aeeaeaiiy. Canoiniaoaaee
nenoaio iieeao i?aeoiieaiiyi, o?e ?ace ia aeiao /a?ac 8 aiaeei.
O?eaae?noue na?oeiaiai ia??iaeo 10-12 aiaeei ia aeiao c ina?oeaiiyi
aaaaoaoe?eii? iiaa?oi? 1000 — 1200 ethen. Aeine?aeaeaiiy aieeao ??cieo
iiaeeaieo ?ic/ei?a aeey caieiaeaiiy iin?ao y/iaith ia ??no caeaii?
iane oa ?? ye?noue i?iaiaeeee c aeei?enoaiiyi i?ia?aeueieo iiaeeaieo
noi?o?a ??ooa?a, Aaeue??aaey, Oiaeaiaea-A?iiia oa 50%-ai aiaeiiai
?ic/eio ?a/iaeie Aeaeinoei acyoi? o e?eueeino? 3 % a?ae iane
caieiaeoth/eo ?ic/ei?a. No?oeoo?o o?iaeath ei?io ae?aaeaee ianith
caeaieo iao?a, aenioith ?ineei, ne?ith ianith eenoy, ianith ei?aiaaiai
oa?o. Ai?no i?ioa?io, aee?o, ee?oeiaeie oa ciee a eeno? caeaiiai ei?io
aecia/aee iaoiaeii ?io?a/a?aiii? niaeo?ineii?? ia aiae?caoi?? ?io?ai?ae
— 61 ( E??uaiei A.I. oa ?io?, 1986).

y ia?anaaeee aiaei?ino? ia na?aea na?aaeiaeua. Aeeiai?eo ?? ?inoo
aecia/aee niaeo?ioioiiao?e/ii i?e 540 ii c 2-ai ii 8-e aeaiue
eoeueoeaoaaiiy. O?e iacaeaaei? aenia?eiaioe o o?ee?aoi?e iiaoi?iino?
ia?iaeyee noaoenoe/ii ca e?eoa???i Nouethaeaioa.

Ia eoeueoo?? aiaei?ino? Paramaecium caudatum L. a?ioanooaaiiyi
aecia/aee o?oiioeeaeio aeoeai?noue na?aeiai ioaie/iiai aeno?aeoo oa eiai
nooiai aiaeiao a caeaaeiino? a?ae ai?noo a ?o neeaae? e?ioiaeth.
Aeny/o e?aieth eoeueoo?e ia i?aaeiaoiiio nee? iii?uaee iaae /aniaei
neeii c ei?aiaaeie aeciiaoaaie?oaie na?aeiai aeno?aeoo oa eiai nooiai
aiaeiao c e?ioiaeai oa aac iueiai. I?yiei i?e?ineiioaaiiyi aecia/aee
/an cae?i/aiiy ?ooo aiaei?inoae oa ?ic?aooieiaei iaoiaeii
anoaiiaethaaee o?oiioeeaeio aeoeai?noue (A?iaeceinueeee A.I., 1973).

Aeine?aeaeaiiy aioeaaeoa??aeueii? aeoeaiino? e?i?aeieo aeno?aeo?a
aeciiaoaaie?o?a i?i?inoe?a ioaieoe? i?iaiaeeee ca caaaeueii aaeeaaiei
iaoiaeii aecia/aiiy ae?aiao?o a?eueii? a?ae i?e?ii?aai?ci?a ciie (
Iaeneiiaa ?.A. oa ?io?, 1984 ), aeey /iai aeei?enoiaoaaee i?e?ii?aai?cie
Escher?chia coli, Staphylococcus aureus, ?seudomonas aeruginosae oa
Streptococcus. Ii??aithaaee c aioea?ioeeaie (aaioii?oeeiii oa
iie?i?eneiii), a oaeiae c /enoei oei?ioi?iii. Aioeaaeoa??aeueio
aeoeai?noue aioea?ioee?a i?eeiaee ca 100 %, aaeoa??oeeaei?noue
e?i?aeieo aeno?aeo?a aeciiaoaaie?o?a aeneiaethaaee o a?aenioeao a?ae
aeoeaiino? aioea?ioee?a.

?ACOEUeOAOE AeINE?AeAEAIUe OA ?O IAAIAI?AIIss

Aeine?aeaeaiiy aeae?eueii? nenoaie
i?i?inoe?a ioaieoe?.

?ic?iaeyth/e i?aeoe/i? iniiae iaea?aeaiiy ei?aiaaeo aeciiaoaaie?o?a
aea/aee iaea?eueoo aeae?eueio caeaoi?noue i?i?inoath/iai ian?iiy.
Anoaiiaeee, ui eiao?oe??io i?iioneaiiy aiaeiiai

?ic/eio ei?aiaaeo aeciiaoaaie?o?a iaea?eueoa ci?ith?oueny c ia?oi? ii
o?aoth aeiao i?i?inoaiiy. Iaea?eueoa aeae?eaiiy aeciiaoaaie?o?a
niinoa??aa?oueny o ia?o? o?e aeiae. I?e oeueiio ?I aiaeiiai ?ic/eio ?o
ci?uo?oueny o a?e i?aeaeuaiiy eeneioiino? aei 4,7 iae. ?I, a ia i(yoo
aeiao eeneioi?noue cieaeo?oueny. Oea iiaiaeaeo?oueny c c?inoaiiyi
aeoeaiino? ?ii?a aiaeith o i?ioean? ?inoo ei?aiy c ia?oi? ii /aoaa?oo
aeiao o 19 ?ac?a oa ciaioaiiyi oeueiai iieacieea ia i(yoo aeiao, ui
a?aeiia?aea? ci?i? ?I. Ieenethaaeueii-a?aeiiaiee iioaioe?ae ?ic/eio
aeciiaoaaie?o?a oaeiae cacia? aeeaieeo ci?i. Oae ia ia?oo aeiao a?i iaa
iaea?eueoa cia/aiiy, iio?i ciaioeany o 13 ?ac?a ?c iacia/iei i?aeeiiii
ia /aoaa?oo oa i(yoo aeiao ?inoo ei?aiy. Aeniea eaa?euei?noue oeeo
iieaciee?a na?ae/eoue i?i oaeaee? ye ye?ni?, oae ? e?euee?ni? ci?ie
neeaaeo eiiiiiaio?a aeciiaoaaie?o?a. Aea/aiiy ?yaeo i?eaaaeaieo
iieaciee?a aeicaiey? iia?/ii ioa?aeoa?ecoaaoe ci?ie aeae?eueii? nenoaie
ei?aiy a i?ioean? eiai ?icaeoeo oa anoaiiaeoe, ui o?aoy aeiaa ?
iaea?eueo ni?eyoeeaith aeey io?eiaiiy ye?niiai ?ic/eio ei?aiaaeo
aeciiaoaaie?o?a. Aieea ??cieo niinia?a ia?aaeiin?aii? ia?iaee
ian?iiy ai?ioo Oeaae?anueeee 10 ia e?euee?noue ei?aiaaeo aeae?eaiue
oa ii?oi — a?iiao?e/i? ci?ie aniieooth/i? ciie ?inoo/iai ei?aiy,
i?aaenoaaeai? a oaae.1.

Oaaeeoey 1

Ieiua neeciaeo aeae?eaiue oa ciia aniieooaaiiy ?inoo/iai ei?aiy a
caeaaeiino?

a?ae niiniao ia?aaeiin?aii? ia?iaee ian?iiy ai?ioo.

Niin?a Ieiua Ei?aiaa? aieinee

i?aaeiin?aii? neeciaiai

ia?iaee /ioea E?euee?noue Oe?eia
Aeiaaeeia

ian?iiy

2 % aei % aei % aei
% aei

i?e?ii eiio?. Oooe eiio?. I?e?ii eiio?. I?e?ii
eiio?.

Nooa ian?iiy,

eiio?ieue 483728,9 100,0 78 100,0 14,94 100,0 251,23
100,0

Caieiaeaiiy

aiaeith 616962,0 127,5 71 91,0 15,25 102,1 245,92
97,2

?iieoeyoe?y

??cioi?o?iii 555458,0 114,8 67 85,9 14,66 98,1 249,21
99,2

Caaaa/aiiy

aeno?aeoii 643870,0 133,1 74 94,9 14,15 94,7 197,57
78,6

Caaaa/aiiy

aeno?aeoii ? 945624,0 195,5 145 185,9 14,17 94,9 323,60
128,8

??cioi?o?iii

I?? 8698,97 9,15 1,18 37,67

05

Ipei?oea: e?euee?noue eipaiaaeo aieine?a, ?o oepeia oa aeiaaeeia
aei??yi? ia

a?aenoai? 1280 i?epii ciie aniieooaaiiy eipaiy.

Ia?aaeiin?aia ia?iaea ian?iiy ai?ioo ioaie/iei aeno?aeoii
i?aeaeueea ieiuo ei?aiaaeo aeae?eaiue o 1,3 ?ace, a noniaic?y
aeno?aeo — ??cioi?o?i o 2 ?ace oa ca?eueoeea e?euee?noue ei?aiaaeo
aieine?a ciie aniieooaaiiy ? ?o aeiaaeeio o 1,9 oa 1,3 ?ace
a?aeiia?aeii, o ii??aiyii? c eiio?ieai. ?io? aeaee ia?iaee ia aeyaeee
a??ia?aeiiai aieeao ia oi?ioaaiiy ciie aniieooaaiiy.

Oaeei /eiii, iaea?eueoa e?euee?noue ei?aiaaeo aeae?eaiue oa
iaeneiaeueia ca?eueoaiiy ciie

aniieooaaiiy o ai?ioa niinoa??aathoueny i?e ia?iaoe? ian?iiy noniaic??th
aeno?aeo — ??cioi?o?i.

Oaoiieia?y iaea?aeaiiy nooi?
?a/iaeie Aeaeinoei.

Na?aeee ioaie/iee aeno?aeo aeei?enoiao?oueny a ?ineeiieoeoa?
aeey ia?aaeiin?aii? ia?iaee

ian?iiy iieueiaeo eoeueoo? c iaoith i?aeaeuaiiy ?o o?iaeaeiino?.
Aea, a?i ia? cia/i? iaaeie?ee — iecueeo oaoiieia?/i?noue ( /a?ac 2-4
aiaeeie ao?a/a? aeoeai?noue, ia caa??aa?oueny ), a ?acoeueoao? /iai ia

io?eiaa aeinoaoiueiai iioe?aiiy o n?euenueeiainiiaea?nuee?e i?aeoeoe?.
Oaoiieia?y iaea?aeaiiy nooi? ?a/iaeie Aeaeinoei ?c aiaeiiai
?ic/eio ei?aiaaeo aeciiaoaaie?o?a i?i?inoath/iai ian?iiy

eiai oeo?o. Oeo? nooi? ?a/iaeie — 0,004472. Nooa ?a/iaeia
Aeaeinoei caa??aa? aeiaaee /an a?ieia?/io aeoeai?noue oa caaacia/o?
aenieo oaoiieia?/i?noue aeei?enoaiiy a ?ineeiieoeoa?.

A?AeAe?EAIIss

CIEA IAAEOEAII?

I AOAI- IAN?IIss I?I?IUOAAIIss AECIIAOA- O?AEOe??

IAN?IIss AIE?O?A
OeAIO?EOOAOAAIIssI

|

— AE?OA — OAIIA?AOO?A — AIAeA
OIEIAe

— NOIA?AE?OA — AIEIA?NOUe

— OAI?ssAA

CAII?IAEOAAIIss

AEOEAII? NOOEA

II AOAI- O?AEOe??

AENO?AEOO

OIEIAe AAEOOI
IOIEIAeAEAIIss I?Ae?A??A

EAIA?E
AENOOOAAIIAI IAOA??AEO

IA?AA?Ae

NOOI? ?A*IAEIE OIAEIAEA OA

III AOAI- O II?IOEOAAOEE AA?IAOECAOe?ss

NOAI I?IAeOEOO

— OIEIAe; OEUeO?ACAOEIAA IIAe??AEAIIss

?en.1 Aeie-noaia oaoiieia?/iiai i?ioeano iaea?aeaiiy nooiai ioaie/iiai
aeno?aeoo

(?a/iaeie Aeaeinoei) ?c aeciiaoaaie?o?a i?i?inoath/iai ian?iiy
ioaieoe?.

E?i?aeiee neeaae ei?aiaaeo
aeciiaoaaie?o?a.

I?iaaaeaii ye?nia oa e?euee?nia aeine?aeaeaiiy e?i?aeiiai neeaaeo
ei?ai?a oa ei?aiaaeo aeciiaoaaie?o?a o?eaeaiieo i?i?inoe?a ioaieoe?
ye iaeiaio aea/aieo eiiiiiaio?a aeae?eueii? nenoaie ( oaae.2 ).
Anoaiiaeaii, ui 54% an?o e?i?ae?a ei?ai?a i?eoiaeeoueny ia
oinoie?i?aee, a 46% ia iaeo?aeuei? e?i?aee.

Oaaeeoey 2

E?i?aeiee oa oinoie?i?aeiee neeaae ei?ai?a o?eaeaiieo i?i?inoe?a

oa ?o aeciiaoaaie?o?a.

E ? i ? ae i e e n e e a ae

O?aeoe?? Ei?ai? Aeciiaoaaie?oe

iea/ia % iea/ia %

Oinoie?i?aee 64,93(1,21 53,7(1,7 52,38(2,70 54,7(1,2

O?eaoeeeae?oea??aee 10,65(0,89 8,7(0,4 0 0

Iaaoapeo?eiaai?

aee?i? eeneioe 11,67(1,59 9,4(0,9 14,08(0,63 15,1(0,6

?ineeii? noa?i?aee 9,47(1,09 5,9(0,7 0 0

Ao??e ?ineeiieo

noa?i?ae?a 28,30(1,10 23,2(0,3 27,96(1,14 30,3(0,3

O i n o i e ? i ? ae i e e n e e a ae

Oinoaoeaeeeoie?i 9,5(0,9 34,0(2,1 11,6(0,3 21,0(1,3

Oinoaoeaeeeaoaiieai?i 4,0(0,8 14,3(2,3 9,6(0,6 17,1(0,8

Oinoaoeaeee?iic?oie 4,3(0,8 15,4(2,9 13,5(1,3 23,9(1,5

Oinoaoeaeia eeneioa 2,9(0,2 10,4(0,8 0 0

Ea?aeeie?i?i 3,2(0,7 11,5(0,6 10,3(0,8 18,4(1,2

Ia ?iaeaioeo?eiaaia

o?aeoe?y 4,0(0,5 14,3(1,3 10,9(0,6 19,6(0,7

A e?i?aeiiio neeaae? aeciiaoaaie?o?a a?aenooi? aea? o?aeoe??
iaeo?aeueieo e?i?ae?a. Oinoie?i?aeiee neeaae ei?ai?a i?aaenoaaeaiee 6
o?aeoe?yie. Iaea?eueoa e?euee?noue, 34 % a?ae an?o oinoie?i?ae?a,
i?eiaaea? ia oinoioeaeeeoie?i. ?io? o?aeoe?? i?aaenoaaeai? i?aeoe/ii a
aeecueeeo ni?aa?aeiioaiiyo ? ia eiaeio c ieo i?eiaaea? a?ae 10 aei 15%.
A oinoie?i?aeiiio neeaae? ei?aiaaeo aeciiaoaaie?o?a a?aenooiy
oinoaoeaeia eeneioa. A?aenooi?noue oinoaoeaeii? eeneioe oa
o?eaoeeeae?oea?eae?a (oi?ie aeaiiioaaiiy e?i?ae?a) na?ae/eoue i?i
iayai?noue niaoeeo?/iiai iaoai?cio aeae?eaiiy e?i?aeieo eiiiiiaio?a o
na?aaeiaeua.

Aeine?aeaeaiiy iieacaee, ui e?i?aeiee neeaae ei?aiaaeo
aeciiaoaaie?o?a a?ae??ciy?oueny a?ae e?i?aeiiai neeaaeo ei?ai?a. Oae,
oinoie?i?aee oa iaeo?aeuei? e?i?aee aoiaeyoue aei ?o neeaaeo o ??cieo
ni?aa?aeiioaiiyo.

Aioeaaeoa??aeueia aeoeai?noue
ei?aiaaeo aeciiaoaaie?o?a.

Aeine?aeaeaiiy aioeaaeoa??aeueii? aeoeaiino? e?i?aeieo aeno?aeo?a
ei?aiaaeo aeciiaoaaie?o?a iieacaee, ui e?i?aee, ia?aaoaath/? o ?o
neeaae?, ia iathoue aaeoa?eoeeaeieo aeanoeainoae (oaae.3). Aiie ia
ooai?thaaee a?eueieo a?ae i?e?ii?aai?ci?a cii, a e?i?aee, aeae?eai? ?c
neeaaeo ei?aiaaeo aeciiaoaaie?o?a oa ca?eueiai? a?ae ca(yceo c ?ioeie
eiiiiiaioaie, i/aaeaeii, a?eeaie oa aoaeaaiaeaie, oae? ciie
ooai?thaaee. Oea aeaco? ia o?, ui e?i?aee o neeaae?
aeciiaoaaie?o?a ciaoiaeyoueny o eiiieaenao, iiaeeeai c a?eeaie. Oae,
e?i?aee, aeae?eai? ?c neeaaeo aeciiaoaaie?o?a, aeey Pseudomonas
aeruginosae ooai?thaaee a?eueio ciio o 14 ii, ui a?aeiia?aea? 88 %
?ia?a??oaaiith ?inoo o ii??aiyii? c aioea?ioeeaie.

Oaaeeoey 3

Aioeaaeoa??aeueia aeoeai?noue e?i?ae?a ei?aiaaeo aeciiaoaaie?o?a oa
aioea?ioee?a.

Ae?aiao? a?eueii? a?ae i?e?ii?aai?ci?a ciie, ii.

Ooaie c aaioi- c iie?- c caaaeuei.c e?i?aeaie,

i?oeeiii i?eneiii aeciiaoa- aeae?eaieie eiio?ieue

i?e?ii?aaieci?a aie?oaie ?c aeciia-

ei?ai?a oaaie?o?a

ei?ai?a

Escherichia coli — — 0 18,0(0,3 0

Staphylococcus — — 0 13,0(0,9 0

aureus

Pseudomonas 16,0(0,2 16,0(0,3 0 14,0(0,5 0

aeruginosae

Streptococcus — — 0 30,0(0,6 0

I?ei?oea. 0 — a?aenooi?noue aioeaaeoa??aeueii? aeoeaiino?; eiio?ieue-

oei?ioi?i (?ic/eiiee, yeei aeno?aaoaaee e?i?aee c aeciiaoaaie?o?a).

??no ?ioeo i?e?ii?aai?ci?a i?eai?/oaaany ua a?eueoa. Aeine?aeaeaiiy
iieacothoue, ui e?i?aee, aeae?eai? ?c neeaaeo aeciiaoaaie?o?a,
aeyaeythoue aaeoa??oeeaeio aeoeai?noue, aeecueeo aei aeoeaiino?
noaiaea?oieo aoea?ioee?a.

Aieea ei?aiaaeo aeciiaoaaie?o?a ia
??ciaaiac eaanie?.

Aeine?aeaeaiiy aieeao ei?aiaaeo aeciiaoaaie?o?a ia aeoeai?noue
??ciaaiaco o ii??aiyii? c oe?iei aeei?enoiaoaaieie o i?aeoeoe?
aaaaoaoeaiiai ?iciiiaeaiiy oei??ithaa/aie (aaoa?iaoeneiii oa
Io?aneaa-Neoaa) iieacaee, ui i?e ia?iaoe? caeaii? ??cee eaanie? 25%-ei
Aeaeinoeiii aeiaeaoeiaeo ei??ioe?a ooai?eeinue o 1,8 ?ace a?eueoa i?ae
ia eiio?ie?, a i?e ia?iaoe? aaoa?iaoeneiii oa Io?aneaa-Neoaa —
a?aeiia?aeii o 1,3 oa 1,7 ?ace. Ia?iaea 25%-ei Aeaeinoeiii ca?eueoeea
aeiaaeeio ciie oei??ithaaiiy ??cee eaanie? o 1,7 ?ace. Ioaea, 25%-ee
Aeaeinoei ia iinooia?oueny ca aeoeai?noth ??ciaaiaco a?aeiiei
oei??ithaa/ai. Iiaeeeai, a?i i?aeneeth? ia?enoaiaoecaoe?th oeaiei,
iaaiaeiaiiy ee?oei oa aeoea?co? iai?i ?a/iaei, ui iiceoeaii
aieeaa? ia ei?aiaooai?aiiy.

Aieea ei?aiaaeo aeciiaoaaie?o?a ia
i?i?inoaiiy ieeeo.

ian.% ) (oaae.4).

Oaaeeoey
4

A?aenioie i?i?inoaiiy (?) oa na?aaeiy aeiaaeeia ieeeiaeo o?oaie
(L)

/a?ac 12 a. i?ney i?i?iuaiiy ieeeo ??cieo aeae?a ?ineei.

A?ooa cae/aeia, ssaeoiy *oaooiee
a?ioeaaee,

Aeaee n?aaeiaeua «Ethaeieoey Eeaiia” neeaieenoia oi?ia
cieioenoa

(aa??aioe)

?,% L,iei ?,% L,iei ?,% L,iei

Eiio?ieue-

20% naoa?ica 4,5 615 76,3 424 41,3 203

Oaae ( Aeaeinoei,

ian.%:

0,1 0,0 — 0,0 — 0,0 —

0,01 19,4 198 55 352 0,0 —

0,001 87* 883 93* 719 45 512

0,0001 89* 914 94* 1099 91* 553

0,00001 95* 1231 97* 1685 89* 743

0,000001 91* 1053 90* 524 82* 563

0,0000001 85* 646 88* 661 69* 407

0,00000001 90,8* 666 92* 711 74* 378

* — ae?ia?aeii ia 5% ??ai? cia/aiiy

ieoeoa? aeey i?aeaeuaiiy o?iaeaeiino? i?e ia?iaoe? aaaeeicaieethaaieo
?ineei, a oaeiae o naeaeoe?eii-aaiaoe/ieo aeine?aeaeaiiyo.

Aieea ei?aiaaeo aeciiaoaaie?o?a oa aaeoa??e ?iaeo
Rhizobium ia o?iaeaei?noue

ca?iiaiaiaeo
eoeueoo?.

A?aeiii, ui e?euee?noue o?eniaaiiai ?ineeiaie acioo i?iii?oe?eia
e?eueeino? aoeueai/eiaeo aaeoa??e, ui ca?aceee ei??iue ?ineei. Aea/aee
aieea ei?aiaaeo aeciiaoaaie?o?a ia ?ioaineai?noue ?iciiiaeaiiy
aoeueai/eiaeo aaeoa??e. ?icaaaeaiiy aoeueai/eiaeo aaeoa??e o ?ic/ei?
ei?aiaaeo aeci-iaoaaie?o?a i?eaiaeeoue aei oaeaeeiai ?o ?iciiiaeaiiy.
Oae, /a?ac 2 aiaeeie ?ieoaaoe?? e?euee?noue ?o o i?aeaioiaeai?e
noniaic?? a 5,8 ?ace aeua, i?ae ia eiio?ie? (?en.2). A iane?aeie oeueiai
niinoa??aa?oueny oaeaeea ?o ?iciiiaeaiiy ? aoaeoeaia ca?aaeaiiy ei??iiy
aoeueai/eiaeie aaeoa??yie. Ia?aaeiin?aia ia?iaea ian?iiy ai?ioo
Oeaae?anueeee 10 noniaic??th aeno?aeo — ??cioi?o?i i?aeaeueea
o?iaeaei?noue

ai?ioo o na?aaeiueiio ca 3 ?iee ia 5,6 oe/aa aai o 1,2 ?ace (?en.3). Ia
eiio?ie? aiia aei??aithaaea 29,0 oe/aa, a i?e ia?iaoe? ei?aiaaeie
aeciiaoaaie?oaie ? ??cioi?o?iii — 34,6, oiaoi iaaeaaaea a?iaeath
neeaea 19,3 %.

?en.3. O?iaeaei?noue ai?ioo Oeaae?anueeee 10 o caeaaeiino? a?ae
niiniao ia?aae-

iin?aii? ia?iaee ian?iiy (1991-1993 ??.).

canoinoaaiiyi ( aa?o?noue aaeoa?ii? ii?ie Aeaeinoeio oa ??cioi?o?io ).
Oiiaii /enoee i?eaooie a?ae canoinoaaiiy niiniao aecia/a?oueny ca
oi?ioeith

O*I=A((Oi-Oa)-(A(A(?),

aea A-oe?ia iaeiiai oeaioia?o ian?iiy ai?ioo + 40% ni?oiaa iaaeaaaea
(ae?th/? oe?ie 1997 ?.), a?i.-26,0+10,40=36,40; Oi-o?iaeae iiaiai
aa??aioo, oe/aa-34,6; Oa -o?iaeae aaciaiai aa??aioo, oe/aa-29,0; A-oe?ia
iaeiiai a?aio ?a/iaeie Aeaeinoei, a?i.-0,61; A-aaeoa?ia ii?ia Aeaeinoeio
i?ae ai?io, a?ai -27,0; ?-oe?ia aaeoa?ii? ii?ie ??cioi?o?io, a?i.-7,0.

O*I=36,4((34,6-29,0)-(0,61(27+7,0)=180,34 a?i.

I?e aeo?aoao ia ai?iaaaeaeaiiy 23,5 a?i/aa oiiaii /enoee
i?eaooie aei??aith? 180,34 a?i/aa.

Ei?aiaa? aeciiaoaaie?oe noeioeththoue ?iciiiaeaiiy aoeueai/eiaeo
aaeoa??e oa i?aeaeuothoue aoa-eoeai?noue aeei?enoaiiy o i?aeoeoe?
?ineeiieoeoaa i?aia?aoo aoeueai/eiaeo aaeoa??e — ??cioi?o?io.

Aeei?enoaiiy ei?aiaaeo aeciiaoaaie?o?a o a?ae?iiiii?e
eoeueoo?? i?e ae?iuoaaii?

caeaiiai
ei?io.

Aeine?aeaeaiiy iiaeeeaino? aeei?enoaiiy ei?aiaaeo aeciiaoaaie?o?a
o oaoiieia?? ae?iuoaaiiy caeaiiai a?ae?iiiiiiai ei?io iieacaee, ui
i?iieai? aiaee, ye? ai?uothoue 50% — ao ?a/iaeio Aeaeinoei,
iathoue ii??aithaai? iieacieee ye ca no?oeoo?ith caeaiiai ei?io, oae ?
ca eiai o?i?/iei neeaaeii c ei?iii, ae?iuaiei i?e aeei?enoaii?
i?ia?aeueieo iiaeeaieo noi?oie ??ooa?a,

Aaeue??aaey oa Oiaeaiaea — A?iiia (?en.4).

Aeei?enoaiiy ei?aiaaeo aeciiaoaaie?o?a o neeaae? caieiaeoth/eo
?ic/ei?a i?e ae?iuoaaii?

caeaiiai ei?io ca a?ae?iiiiiith oaoiieia??th aeicaiey? ia
aeei?enoiaoaaoe o?i?/i? ?a/iaeie oa ni?inoeoe oaoiieia?th.

Aieea Aeaeinoeio ia ?iciiiaeaiiy
aiaei?ino? Dunaliella.

i?ecaiaeeei aei niia?eueiaiiy ?inoo eoeueoo?e ia

Ia?niaeoeai?noue aeei?enoaiiy ?a/iaeie Aeaeinoei.

Ia?aaaae nooi? ?a/iaeie Aeaeinoei aeey n?euenueeiainiiaea?nueeiai
ae?iaieoeoaa iieyaathoue o oiio, ui a?i: 1) aeiaai caa??aa?
a?ieia?/io aeoeai?noue (0,5-1 ??e); 2) aeei?enoiao?oueny a iioeiaeuei?
no?iee oa ca ni?eyoeeaeo oiia (iiaiaea, oaoi?ea oa ?i.); 3) ia? iecueeo
nia?aa?o?noue.

AENIIAEE

1. Aeine?aeaeaiiy aeae?eueii? ae?yeueiino? i?i?inoath/iai ian?iiy
ioaieoe? iieacaei, ui iaeaeua aeae?eueia ni?iiiaei?noue niinoa??aa?oueny
ia 3 aeiao i?i?inoaiiy, ? oeae ia??iae ? iaea?eueo ni?eyoeeaei aeey
io?eiaiiy ye?nieo ei?aiaaeo aeciiaoaaie?o?a. Ia?iaea ian?iiy
n?euenueeiainiiaea?nueeeo ?ineei ioaie/iei aeno?aeoii aoaeoeaii aieeaa?
ia ei?aiaa? aeae?eaiiy oa oi?ioaaiiy ciie aniieooaaiiy o ioaieoe?, a
noniaic?y aeno?aeo-??cioi?o?i o ai?ioa.

2. ?ac?iaeaia oaoiieia?y iaea?aeaiiy nooi? ?a/iaeie Aeeaeinoei ?c
aiaeiiai ?ic/eio ei?aiaaeo aeciiaoaaie?o?a, yea ia? no?eeo a?ieia?/io
aeoeai?noue, iie?ooieoe?iiaeuei?noue oa oaoiieia?/i?noue a
n?euenueeiainiiaea?nueeiio ae?iaieoeoa? (iaoaio 18419 (UA) Oe?a?ie,
2095956 (RUS) ?in??).

3. Aeine?aeaeaii ye?niee oa e?euee?niee e?i?aeiee neeaae ei?ai?a oa
ei?aiaaeo aeciiaoaaie?o?a o?eaeaieo i?i?inoe?a ioaieoe?. Io?eiai?
?acoeueoaoe iieacaee, ui e?i?aeiee neeaae ei?aiaaeo aeciiaoaaie?o?a
a?ae??ciy?oueny a?ae e?i?aeiiai neeaaeo ei?ai?a.

4. E?i?aee, aeae?eai? ?c ei?aiaaeo aeciiaoaaie?o?a oa ca?eueiai? a?ae
ca(yceo c ?ioeie eiiiiiaioaie, aeyaeythoue aioeaaeoa??aeueio
aeoeai?noue, a oie /an ye e?i?aee, ia?aaoaath/? o neeaae?
ei?aiaaeo aeciiaoaaie?o?a oe? aeanoeaino? ia aeyaeythoue.

5. Aeine?aeaeaiiy aieeao ei?aiaaeo aeciiaoaaie?o?a ia ?ineeii?
ia(?eoe iieacaei(

a) ui iecuee? eiioeaio?aoe?? (1(10 ia/ie) noeioethaaee ??no oa
?iciiiaeaiiy ea?ioeiinii? aiaei?ino? Dunaliella viridis Teod, a
i?aeaeuaiiy eiioeaio?aoe?? aei 1(10 ia/ie aaeueioaaei oeae i?ioean;

a) a eiioeaio?aoe?? 1(10 ian.% aeciiaoaaie?oe aoaeoeaii ae?thoue
a yeino? eiiiiiaioa ?ic?iaea-iiai na?aaeiaeua aeey i?i?iuoaaiiy ieeeo,
ca?eueooth/e a?aenioie eiai i?i?inoaiiy (iaoaio 9982 (UA) Oe?a?ie);

a) ca aeoeai?noth ??ciaaiaco 25% Aeaeinoei ia iinooia?oueny a?aeiiei
oei??ithaa/ai ( aaoa?iaoeneio oa Io?aneaa — Neoaa );

a) aaaaeaiiy ei?aiaaeo aeciiaoaaie?o?a aei neeaaeo caieiaeoth/eo
?ic/ei?a i?e ae?iuoaaii? a?ae?iiiiiiai ei?io iie?ioo? eiai ye?noue oa
aeicaiey? ia aeei?enoiaoaaoe i?ia?aeuei? iiaeeai? noi?o?;

ae) na?aeii?eaioiaeaiee aiaeiee ?ic/ei ei?aiaaeo aeciiaoaaie?o?a ia?
a?eueo aenieo o?oiioeeaeio aeoeai?noue, i?ae eiai nooee aiaeia;

a) ?ic?iaeaiee niin?a aeey ia?aaeiin?aii? ia?iaee ian?iiy
ai?ioo i?aeaeuo? acioo?enaoe?th oa o?iaeaei?noue oe??? eoeueoo?e ca
?aooiie noeioeyoe?? ?iciiiaeaiiy aoeueai/eiaeo aaeoa??e oa ?inoo
aoaeoeaiino? ca?aaeaiiy ei?aiaai? nenoaie (iaoaio 8741 (UA) Oe?a?ie).

6. ?a/iaeia Aeaeinoei ? ei?aiaaei aeciiaoaaie?oii iaoo?aeueiiai
iioiaeaeaiiy. Aiia ia? eiiieaeniee neeaae oa aeyaey?
iie?ooieoe?iiaeuei?noue ae??, ui iiaea oe?iei aeei?enoiaoaaoeny o
?ineeiieoeoa?.

I?IIICEOe??
AE?IAIEOeOAO

1. Aeey ia?aaeiin?aii? a?inoeioeyoe?? ian?iiy iieueiaeo eoeueoo?
aeioe?eueii aeei?enoiaoaaoe i?e?iaeiee ei?aiaaee aeciiaoaaie?o,
aenieioaoiieia?/io ?a/iaeio Aeaeinoei.

2. C iaoith i?aeaeuaiiy a?iaeaeiino? aiaiaeo eoeueoo?, aeei?enoiaoaaiee
o ?ineeiieoeuee?e i?aeoeoe? acioo?enoth/ee aaeoa??aeueiee i?aia?ao
??cioi?o?i iaiao?aeii ?icaiaeeoe o aiaeiiio ?ic/ei? ei?aiaaeo
aeciiaoaaie?o?a, ui ni?ey? i?aeaeuaiith eiai aoaeoeaiino?.

3. Aeey noeioeyoe?? ?iciiiaeaiiy ? c?inoaiiy a?iiane aiaei?ino?
Dunaliella, i?aeoe/iiai aeaea?aea iaea?aeaiiy (-ea?ioeio, ia?niaeoeaii
aeei?enoiaoaaoe iecuee? eiioeaio?aoe?? ?a/iaeie Aeaeinoei.

4. Iiaeeaia na?aaeiaeua aeey i?i?iuoaaiiy ieeeo, aei neeaaeo yeiai
aoiaeeoue ?a/iaeia Aeaeinoei, aeioe?eueii aeei?enoiaoaaoe aeey ia?ia?oeo
aaaeeicaieethaaieo ?ineei ? a naeaeoe?eii-aaiaoe/ieo aeine?aeaeaiiyo.

5. I?e?iaeio ? iie?ooieoe?iiaeueio ?a/iaeio Aeaeinoei, io?eiaio ?c
ei?aiaaeo aeciiaoaaie?o?a i?i?inoath/iai ian?iiy, aeioe?eueii ?
ia?niaeoeaii aeei?enoiaoaaoe ia o?eueee a n?euenueeiainiiaea?nueeiio
ae?iaieoeoa?, a ? a oa?/ia?e oa i?e?ia?ieia?/i?e i?iieneiaino?,
a?ioaoiieia??, einiaoieia??, iaaeeoeei? oa ?ioeo aaeocyo ia?iaeiiai
ainiiaea?noaa.

NIENIE IAOEIAEO I?AOeUe, IIOAE?EIAAIEO II OAI?
AeENA?OAOe??:

1. Iaoiia A.O., Aiaeeia A.E., Eaiioiae/ A.I., Neey? A.E., Aaeion A.I.
Iieeooieoeeiiaeueiinoue aeeaeiiaoe/aneiai aauanoaa aeeaeinoei //
AeAI Oe?aeiu. -1993. -?11.-N.166-169.

2. Aiaeeia A.E., Iaicyiiaa I.A., Eaiioiae/ A.I. Eeieaeiue ninoaa e
aioeaaeoa?eaeueiay aeoeaiinoue ei?iaauo yeciiaoaaieeoia i?i?inoeia
ioaieoeu // Oeceieiaey ?anoaiee. -1996. -O.43, ?6. -N.920-925.

3. Aiaeeia A.E., Iaicyiiaa I.A., Eaiioiae/ A.I., Aaeion A.I.
Aeeiaieea eeieaeiiai ninoaaa ei?iaauo yeciiaoaaieeoia a i?ioeanna
?acaeoey Triticum aestivum L. // AeAI Oe?aeiu. -1998. -?3.

-N.166-170.

4. Iaoaio 18419 Oe?a?ie, IEA 5A01N1/00. Niin?a iaea?aeaiiy
nooiai i?aia?aoo aeey ia?iaee ian?iiy c aeno?aeoo ian?iiy, ui
i?i?inoa? / Iaoiia A.O., Eaiioiae/ A.I., Eacaeiaa E.?. (UA); Iioae.
25.12.97; Athe.?6.

5. Iaoaio 9982 Oe?a?ie, IEA 5A01I1/04, A01I4/00. Iiaeeaia
na?aaeiaeua Iaoiiaa A.O. aeey i?i?iuoaaiiy ieeeo / Naii?iaeia A.I.,
Iaoiia A.O., Iiniaeia N.A., Eacaeiaa E.?., Eaiioiae/ A.I. (UA);
Iioae.30.09.96; Athe.?3.

6. Iaoaio 8741 Oe?a?ie, IEA 5A01N1/00. Niin?a aeey ia?aaeiin?aii? ia?iee
ian?iiy ca?iiaiaiaeo eoeueoo? / Iaoiia A.O., Een?eue I.N., Oa??i N.A.,
Eaiioiae/ A.I., ?euei?eiaa A.A. (UA); Iioae. 30.09.96; Athe.?3.

7. Iaoaio 2095956 ?innee, IEE 6A01N1/00. Niinia iieo/aiey nooiai
i?aia?aoa aeey ia?aaioee naiyi ec yeno?aeoa i?i?anoathueo naiyi / Iaoiia
A.O., Eaiioiae/ A.I., Eacaeiaa E.?. (RUS); Iioae. 20.11.97; Athe.?32.

8. Iaoiia A.O., Aeueieeiaa A.A., Eaiioiae/ A.I. Aeieiae/anee aeoeaiua
aauanoaa i?i?anoathueo naiyi, niiniau eo iieo/aiey e i?eiaiaiey a
naeueneii oicyenoaa // O?. Iaaeaeoia?. eiio. “Aeioaoiieiaey iieo/aiey
ei?iiaiai aaeea, yeieiae/anee /enouo i?aia?aoia iiauoathueo
o?iaeaeiinoue, i?aieenia, oa?iaioia e aeoaieiia ei?iiaiai iacia/aiey” /
A?ioao-95 /. -Aeiai?iiao?iane: -1995.-N.28.

Eaiioiae/ A.I. “Ei?aiaa? aeciiaoaaie?oe i?i?inoath/iai ian?iiy
ioaieoe?, ?o iaea?aeaiiy, aeine?aeaeaiiy oa aeei?enoaiiy o
?ineeiieoeoa?”.

-?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
n?euenueeiainiiaea?nueeeo iaoe ca niaoe?aeuei?noth 06.01.09 —
?ineeiieoeoai. -?inoeooo ?ineeiieoeoaa ?i. A.ss. TH?(?aa, OAAI, Oa?e?a,
1999.

Aeena?oaoe?y i?enay/aia ?ic?iaoe? oai?aoe/ieo iniia oa i?aeoe/ieo
iaoiae?a iaea?aeaiiy, aeine?aeaeaiiy oa aeei?enoaiiy ei?aiaaeo
aeciiaoaaie?o?a i?i?inoath/iai ian?iiy ioaieoe? o ?ineeiieoeoa?. Ia
iniia? eiiieaeniiai aeine?aeaeaiiy ei?aiaaeo aeciiaoaaie?o?a, ?ic?iaeaia
oaoiieia?y iaea?aeaiiy a?ieia?/ii aeoeaii?, iie?ooieoe?iiaeueii? oa
oaoiieia?/ii? ?a/iaeie Aeaeinoei. Ia ??cieo a?ieia?/ieo ia(?eoao (
ian?ii?, i?e?ii?aai?ciao, ?ineeiao) iieacaia iie?ooieoe?iiaeuei?noue
ae??, aoaeoeai?noue oa ia?niaeoeai?noue aeei?enoaiiy ei?aiaaeo
aeciiaoaaie?o?a o ?ineeiieoeoa? oa ?ioeo

aaeocyo.

Eeth/ia? neiaa: aeciiaoaaie?oe, Aeaeinoei, ?ineeiieoeoai, i?i?inoath/a
ian?iiy, iie?ooieoe?iiaeuei?noue, aeoeai?noue.

Leontovitch V.P. (The roots exometabolities from germinating
wheat seeds, how they are obtained,

observed and used in plant-growing(.

-Manuscript.

Dissertation on the scientific degree competition of candidate of
agricultural sciance in the speciality 06.01.09 — plant-growing.

-Institute of plant-growing in the name of V.Y. Yurieva, UAAN, Kharkov,
1999.

The dissertation is dedicated on the theoretical, preparations and
practical methods to obtain, observe and to the usage of biological
active substances from the roots of germinating wheat seeds
exometabolities. Complex researches on germinating wheat seeds roots
exometabolitits have been worked out. The techno-logy is to obtain
from them biological active polyfunctional dry substance of the main
orgin Allelostim. On different biological objects ( seeds,
microorganisms, plants ) exometabolities of germinating seeds, shows
polifunctional reactions, effectivness and prosptctiveness for their
usage in germinating plant-growing and in many other field.

Key words( exometabolities, seeds, Allelostim, polyfunctional,
plant-growing, active.

Eaiioiae/ A. I. “Ei?iaaua yeciiaoaaieeou i?i?anoathueo naiyi
ioaieoeu, eo iieo/aiea,

enneaaeiaaiea e eniieueciaaiea a ?anoaieaaiaenoaa”.

— ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa
naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe
06.01.09-?anoaieaaiaenoai.-Einoeooo ?anoaieaaiaenoaa ei. A.ss. TH?ueaaa,
OAAI, Oa?ueeia, 1999.

iieeooieoeeiiaeueiinoue eo aeaenoaey. Aaaaeaiea ei?iaauo
yeciiaoaaieeoia a eoeueoo?aeueioth n?aaeo aiaei?inee Dunaliella
noeioee?oao ?aciiiaeaiea e ?ino aeiiannu aiaei?inee, i?aeoe/aneiai
enoi/ieea iieo/aiey ( — ea?ioeia. Enneaaeiaaiea aioeaaeoa?eaeueiie
aeoeaiinoe aeciiaoaaieeoia iieacaee, /oi aaeoa?eoeeaeiinoueth
iaeaaeatho oieueei eeieaeu, auaeaeaiiua ec eo ninoaaa. ?ac?aaioaia
ieoaoaeueiay n?aaea aeey i?i?aueaaiey iueueoeu eoeueoo?iuo ?anoaiee, a
eiiiiiaioiue ninoaa eioi?ie aoiaeyo ei?iaaua yeciiaoaaieeou,
oaaee/eaathuea i?i?anoaiea iueueoeu. ?ac?aaioai niinia aeey
i?aaeiinaaiie aeinoeioeyoeee naiyi ca?iiaiaiauo eoeueoo?. Niaianoiia
eniieueciaaiea ei?iaauo yeciiaoaaieeoia e eeoaaiueeiauo aaeoa?ee ?iaea
Rhizobium aeey i?aae-iinaaiie ia?aaioee naiyi aiaiauo ?anoaiee iiauoaao
o?iaaiue aciooeenaoeee e eo o?iaeaeiinoue. Ia ?acee/iuo aeieiae/aneeo
iauaeoao ( naiaiao, iee?ii?aaieciao, aiaei?ineyo, ?anoaieyo )
iieacaia iieeooieoeeiiaeueiinoue aeaenoaey, yooaeoeaiinoue e
ia?niaeoeaiinoue eniieueciaaiey yeciiaoaaieeoia i?i?anoathueo naiyi
ioaieoeu a ?anoaieaaiaenoaa e ae?oaeo io?aneyo.

Eeth/aaua neiaa: yeciiaoaaieeou, Aeeaeinoei,
iieeooieoeeiiaeueiinoue, ?anoaieaaiaenoai,

i?i?anoathuea naiaia, aeoeaiinoue.

A?aeiia?aeaeueiee ca aeione: eaiaeeaeao
n?euenueeiainiiaea?nueeeo iaoe Ne?ee I.I.

I?aeienaii aei ae?oeo 08.12.98 ?. Oi?iao 60(84/16. Ianya: 1,0
oi.-ae?oe.a?e., 1,0 iae.-aeae. a?e.

Caiiaeaiiy .
Oe?aae 100 i?ei.

Oa?e?anueeee aea?aeaaiee aa?a?iee oi?aa?neoao
?i. A.A.Aeieo/aaaa

312131, Oa?e?a, i/a
“Eiioi?no-1”

Ae?eueieoey iia?aoeaiiai ae?oeo OAeAO

PAGE

1

2

3

4

-2

-8

2

2

5

-1

-7

-9

-6

6

7

8

9

-2

-8

-5

10

?en.2.?ioaineai?noue ?iciiiaeaiiy aoeueai/eiaeo aaeoa??e Rh?zobium o
aiae? (1)

oa o aiaeiiio ?ic/ei? ei?aiaaeo aeciiaoaaie?o?a (2) o
caeaaeiino? a?ae

/ano ?ieoaaoe??.

Oe/aa

11

?en.4 Aieea ??cieo iiaeeaieo noi?oie ia no?oeoo?o caeaiiai
a?ae?iiiiiiai

ei?io ae?iuaiiai ca 7 ae?a ( y/i?iue, ni?o Iaoai?).

1 — ??ooa?a

2 — Aaeue??aaey

3 — Oiaeaiaea-A?iiia

4 — 50% Aeaeinoei

a?ai

12

-9

-7

-6

-9

7 aeiao e?euee?noue ee?oei aoea ieae/a eiio?iey ia 40%. Ei?aiaa?
aeciiaoaaie?oe i?i?inoath/iai ian?iiy i?iyaeythoue noeioethth-/ee ?
?ia?aoth/ee aoaeoe, ye? caeaaeaoue a?ae eiioeaio?aoe?? Aeaeinoe-io
(?en.5). Aeine?aeaeaiiy aeiciai? caeaaeiino? aeicaieeei aeyaeoe
iioeiaeueio noeioethth/o eiioeaio?aoe?th. I?e eiioeaio?aoe??
Aeaeinoeio 1(10 ia/ie niinoa??aa?oueny i?aeneeaiee ??no
a?iiane aiaei?ino?.

?en.5 ?ioaineai?noue ?iciiiaeaiiy aiaei?ino? Dunaliella o eiio?ie? (1)
oa o i?enooiino?

Aeaeinoeio o eiioeaio?aoe?? 1(10 (2), 1(10 (3), 1(10
ia/ie (4).

-9

-7

-6

13

-6

-5

-9

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