Біоекологічні особливості насіннєвості хвойних інтродуцентів на південному сході України: Автореф. дис… канд. біол. наук / О.П. Суслова, УААН. Держ.

AeA?AEAAIEE I?E?ONUeEEE AIOAI?*IEE NAAe

OAAI

NONEIAA Ieaia Iao??aia

OAeE 633.812: 665.527. 64
(477.75)

A?IAEIEIA?*I? INIAEEAINO? IAN?II?AINO? OAIEIEO ?IO?IAeOOeAIO?A IA
I?AAeAIIIIO NOIAe? OE?A?IE

03.00.05 — aioai?ea

AAOI?AOA?AO

aeenna?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

sseoa — 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia o a?aeae?e? aeaiae?ieia?? oa ea?oieea?noaa Aeiiaoeueeiai
aioai?/iiai naaeo IAI Oe?a?ie

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe

Iieyeia Ieaen?e Einoyioeiiae/,

Aeiiaoeueeee aioai?/iee naae IAI
Oe?a?ie,

caa?aeoaa/ a?aeae?eo

Io?oe?ei? iiiiaioe: aeieoi? n?euenueeiainiiaea?nueeeo iaoe

Iieyeia Ieaen?e Oaaei?iae/,

E?einueea a??nueei-e?niaa
iaoeiai-aeine?aeia

noaioe?y Oe?IAeOEAA,

aee?aeoi?

eaiaeeaeao a?ieia?/ieo iaoe

?oaocia ?ai? Ieaenaiae?iae/,

Aea?aeaaiee I?e?onueeee
aioai?/iee naae OAAI,

noa?oee iaoeiaee ni?a?ia?oiee

I?ia?aeia onoaiiaa — Oeaio?aeueiee aioai?/iee naae ?i. I.I. A?eoea

IAI Oe?a?ie, i. Ee?a

Caoeno a?aeaoaeaoueny «07» o?aaiy 1999 ?. i 1300 aiae. ia can?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae 53.369.01 o I?e?onueeiio aioai?/iiio
naaeo OAAI ca aae?anith: 334267, Aaoiiiiia ?anioae?ea E?ei, i. sseoa,
Aea?aeaaiee I?e?onueeee aioai?/iee naae OAAI.

C aeena?oaoe??th iiaeia iciaeiieoenue o iaoeia?e a?ae?ioaoe?
Aea?aeaaiiai I?e?onueeiai aioai?/iiai naaeo OAAI ca aae?anith: 334267,
Aaoiiiiia ?anioae?ea E?ei, i. sseoa, Aea?aeaaiee I?e?onueeee aioai?/iee
naae OAAI.

Aaoi?aoa?ao ?ic?neaiee «30» aa?aciy 1999 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee _______________
N.TH.Naaeiao?nueeee

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

 

Aeooaeuei?noue oaie. ?io?iaeoeoe?y ?ineei o i?iieneia? ?aa?iie ia?
aaaeeeaa oai?aoe/ia oa i?aeoe/ia cia/aiiy. A iiaeo i?e?iaeii-ee?iaoe/ieo
oiiaao ia ?io?iaeoeiaai? ?ineeie aieeaa? oe?eee eiiieaen aeieia?/ieo
oaeoi??a, yeee iii?oii ci?ith? ?o ?icaeoie. I?aaeaiiee No?ae Oe?a?ie
a?ae??ciy?oueny ?yaeii iniaeeainoae, ye? neeaeeny o i?ioean?
aaieia?/iiai oi?ioaaiiy oa ?noi?e/iiai ?icaeoeo. Cia/ia ainiiaea?/a
inai?ii?noue oa?eoi???, iane/aii?noue i?iieneiaeie i?aei?e?inoaaie oa
aeniea aonoioa ianaeaiiy iaeeaaeathoue na?e a?aeaeoie ia i?e?iaei?
oiiae. Aea?aai? ianaaeaeaiiy a Aeiiaan? ? iaeiei c oaeoi??a iioei?caoe??
oaoiiaaiiiai na?aaeiaeua, a noai?aiiy no?eeeo caoenii-aeaei?aoeaieo
ianaaeaeaiue — ? ia?oi/a?aiaei caaaeaiiyi. Iaeiei c iieaciee?a
aaeaioaoe?? ?ineei a oiiaao ?io?iaeoeoe?? ? anooi ?o a aaia?aoeaio
oaco, ii?iaeueia i?ioiaeaeaiiy oeeeeo ?ai?iaeoeoe?? oa oi?ioaaiiy
iiaiioe?iiiai ian?iiy (Iae?ania, 1973; Eioii, 1980; Eoeaaei, 1984).
I?e oeueiio, ian?iiy, ui ooai?eeinue a oiiaao ?io?iaeoeoe??, iiaea
iaoe iciaeaie, iai?aaeai? ia aeaeeaaiiy ?ineei a iiaeo ee?iaoe/ieo
oiiaao oa a oiiaao oaoiiaaiiiai caa?oaeiaiiy. Oae? yeino? iiaeooue
aeieeaoe a iane?aeie iaeioe/iei ?aeiia?iaoe?yi oa iooaoe?yi, /anoioa ?
niaeo? yeeo caeaaeaoue a?ae aa?ioe/ieo oa a?ioe/ieo no?an?a (AEo/aiei,
1995). I?e ?iciiiaeaii? ian?iiyi a?aee?eaa?oueny iiaeeea?noue
cae??ieaiiy ? ia?aaea/? iiaiio iieie?iith oeeo yeinoae /a?ac aaiaoe/io
?ioi?iaoe?th. Ie??i oiai, ian?ii?aa ?iciiiaeaiiy oaieieo ? iaea?eueo
aeaoaaei oa ?acoeueoaoeaiei niiniaii ?iciiiaeaiiy.

Aaeeea e?euee?noue ?io?iaeoeiaaieo oaieieo no?ee? aei i?iieneiaeo
aeeeae?a, oiio ?o aeioe?eueii aeei?enoiaoaaoe aeey icaeaiaiiy i?no.
Oaeei /eiii, aeooaeueiei ? aea/aiiy iniaeeainoae ian?ii?aino? oeeo
?ineei aeey ae??oaiiy oai?aoe/aneeo ieoaiue aaeaioaoe?? ?o i?e
?io?iaeoeoe??, a oaeiae iaea?aeaiiy ian?iiy i?noeaai? ?ai?iaeoeoe?? c
iaoith ?ic?iaee aoaeoeaieo iaoiae?a eoeueoeaoaaiiy oaieieo
?io?iaeooeaio?a aeey aeei?enoaiiy ?o o caeaiiio aoae?aieoeoa? oa
e?niaiio ainiiaea?noa?.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie.  ?iaioa aeeiiaia
o a?aeae?e? aeaiae?ieia?? oa ea?oieea?noaa Aeiiaoeueeiai aioai?/iiai
naaeo IAI Oe?a?ie (AeAN) a ia??iae c 1992 ii 1997 ??. Aeena?oaoe?y
aaciina?aaeiuei iia’ycaia c ieaiiaeie aeine?aeaeaiiyie AeAN c aeaio
athaeaeaoieo oai: 1991 — 1995 ??. — “?ic?iaea oai?aoe/ieo iniia
?io?iaeoeoe?? ?ineei a ?iaeono??aeuei? ?aa?iie c iaoith caaaa/aiiy oa
aeei?enoaiiy ?ineeiieo ?ano?n?a a ia?iaeiiio ainiiaea?noa? oa
iioei?caoe?? oaoiiaaiiiai na?aaeiaeua”, N Ae? 0193U024295, ?ia. N
0296U004885; 1996 -2000 aa. — “Iaoeia? iniiae iia?e?caoe??, caa?aaeaiiy
oa aeei?enoaiiy ?ineei a oiiaao aio?iiiaaiii o?ainoi?iiaaiiai
na?aaeiaeua I?aaeaiiiai Noiaeo Oe?a?ie”, N Ae? 0196U001211.

 Iaoa ? caaea/? aeine?aeaeaiiy.  Iaoa aeine?aeaeaiiy — aeyaeoe
iniaeeaino? aaia?aoeaiiai ?icaeoeo oaieieo, ?io?iaeoeiaaieo ia
I?aaeaiiee No?ae Oe?a?ie oa anoaiiaeoe iiaeeea?noue ?iciiiaeaiiy ?o ca
aeiiiiiaith ian?iiy. Aeey aeinyaiaiiy iinoaaeaii? iaoe aoee iai?/ai?
ianooii? caaaeaiiy:

· ?aeaioeo?eoaaoe ?io?iaeoeiaai? aeaee oaieieo, ui ?inoooue a
eieaeoe?eieo ianaaeaeaiiyo AeAN;

· aeyaeoe oaiiaeieia?/i? iniaeeaino? ?icaeoeo ?ineei;

· aea/eoe iniaeeaino? ii?oiaaiaco aaia?aoeaieo a?oiueie ?
i?e?inii?iaaiaco;

· anoaiiaeoe n?iee “oea?o?iiy” oa ye?noue ieeeo oaieieo,
?io?iaeoeiaaieo ia I?aaeaiiee No?ae Oe?a?ie;

· i?iaiae?coaaoe aeeiai?eo aec??aaiiy ian?iiy;

· anoaiiaeoe ye?ni? iieacieee ian?iiy;

· aeaoe ioe?ieo ?ai?iaeoeoeaii? caeaoiino? iaea?eueo ia?niaeoeaieo
aeae?a oaieieo ?io?iaeooeaio?a.

Iaoeiaa iiaecia io?eiaieo ?acoeueoao?a.  Aia?oa aea/aii a?iaeieia?/i?
iniaeeaino? ian?ii?aino? ?yaeo aeae?a oaieieo, ?io?iaeoeiaa-ieo ia
I?aaeaiiiio Noiae? Oe?a?ie ( Pinus pallasiana  D. Don,
Pseudo-tsuga menziesi  (Mirb.) Franco, Larix dahurica  Turcz. ex
Trautv.,  Platycladus orientalis  (L.) Franco, Thuja occidentalis
L., Juniperus virgi-niana  L., Chamaecyparis lawsoniana  (A. Murr.)
Parl., Taxus canadensis Marsch. Anoaiiaeaii ?yae ci?i a aaia?aoeaiiio
?icaeoeo oaieieo: ci?uaiiy oa?i?i?a anooio oa o?eaaeino? oac ii?oiaaiaco
aaia?aoeaieo a?oiueie oa cieaeaiiy ye?nieo iieaciee?a ian?iiy noiniaii
?o i?e?iaeieo a?aae?a, a oaeiae ii?ooaiiy a i?ioean? iaeico. Oe? ci?ie
? ?aaeoe??th i?enoinoaaiiy aei ci?iaieo i?e?iaeii-aeieia?/ieo oiia
c?inoaiiy. Iiaeaia ioe?iea ?ai?iaeoeoeaii? caeaoiino? aeae?a, ye?
aea/aeenue.

I?aeoe/ia cia/aiiy io?eiaieo ?acoeueoao?a.  Ia i?aenoaa? io?eia-ieo
?acoeueoao?a anoaiiaeaii iiaeeea?noue oa aeioe?euei?noue aeei?enoaiiy
ian?iiy i?noeaai? ?ai?iaeoeoe?? aeey ianiaiai ?iciiiaeaiiy ? aaaaeaiiy a
eoeueoo?o oe?iieo ?io?iaeoeiaaieo aeae?a oaieieo. ?ic?iaeai?
?aeiiaiaeaoe?? c ae?iuoaaiiy iinaaeeiaiai iaoa??aeo Picea pungens,
Pinus pallasiana, Juniperus virginiana,  Platycladus orientalis
oa  Thuja occidentalis c ian?iiy i?noeaai? ?ai?iaeoeoe??. ?aeiiaiaeaoe??
ia?aaeaii Aea?aeaaiiio eiioiaeueiiio i?aei?e?inoao caeaiiai
aoae?aieoeoaa i. Aeiiaoeueea.

Iniaenoee aianie caeiaoaa/a. Aeena?oaoe?eia ?iaioa aeeiiaia aaoi?ii
i?ae ea??aieoeoaii ae.a.i. I.E. Iieyeiaa ? ? naiino?eiei
aeine?aeaeai-iyi caeiaoaa/a. Aaoi?ii ?ic?iaeaia i?ia?aia
aenia?eiaioaeueieo ?ia?o, aeeiiaii aeine?aeaeaiiy, i?iaaaeaii aiae?c ?
ocaaaeueiaiiy io?eiaiiy aenia-?eiaioaeueieo aeaieo. ?acoeueoaoe
aeine?aeaeaiue a?aeia?aaeai? a aeena?oaoe?? oa ioae?eaoe?yo. A iaoeiaeo

?iaioao, iaienaieo o ni?aaaoi?noa?, aaoi? ? iiaiii?aaiei /eaiii oai?/iai
eieaeoeao. I?aaa ni?aaaoi??a ia ii?ooai?.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??.  ?acoeueoaoe aeine?aeaeaiue
aeieeaaeai? ia eiioa?aioe?yo iieiaeeo a/aieo oa niaoe?ae?no?a «No/ani?
i?iaeaie aioai?ee» (Aeiiaoeuee, 1995), «Aeooaeueiua i?iaeaiu aioaieee e
yeieiaee» (Oa?e?a, 1996), «I?iieneiaa aioai?ea: noai oa ia?niaeoeae
?icaeoeo» (Aeiiaoeuee, 1998). Iaoa??aee oa iniiai? iieiaeaiiy ?iaioe
i?ieoee ai?iaaoe?th ia can?aeaiiyo a?aeae?eo aeaiae?ieia?? oa ia
can?aeaiiyo a/aii? ?aaee AeAN IAI Oe?a?ie.

Ioae?eaoe??.  Ca oaiith aeena?oaoe?? iioae?eiaaii 10 iaoeiaeo ?ia?o.

No?oeoo?a oa ianya ?iaioe. No?oeoo?a aeena?oaoe?? neeaaeaoueny ?c
anooio, oanoe ?icae?e?a, i?aeoe/ieo ?aeiiaiaeaoe?e, aeniiae?a, nieneo
aeei?enoaieo e?oa?aoo?ieo aeaea?ae. Iiaiee ianya aeena?oaoe?? noaiiaeoue
157 noi?., c yeeo ia oaaeeoe? i?eiaaea? 24, ?ethno?aoe?? — 10, nienie
ia?aoiao? 190 e?oa?aoo?ieo aeaea?ae — 20 noi?.

INIIAIEE CI?NO ?IAIOE

?ICAe?E 1

O?CEEI-AAIA?AO?*I? OIIAE I?AAeAIIIAI NOIAeO OE?A?IE

I?aaeaiiee No?ae Oe?a?ie a iaaeao Aeiiaoeueei? oa Eoaainueei? iaeanoae
caeia? ieiuo 53,2 oen. ei2 ? a?ae??ciy?oueny cia/iith aaieia?/iith oa
aaiii?-oieia?/iith ??ciiiai?oi?noth. Iniiaieie aeaiaioaie aaieia?/ii?
no?oeoo-?e ? Aeiiaoeueeee e?yae, I?eacianueea aeni/eia oa I?eacianueeee
ieceiiee noai. Aaieia?/ieie oa aaiii?oieia?/ieie i?ioeanaie ?
aea?o?thaaiiy, oi?ioaaiiy aetha?th, ianiaee ?oo iooeiai iaoa??aeo ia
noeeao, cieaaiiy oa ?icieaaiiy, a?o?iaa a?ic?y oa aeoioeyoe?y, ea?noia?
yaeua, no/ani? oaeoi-i?/i? ?ooe, a oaeiae i?ioeane ?aeue?oiooai?aiiy,
iia’ycai? c ainiiaea?nueeith ae?yeuei?noth ethaeeie.

Ca ee?iaoe/iei ?aeiioaaiiyi Oe?a?ie ?aa?ii a?aeianaii aei
eiioe-iaioaeueiie noaiiai? iaeano? iii??ieo oe?io (Aaae/aiei e ae?.,
1984). Ee?iao eiioeiaioaeueiee, c ae?aaeaieie iinooeeai-nooia?eieie
yaeua-ie oa ia??aiii??iei ?iciiae?eii iiaae?a i?ioyaii ?ieo (460-540 ii)
. Iae-oieiaei?oee i?nyoeue — n?/aiue (-0,6 — 7,8iN), iaeoaie?oee —
eeiaiue (20,9 — 22,9iN). Aaaaoaoe?eiee ia??iae — 200 -215 aeiae.

Ca?aeii noaie a?oioiaiai ?aeiioaaiiy Oe?a?ie, oa?eoi??y
a?aeiineoue-ny aei ciie noai?a c /a?iicaiaie cae/aeieie, i?aaeaiieie oa
i?oeaeey?ii-ea?aiiaoieie (Aoean ii/a Oe?aeineie NN?, 1979).
A?ae?ieia?/ia n?oea i?aaenoaaeaia ??/eaie, ica?aie, i?aecaiieie aiaeaie,
aiaeinoiaeuaie oa noaaeaie. Aeiiaan — nai???aeiee o?ceei-aaia?ao?/iee
?aeii, o yeiio ae?y aeno?aiaeueieo i?e?iaeieo oaeoi??a iaoiiaeth?
oi?ioaaiiy aei?noeeo e?ni?ineeiieo oiia. Aea aenieee ??aaiue
?iaeth/ino? ciiaeueiiai a?oioo, aaeeeee oaieiaee aaeain (2800-3200iN
aeoeaieo oaiia?aoo?) oa caaeia?eueia aieiaicaaacia/ai?noue aeicaieythoue
eoeueoeaoaaoe a oiiaao I?aaeaiiiai Noiaeo Oe?a?ie oe?eee ?yae
?io?iaeoeiaaieo aeae?a aea?aaieo ?ineei.

?ICAe?E 2

NOENEEE IA?EN ?NOI??? ?IO?IAeOEOe?? OAIEIEO IA I?AAeAIIEE NO?Ae
OE?A?IE

Iaoeiaee i?aeo?ae aei i?iaeaie ?io?iaeoeoe?? oaieieo ia I?aaeaiiee No?ae
Oe?a?ie ii/aany c 1843 ?ieo, c canioaaiiy ia?oiai noaiiaiai
Aaeeei-Aiaaeieuenueeiai aeine?aeieoeueeiai e?nieoeoaa. Cia/io ?ieue o
?iciianthaeaeaii? oaieieo a?ae?a?aee Ia??oiieuenueea, Noa?ia?euenueea
e?ni-eoeueoo?i? ae?eyiee a oaeiae Aaioeoaanueeee ia?e, a yeiio aoei
aenaaeaeaii 26 aeae?a oa oi?i oaieieo (?iaaioa??caoe?y 1936-1939 ??.).

A Aeiiaoeueeiio aioai?/iiio naaeo IAI Oe?a?ie c?a?aia eieaeoe?y oaieieo,
yea ia?aoiao? 98 aeae?a ? oi?i ?io?iaeoeiaaieo aeae?a. Iaeiae, o
i?nueeeo ianaaeaeaiiyo Aeiiaano eoeueoeao?oueny anueiai 14 aeae?a oa 3
oi?ie oaieieo aecio?a, ui iaaeinoaoiuei aeey caaeiaieaiiy iio?aa
caeaiiai aoae?aieoeoaa (A.E.Iieyeia, E.A.Iaethaei, A.I.Oa?aa?ei, 1992
a.; A.A.Aii-oeia, 1993 a.). O ca’yceo c oeei aeieeea iaiao?aei?noue
aaaaeaiiy o neeaae ??cieo eaoaai??e ianaaeaeaiue iaea?eueo
ia?niaeoeaieo aeae?a oaieieo ?ineei, ye? i?ioiaeyoue ?io?iaeoeoe?eia
aei?iaoaaiiy a Aeiiaoeueeiio aioai?/iiio naaeo.

?ICAe?E 3

IA’?EOE OA IAOIAeE AeINE?AeAEAIUe

3.1. Ia’?eoe aeine?aeaeaiue

Ia’?eoe aeine?aeaeaiue — ?io?iaeoeiaai? aeaee oaieieo, ui i?aaenoaaeai?
a eieaeoe?eiiio oiiae? AeAN, ye? aeinyaee ?ai?iaeoeoe?eiiai a?eo,
anooieee a aaia?aoeaio oaco oa i?iaeoeothoue ian?iiy. Aeaee, ye?
aea/aeenue, a?aeiinyoueny aei ?iaeei: Cupressaceae — Chamaecyparis
lawsoniana (A. Mirr.) Parl., Juniperus virginiana L., Platycladus
orientalis (L.) Franco, Thuja occidentalis L.; Pinaceae — Larix
dahurica Turcz. ex Trautv., Picea pungens Engelm., Pinus pallasiana
D.Don, Pseudotsuga menziesi (Mirb.) Franco; Taxaceae — Taxus
canadensis Marsch.

3.2. Iaoiaee aeine?aeaeaiue

Aea/aiiy a?iaeieia?/ieo iniaeeainoae ?inoo oa aaia?aoeaiiai ?icaeoeo
oaieieo aeeiiai? iaoiaeaie, ye? canoiniaothoueny a ?io?iaeoeoe?ii-ieo
aeine?aeaeaiiyo. Oaii?eoieeo naciiiiai ?icaeoeo aea/aee iaoiaeeeith,
yea ?ic?iaeaia N.?. Eociaoeiaei oa ?.I. ?oaociaei a Aea?aeaaiiio
I?e?onueeiio aioai?/iiio naaeo OAAI (1973). Ceiino?ee?noue ?ineei
anoaiiaethaaee ca naieaaeueiith oeaeith I.I. Eaio?aa (1974) a?aeiia?aeii
yei? iieacieeii ceiino?eeino? ? nooi?iue iioeiaeaeaiiy iaaecaiii?
/anoeie aea?aa i?ney ia?aceioaaiiy; iinooino?ee?noue ioe?ithaaee
a?coaeueii ca naieaaeueiith oeaeith ?.O. A?eoeaiei (1953).
Aeaei?aoeai?noue ?ineei, a iniiao yei? iieeaaeaii ni?eeiaiiy ?ineei ye
aeaiaioa naaeiai? a?o?oaeoo?e — ca /ioe?eaaeueiith oeaeith I.A.
Eio?eiaie oa I.N. A?a/ei (1969). ?yni?noue caeeaaeaieo no?ia?e?a
aecia/aee ca oeaeith, cai?iiiiiaaiith A.I. Eiaociaei (1962).
I?e?inii?iaaiac oa ?icaeoie ieeeo aea/aee ia iino?eieo i?aia?aoao,
i?eaioiaeaieo ca caaaeueiii?eeiyoith o oeeoieia?? iaoiaeeeith (Iaooaaa,
1988), aeeoo?caeao-i?noue ieeeo — iaoiaeii i?i?iuoaaiiy eiai ia
niiaeeaieo na?aaeiaeuao, oa?oeeuei?noue — ie?aooaaiiyi ieeeiaeo ca?iei
aoeaoiea?i?iii. Iioaioe?e-io ian?ii?ao i?iaeoeoeai?noue aea/aee ca
iaoiaeeeith, ?ic?iaeaiith ?.I. ?oaociaei, O.I. Eaaiiii oa E.O. Neeiiiith
a Aea?aeaaiiio I?e?onueeiio aioai?/iiio naaeo OAAI (1986). ?aaeueio
ian?ii?ao i?iaeoeoeai?noue aecia/aee a i?ioeaioao ca ni?aa?aeiioaiiyi
iioaioe?eii? ian?ii?ai? i?iaeoeoeaiino? aei e?eueeino? aec??aoiai
ian?iiy. sse?noue ian?iiy a i?ioean? ?o oi?ioaaiiy aecia/aee i?e
ia?aaeyae? ?ic??caiiai ian?iiy i?ae i?e?ineiiii IAN-10. Iano 1000 oooe
ian?iei aecia/aee oeyoii ia?a?aooaaiiy iane 100 oooe aeia?eueii
aea?aiiai ian?iiy o aeanyoee?aoi?e iiaoi?iino? (Eeuoe N.N., 1991).
Noaoenoe/io ia?iaeo io?eiaieo ?acoeueoao?a i?iaiaeeee iaoiaeaie
aa??aoe?eii? noaoenoeee (Caeoeaa, 1983) ca aeiiiiiaith IBM PC
“Statistical Graphic System” (version: 2.9, 1985, 1986, STST,
Statistical Graphics Corporation, EXEC*U*STAT’’).

?ICAe?E 4

NACIIIEE ?ICAEOIE OAIEIEO ?IO?IAeOOeAIO?A IA I?AAeAIIIIO NOIAe? OE?A?IE

Aea/aii oa?i?ie anooio oa o?eaae?noue oaiieia?/ieo oac. Anoaiiaeaii
cia/i? eieeaaiiy oa?i?i?a ii/aoeo oa o?eaaeino? oaiioac o an?o aeae?a,
ye? aea/aeenue, o ??ci? ?iee, ui iaoiiaeaii i?ieea?noth ee?iaoe/ieo
oiia ceiiaiai oa aaniyiiai ia??iae?a. An? aeaee ca oa?i?iaie aaaaoaoe??
a?aeiia?aeathoue aaaaoaoe?eiiio ia??iaeo I?aaeaiiiai Noiaeo Oe?a?ie,
oa?aeoa?ecothoueny nai?/aniei i?eieiaiiyi ?inoo oai? oa iaaii?a aei /ano
iioieiaeaiiy.

Aeine?aeaeaiiy iieacaee, ui an? aea/ai? aeaee ia?aiinyoue ceiiaee
ia??iae aac iioeiaeaeaiue ? ioe?iai? I oa II aaeaie ceiino?eeino?. A
iaea?eueo noai?? ceie o Platycladus orientalis oa Thuja occidentalis
iaia?cathoue e?ioe? iaaii?a inoaiiueiai ?ieo aaaaoaoe??, a iaeii??/i?
iaaiie Chamaecyparis lawsoniana — iaia?cathoue ia anth aeiaaeeio.

I?ae ae??th e?oiuei? iinooe iaea?eueo iioeiaeaeo?oueny Chamaecyparis
lawsoniana (a ia??iae iinooe a’yia oaiy), o Taxus canadensis a iniaeeai
iinooeea? ?iee ieaei? oai?iee iaaii?a i?eeiathoue in?ii?e eie?? ?
iiaaeathoue. Inoaii? aeaee aeineoue iinooino?ee?, iioeiaeaeaiue o ieo ia
ciaeaeaii.

An? aeaee ioe?iai? aeuei aaeii aeaei?aoeaiino? — Ae4. Nthaee ae
a?aeianaii ? Larix dahurica, yea ia ? ceiiaicaeaiei aeaeii, aea ia?
i?ea?iaeueiee aaa?oon e?iie oa na?oeee eie?? a?eie.

?ICAe?E 5

AAIA?AOEAIEE ?ICAEOIE OAIEIEO

?IO?IAeOOeAIO?A IA I?AAeAIIIIO NOIAe? OE?A?IE

5.1. Aaia?aoeaiee ?icaeoie ye e?eoa??e aaeaioaoe?? ?ineei i?e

?io?iaeoeoe?? (iaeyae e?oa?aoo?e)

Aaia?aoeaiee ?icaeoie ?, c iaeiiai aieo, aieiaiith eaieith a ooai?aii?
aaeaioeaieo i?enoinoaaiue, ui aeieeathoue ye iane?aeie ii?ooaiue iaeico,
a c ae?oaiai — iaeiei c e?eoa???a aaeaioaoe?? ?ineei a iiaeo oiiaao,
ine?eueee ian?ii?a?noue caaa?oo? an? noaae?? iioiaaiaco oa caaacia/o?
iiyao iiaiai iieie?iiy (Aa?i?ei, 1973; Iae?ania, 1973; Eioii, 1980;
Eoeaaei, 1984; AEo/aiei, 1995).

O ?icae?e? iieacaii, ui i?e ?io?iaeoeoe?? a aaia?aoeai?e noa?? ?ineei
iiaeooue i?ioiaeeoe ??ci? ii?ooaiiy, i?e/eiith yeeo ? iia?
i?e?iaeii-aeieia?/i? oiiae (?oaocia, Neeiiiay, *aaioa?ue, 1992; Iacaia,
1995; Eriksson, 1968). I?eaaaeaii e?oa?aoo?i? aeai? a?aeiinii aea/aiiy
oeeeeo ?ai?iaeoeoe?? oaieieo a ??cieo i?noeac?inoaiiyo
(Aeaenaiae?ianeee, 1971; Eiaocia, 1972, 1974; Eociaoeia, 1984; O?aiei,
1986; O?aoueyeiaa, 1990, 1991; *aaioa?ue, 1995; Kormutak, Hudak, 1994;
Owens, 1972; Salaj, Kormutak, 1994). A?aei?/aii, ui ia I?aaeaiiiio
Noiae? Oe?a?ie aaia?aoeai? i?ioeane oaieieo ia aea/aeeny.

5.2. A?e anooio oaieieo ?io?iaeooeaio?a a aaia?aoeaio oaco ?

?yni?noue no?ia?e?a

Anoaiiaeaii, ui a?e ii/aoeo ?ai?iaeoeoe?? oaieieo a?aeiinii a?eo
eoeuei?iaoe?? iioi/iiai i?e?inoo a aenioo iiaeeeai ?iciiae?eeoe ia o?e
a?oie (oaae. 1).

A?aei?/aia a?aei?ii?noue a ?yniino? /ieia?/eo oa ae?ii/eo no?iaeeia: o
a?eueoino? aea/aieo aeae?a na?aaei?e aae /ieia?/iai “oea?o?iiy” aeua
ae?ii/iai, ui iiynith?oueny a?ieia?/ieie iniaeeainoyie aeae?a. Ie??i
oiai, anoaiiaeaii ??cieoeth o oi?ioaaii? no?ia?e?a ii ?ieai, ia ui
aieeaathoue iiaiaei? oiiae in?iiuei-ceiiaiai ia??iaeo, yeee ia?aaeo?
“oea?o?iith”. Oae, ceie 1994 oa 1996 ??. c ii?icaie aei -22,5iN ?
-21,5iN a?aeiia?aeii oa i?ci?ie caii?iceaie aei e?ioey aa?aciy (-1,6iN)
eiia??ii aecaaee cia/i? iioeiaeaeaiiy oa caaeaaeue no?ia?e?a.

5.3. Ii?oiaaiac aaia?aoeaieo a?oiueie oaieieo ?io?iaeooeaio?a,

ye? c?inoathoue a AeAN

A ?acoeueoao? aiae?co io?eiaieo aeaieo a?aei?/aii, ui aaia?aoeai?
a?oiueee oaieieo caeeaaeathoueny ca ??e aei “oea?o?iiy” oa cae?i/othoue
na?e ?icaeoie aei /ano cieaeaiiy oaiia?aoo? (ca aeiyoeii Pinus
pallasiana, o yei? iiaia oi?ioaaiiy iaaano?ia?eia caaa?oo?oueny aaniith
ianooiiiai ?ieo). I?e ii??aiyii? aeaieo c e?oa?aoo?ieie anoaiiaeaii, ui
ia I?aaeaiiiio Noiae? Oe?a?ie a?aeaoaa?oueny ci?uaiiy oa?i?i?a
i?ioiaeaeaiiy

Oaaeeoey 1

A?e anooio oaieieo ?io?iaeooeaio?a AeAN a aaia?aoeaio oaco

Aeae A?e aea?aa, ?ieeA?e ia?oi? iiyae
no?ia?e?a, ?ieeA?e eoeuei?iaoe?? iioi/iiai i?e?inoo a aenioo, ?iee I
a?oiaLarix dahurica 19 15 18Pseudotsuga
menziesi (Mirb.) Franco 29 24 25Juniperus
virginiana L. 26 14 18Thuja occidentalis
L. 26 15 17
II a?oiaPicea pungens Engelm. 31
20 20Chamaecyparis lawsoniana (A. Mirr.) Parl 12
10 10
III a?oiaPinus pallasiana D.Don 27 18
15Platycladus orientalis(L.) Franco. 26 16
14Taxus canadensis Marsch. 17 12 8iniiaieo
oac ii?oiaaiaco aaia?aoeaieo a?oiueie a?aeiinii i?e?iaeieo
i?noeac?inoaiue aeae?a, ye? aea/aeenue, ui na?ae/eoue i?i i?enoinoaaiiy
?ineei aei ee?iaoe/ieo oiia ?aa?iio ?io?iaeoeoe??.

5.4. I?e?inii?iaaiac oaieieo ?io?iaeooeaio?a

I?e aea/aii? i?e?inii?iaaiaco anoaiiaeaii, ui nooi?iue ?icaeoeo
nii?iaaiii? oeaieie o aeae?a, ye? aea/aeenue, aei ia??iaeo ceiiaiai
niieith ciaoiaeeoueny ia ??cieo noaae?yo, i?ioa, noaae?? ?icaeoeo
nii?iaaiii? oeaieie a?aeiia?aeathoue aiaeia?/iei noaae?yi a i?e?iaeieo
i?noeac?inoaiiyo. I?ae /an iaeico a i?e?inii?ioeeoao a?aeaoaathoueny
ii?ooaiiy, o ?acoeueoao? yeeo ooai?ththoueny aiiiaeuei? i?e?inii?e.
Iaea?eueoa e?euee?noue aeaeeieo iaeioe/ieo ii?ooaiue a?aei?/aia o aeae?a
c i?ioiaeaeaieai iaeico ?aiiueith aaniith (Pseudotsuga menziesi,
Platycladus orientalis, Thuja occidentalis). Na?aae aeaeeieo ii?ooaiue
anoaiiaeai? aac’yaea?i? i?e?inii?e, i?e?inii?e c oeeoiieaciith, ui
a?ae?eoea a?ae iaieiiee, ae?aaee oa o?eaaee (oaae. 2). E?euee?noue
ii?iaeueii ?icaeiooeo i?e?inii? eieeaa?oueny a?ae 84% o Platycladus
orientalis aei 92% o Juniperus virginiana. Oea na?ae/eoue i?i
a?aeiia?aei?noue oiia ?io?iaeoeoe?? ii?iaeueiiio i?ioiaeaeaiith
i?e?inii?iaaiaco oaieieo.

5.5. Oa?oeeuei?noue oa aeeoo?caeaoi?noue ieeeo

Aeey anoaiiaeaiiy i?e/ei ooai?aiiy ionoiai ian?iiy aoea ia?aa??aia
oa?oeeuei?noue oa aeeoo?caeaoi?noue ieeeo oaieieo. Aea/ai? aeaee
oa?aeoa?ecothoueny aeinoaoiuei aenieei i?ioeaioii oa?oeeueieo ieeeiaeo
ca?iei (oaae. 3). Anoaiiaeaia ?iaeea?aeoaeueia i?ieea?noue yeino? ieeeo
iaoiiaeaia a?ieia?/ieie iniaeeainoyie ?ineei. ??/ia i?ieea?noue —
iiaiaeieie oiiaaie a ia??iae i?e?inii?iaaiaco. O i?ai?/iiaia?eeainueeeo
aeae?a, ui ?io?iaeoeiaai? ia I?aaeaiiiio Noiae? Oe?a?ie aiie?ooaea
??/ii? i?ieeaino? yeino? ieeeo iacia/ia (oaae. 3), ui na?ae/eoue i?i
ni?eyoeea? iaoai?ieia?/i? oiiae a ia??iae i?ioiaeaeaiiy
i?e?inii?iaaiaco o oeeo aeae?a.

5.6. Aeeiai?ea aec??aaiiy ian?iiy

A oiae? aia??iaaiaco o aea/aieo aeae?a oaieieo ca?eueoo?oueny
e?euee?noue ionoiai ian?iiy (?en. 1). Caaeaaeue ca?iaee?a
a?aeaoaa?oueny ia ?aii?o noaae?yo aia??iaaiaca o Pseudotsuga menziesi;
/anoeiai ia ?aii?o, /anoeiai ia i?ci?o — o Pinus pallasiana; ia i?ci?o
noaae?yo — o Platycladus orientalis, Thuja occidentalis, Juniperus
virginiana. Iaea?eueoa e?euee?noue ionoiai ian?iiy ciaeaeaia o
Chamaecyparis lawsoniana, Larix dahurica oa Pseudotsuga menziesi.
I?e/eiith oeueiai iiaea aooe ae?y eaoaeueieo aai?a oa iaa?aeiia?aei?noue
i?ae ca?iaeeaie, ye? ?icaeaathoueny oa ae?ii/ei aaiaoio?oii.

Oaaeeoey 2

 Iii?oi? ii?ooaiiy iaeico a i?e?inii?ioeeoao oaieieo, ?io?iaeoeiaaieo
ia I?aaeaiiiio Noiae? Oe?a?ie (1996 ?.)

E?euee?noue Ii?iaeue-
Aiiiaeuei? i?e?inii?eAeaeaeine?aeaeai-i? i?e?i-oo.%, I maacuyaea?i?
ae?aaee, i?e?inii?e eo i?e?i- nii?, oo.nii?e, oo. i?e?inii?e,
%, I mo??aaee, %, I mc oeeoiieac-iith, ui a?ae-ae?eeeany a?ae
iaieiiee, %, I mPicea pungens Engelm. 720 648 72 9,9 0,64
8,2 0,10 0,4 0,01 1,4 0,03Pinus pallasiana D.Don 847 780
76 7,8 0,84 6,0 0,09 0,4 0,02 1,5 0,03Pseudotsuga menziesi
(Mirb.) Franco 333 295 38 10,8 1,85 7,5 0,12 0
3,9 0,08Chamaecyparis lawsoniana (A. Mirr.) Parl. 602 552 50
8,3 0,54 2,7 0,05 0 5,6 0,07Juniperus virginiana L.
746 686 60 8,0 0,42 3,3 0,04 0,5 0,03 4,8
0,03Platycladus orientalis (L.) Franco 831 701 130 15,6 0,43
10,3 0,14 0,3 0,01 5,0 0,06Thuja occidentalis L. 850 725 125
14,5 0,99 11,0 0,18 0 3,5 0,02Taxus canadensis Marsch.
757 682 75 10,0 0,49 6,3 0,15 0,3 0,01 3,3 0,08

?ICAe?E 6

IAN?II?AA ?ICIIIAEAIIss OAIEIEO

?IO?IAeOOeAIO?A IA I?AAeAIIIIO NOIAe? OE?A?IE

6.1. Ian?ii?aa i?iaeoeoeai?noue

Aeey ioe?iee ?ai?iaeoeoeaiie caeaoiino? oaieieo aoei anoaiiaeaii
iioaioe?eio oa ?aaeueio ian?ii?ao i?iaeoeoeai?noue. Anoaiiaeaii, ui ia
I?aaeaiiiio Noiae? Oe?a?ie ?io?iaeoeiaai? aeaee oi?iothoue iaaano?ia?ee
c aenieei i?ioeaioii oa?oeeueieo eiiieaen?a eonie c ian?iieie
ca?iaeeaie, ui ? aeieacii ?o ii?iaeueiiai ?icaeoeo (io 47% o
Pseudotsuga menziesi aei 95% o Juniperus pallasiana). ?aaeueia
ian?ii?aa i?iaeoeoeai?noue aeinyaa? 53% -85 %. Anoaiiaeaia noaoenoe/ii
aeinoia??iee i?yiee ca’ycie ?aaeueii? ian?ii?ai? i?iaeoeoeaiino? c
e?euee?noth ian?iieo ca?iaee?a, ye? caeeaeenue, oa aeeoo?caeaoi?noth
ieeeo.

Oaaeeoey 3

Oa?oeeuei?noue ieeeiaeo ca?ai oaieieo ?io?iaeooeaio?a ia I?aaeaiiiio
Noiae? Oe?a?ie

Aeae Oa?oeeuei?noue ieeeo, % 1994 a.
1995 a. 1996 a.Larix dahurica Turcz. ex Trautv. — 79
2,78 -Picea pungens Engelm. — — 78
2,54Pinus pallasiana D.Don80 1,65 83 2,09 60
1,56Pseudotsuga menziesi (Mirb.) Franco83 1,32 83 1,87 80
1,65Chamaecyparis lawsoniana (A. Mirr.) Parl65 1,05 — 70
1,56Juniperus virginiana L.93 1,45 93 1,07 90 1,83Platycladus
orientalis (L.) Franco.65 1,23 86 2,76 63 1,54Thuja
occidentalis L.57 1,63 81 1,79 57 1,81Taxus canadensis
Marsch.53 0,96 62 0,97 47 1,03

6.2. Iin?ai? yeino? ian?iiy

I?e ian?ii?aiio ?iciiiaeaii? aeae?a, ye? aea/aeenue, anoaiiaeai?
iin?ai? yeino? ian?iiy: iiaiica?ieno?noue oa a?oioiaa noiae?noue.
Ian?iiy Chamaecyparis lawsoniana, Larix dahurica e Pseudotsuga menziesi
oa?aeoa?ecothoueny iecueeei i?ioeaioii iiaiica?ienoino? oa noiaeino?.
?aooa aeae?a oaieieo oi?iothoue iiaiioe?iia noiaea ian?iiy. A?aei?/aia
??cieoey a e?eueeino? noiaeiai ian?iiy, yea oi?io?oueny, ia I?aaeaiiiio
Noiae? Oe?a?ie oa a i?e?iaeieo a?aaeao. I?e/eiith oeueiai ? iaaaoeaiee
aieea oiia ?io?iaeoeoe?? ia i?ioeane ?ai?iaeoeoe??.

 

I?AEOE*I? ?AEIIAIAeAOe??

1. Iaeiei ?c cania?a ?iciiiaeaiiy ia I?aaeaiiiio Noiae? Oe?a?ie
 Picea pungens, Pinus pallasiana, Pseudotsuga menziesi,  Juniperus
virginiana, Platycladus orientalis, Thuja occidentalis oa Taxus
canadensis ? iin?a ian?iiy i?noeaai? ?ai?iaeoeoe??.

2. Iin?a ian?iiy  Picea pungens, Pinus pallasiana, Pseudotsuga
menziesi, Platycladus orientalis, Thuja occidentalis iaiao?aeii
i?iaiaeeoe a aaniyiee ia??iae, a Juniperus virginiana oa Taxus
nanadensis — ainaie.

3. No?aoeo?eaoe?y ian?iiy Platycladus orientalis oa
Thuja occidentalis ai?iaeiaae iaeiiai i?nyoey i?e oaiia?aoo?? 0-5iN
iie?auo? ?o noiae?noue.

4. Iaea?eueo aeioe?eueiith aeeaeiith caae?eo ian?iiy Pinus pallasiana,
 Pseudotsuga menziesi, Platycladus orientalis ? 4 — 5 ci;
 Juniperus virginiana oa Taxus canadensis — 2 — 3 ni; Thuja
 occidentalis — 0,5-1 ni.

5. I?e ae?iuoaaii? ian?iiy ia I?aaeaiiiio Noiae? Oe?a?ie iaiao?aeii
a?aoiaoaaoe, ui ia ?o c??no, noai oa no?ee?noue aei iani?eyoeeaeo
oaeoi??a na?aaeiaeua (iniaeeai aei iecueeeo oaiia?aoo?) cia/ii aieeaa?
aeeiiaiiy iniiaieo aa?ioaoi?/ieo i?eeii?a.

6. A oiiaao I?aaeaiiiai Noiaeo Oe?a?ie iaeiie?oi? n?yioe? ?inus
?allasiana, ?seudotsuga menziesi oa Thuja occidentalis aceieo ia
iaia?cathoue, Platycladus orientalis — /anoeiai iaia?cathoue, oiio a
ceiiaee ia??iae ?o ?aeiiaiaeo?oueny ae?eaaoe oa?ii eenoy aei 10-15 ni.
Iaeiie?oi? n?yioe? Taxus canadensis iioa?iathoue a?ae e?oi?o iinoo,
oiio iaiao?aeii ?o i?eo?iyoe oa iieeaaoe a iniaeeai iinooeeaee e?oi?e
ia??iae.

AENIIAEE

1. Ca ?acoeueoaoaie e?eoe/iiai aea/aiiy ?io?iaeooee?iaaieo a
Aeiiaoeueeiio aioai?/iiio naaeo IAI Oe?a?ie aeae?a aieiian?iieo ?ineei
anoaiiaeaii, /oi eieaeoe?y ia?aoiao? 98 aeae?a oa oi?i, ye?
a?aeiinyoueny aei 14 ?iae?a oa 6 ?iaeei. C ieo 60 aeae?a anooieee a
aaia?aoeaioth oaco.

2. Ia I?aaeaiiiio Noiae? Oe?a?ie o ?io?iaeoeiaaieo aeae?a oaieieo
a?aei?/aii ci?io oa?i?i?a i?ioiaeaeaiiy ii?oiaaiaco aaia?aoeaieo
a?oiueie a?aeiinii i?e?iaeieo i?noeac?inoaiue, ui na?ae/eoue i?i
canoiniaio ?aaeoe?th ?ineei ia iia? i?e?iaeii-aeieia?/i? oiiae
c?inoaiiy.

3. I?e oi?ioaaii? ieeeiaeo ca?ai a i?e?inii?ioeeoao oaieieo
anoaiiaeaii ?yae ii?ooaiue, iaoiiaeaieo aieeaii i?e?iaeii-ee?iaoe/ieo
oaeoi??a ia i?ioeann iaeico. Iane?aeeii oeueiai ? ooai?aiiy noa?eeueieo
ieeeiaeo ca?ai. Aei oaeeo ii?ooaiue iaeaaeaoue aac’yaea?i? i?e?inii?e,
i?e?inii?e c oeeoiieaciith, yea a?aeae?eeeanue a?ae oeeoiieacie,
ae?aaee ? o??aaee. I?ioa, i?ioeaio ii?iaeueii ?icaeiooeo i?e?inii? o
an?o aeae?a caeeoa?oueny aenieei — a?ae 84% o Platycladus orientalis
 aei 92% o  Pinus pallasiana.

4. Aiaeiaa i?ieea?noue aeeoo?caeaoiino? ieeeo aeae?a, ye? aea/aeenue,
coiiaeaia iniaeeainoyie iaoai?ieia?/ieo oiia. Aiie?ooaea eieeaaiue o
aeayeeo aeae?a aeinyaa? cia/ieo aaee/ei. Iaea?eueoa aa??aaaeuei?noue
niinoa??aa?oueny o Platycladus orientalis — 63 — 87% aeeoo?caeaoii?
ieeeo a?ae caaaeueii? e?eueeino? ieeeo, yeee noi?ioaaany, Thuja
occidentalis — 57 — 81% e  Pinus pallasiana  — 60-81%. Iaeiaio? ci?ie
aeeoo?caeaoiino? ii ?ieai a?aei?/ai? o Pseudotsuga menziesi  81-83%.

5. Anoaiiaeaii, ui ia I?aaeaiiiio Noiae? Oe?a?ie o ?io?iaeoeiaaieo
aeae?a oaieieo iioaioe?eia ian?ii?aa i?iaeoeoeai?noue ci?ith?oueny a?ae
66% o Pseudotsuga menziesi  aei 90% o  Juniperus virginiana; ?aaeueia —
io 53% aei 84% a?aeiia?aeii. ?aaeueia ian?ii?aa i?iaeoeoeai?noue
ciaoiaeeoueny a i?yi?e caeaaeiino? a?ae aeeoo?caeaoiino? ieeeo.

6. O ia??iae oi?ioaaiiy ian?iiy a oiae? aia??iaaiaco o oaieieo
ca?eueoo?oueny e?euee?noue ii?iaei?o ian?iei, i?e/eiith /iai ? ae?y
eaoaeueieo aai?a oa iaa?aeiia?aei?noue i?ae ca?iaeeii, ui
?icaeaa?oueny, oa ae?ii/ei aaiaoio?oii. I?e caaeaae? ca?iaee?a ia
?aii?o noaae?yo “aiaeinia?i” aeaaaia?o? oa oi?iothoueny ii?iaei?
ian?ieie (?seudotsuga menziesi -70-80%, Chamaecyparis lawsoniana
89-92%); ia i?ci?o — “aiaeinia?i” ia aeaaaia?o?, a ooai?ththoueny
aeiiaiai? ian?ieie aac ca?iaee?a (Thuja occidentalis — 28-90%, Juniperus
virginiana — 70-98%, Taxus canadensis — 90-95%).

7. I?ae aieeaii i?e?iaeii-ee?iaoe/ieo oiia ia I?aaeaiiiio Noiae?
Oe?a?ie o ?io?iaeoeiaaieo aeae?a oaieieo oi?io?oueny ian?iiy,
noiae?noue yeeo ieua, i?ae o i?e?iaeieo a?aaeao: Larix dahurica —
o Aeiiaoeueeiio aioai?/iiio naaeo IAI Oe?a?ie 1-2%, o i?e?iaeiiio
a?aae? — 55%; Juniperus virginiana — 30% e 87% a?aeiia?aeii;
Chamaecyparis lawsoniana — ia aaouee?auei? — 57%, a Aeiiaoeueeiio
aioai?/iiio naaeo IAI Oe?a?ie — ia noiaeyoue; Pinus ?allasiana ia
I?aaeaiiiio Noiae? Oe?a?ie — 46-56% oa 30-80% a E?eio.

8. Oa?aeoa? naciiiiai ?icaeoeo, iinooino?ee?noue, ceiino?ee?noue,
aeaei?aoeai?noue oa iniaeeaino? aaia?aoeaiiai oeeeeo aea/aieo aeae?a
I?ai?/ii? Aia?eee, No?aeii? Ac??, E?eio oa Aeaeaeiai Noiaeo na?ae/aoue
i?i ia?niaeoeai?noue ?o canoinoaaiiy a caeaiiio aoae?aieoeoa?, e?niaiio
ainiiaea?noa? oa iieacaoeniiio e?ni?icaaaeaii?.

9. Aeey ianiaiai ?iciiiaeaiiy oa aaaaeaiiy a eoeueoo?o oe?iieo
?io?iaeooee?iaaieo aeae?a oaieieo (Picea pungens, Pinus pallasiana,
Juniperus virginiana, Platycladus orientalis oa Thuja incidentalis)
aeioe?eueii aeei?enoiaoaaoe ian?iiy i?noeaai? ?ai?iaeoeoe??.

NIENIE IIOAE?EIAAIEO CA OAIITH AeENA?OAOe?? ?IA?O

1. Noneiaa A.I., Iieyeia A.E. Iniaaiiinoe naciiiiai ?acaeoey oaieiuo
a Aeiiaoeeii aioaie/aneii naaeo //Eio?iaeoeoeey e
aeeeeiaoecaoeey ?anoaiee. — 1995. — Aui. 24. — N.21-25.

2. Noneiaa I.I. Aeaye iniaeeaino oeeeeo aaia?aoeaiiai ?icaeoeo
oaieieo a oiiaao Iaaeaiiiai Noiaeo Oe?aie
//Oe?. aioai. aeo?i. -1996. — N5. — N.619-621.

3. Noneiaa A.I. Naiaiioaiea oaieiuo eio?iaeooeaioia eae e?eoa?ee
aaeaioaoeee eo a iiauo oneiaeyo //I?iaeaie aenia?eiaioaeueii aioaiee oa
aeieia ?ineei. — Eea: Iaoe. aeoiea, 1997. — N.272-274.

4. Noneiaa A.I. ?ai?iaeoeoeaiay niiniaiinoue oaieiuo eio?iaeooeaioia
eae e?eoa?ee eo onoie/eainoe // Ioi?iia aaiioiiaeo ?ineei a Oe?a?i?.
Oac. aeiiia?aeae iaoe. eiio., E?eaee ??a, o?aaaiue, 1994 ?. —
Aeiiaoeuee: AeIIII, 1994. — N.183-184.

5. Noneiaa I.I. Aaia?aoeaiee ?icaeoie oaieieo ia I?aaeaiiiio
Noiae? Oe?a?ie //Aioaie/aneea naaeu — oeaio?u nio?aiaiey aeieiae/aneiai
?aciiia?acey ie?iaie oei?u. Oac. aeiee. nannee Niaaoa aioai. naaeia
Oe?aeiu. — 13-16 ethiy 1995 a., E?ui. — sseoa: Ia/aoiue oeao Ieeeoneiai
aioaie/aneiai naaea, 1995. — N.196-197.

6. Noneiaa I.I. Aeaye? ii?ooaiiy oiaeo aia??iaaiaco oaieieo
?io?iaeooeaio?a ia I?aaeaiiiio Noiae? Oe?a?ie //Aea/aiiy iioiaaiaco
i?e?iaeieo oa eoeueoo?ieo oei? o aioai?/ieo caeeaaeao ?a?ac??. Iao. 8-?
I?aeia?iaeii? eiioa?aioe??. Ee?a, 1995. — N. 145.

7. Noneiaa A.I. Ea/anoai naiyi niniu e?uineie ia thai-ainoiea
Oe?aeiu //Aeooaeueiua aii?inu aioaieee e yeieiaee. Oac. aeiee. eiio.
iieiaeuo o/aiuo e niaoeeaeenoia. Oa?ueeia, 5-7 ethiy, 1996. —
Oa?ueeia: AI Iieea?aoe/aneay oe?ia «I?eioae», 1996. — N.104.

8. Noneiaa I.I. ?icaeoie ian?iiy Pseudotsuga menziesi (Mirb.)
Franco a oiiaao I?aaeaiiiai Noiaeo Oe?a?ie //I?iaeaie aioai?ee ?
i?eieia?? ia ii?ic? o?aoueiai oeny/ie?ooy. Iao. X c’?caeo Oe?.
aioai. oiaa?enoaa. — Iieoaaa, 22-23 o?aaiy, 1997 ?. — Ee?a-Iieoaaa:
OIA «I?aeia?iaeia o?iainiaa aaaioe?y», 1997. — N.155.

9. Noneiaa A.I., Iieyeia A.E. Ea/anoai naiyi eio?iaeooee?iaaiiuo aeaeia
aee a oaoiiaaiiuo oneiaeyo Aeiiaanna //I?iaeaiu aeaiae?ieiaee,
oeaaoiaiaenoaa, ieiaeiaiaenoaa. Iao. V Iaaeaeoia?iaeiie eiio. 6-10
ieoya?y 1997 a. — sseoa: Ia/aoiue oeao Ieeeoneiai aioaie/aneiai naaea,
1997. — *. I. — N.151-152.

10. Noneiaa A.I. Iniaaiiinoe ii?oiaaiaca aaia?aoeaiuo ii/ae Larix
dahurica Turcz. ex Trautv. ia thai-ainoiea Oe?aeiu //I?iieneiaa
aioai?ea: noai oa ia?niaeoeae ?icaeoeo. Iao. o?aouei? i?aeia?iaeii?
iaoe. eiio. — Aeiiaoeuee: Aaainoai «Ioeueoei?ann», 1998. — N.217-218.

AIIOAOe??

Noneiaa I.I. A?iaeieia?/i? iniaeeaino? ian?ii?aino? oaieieo
?io?iaeooeaio?a ia I?aaeaiiiio Noiae? Oe?a?ie. — ?oeiien.

Ae?na?oaoeey ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo
iaoe ca niaoe?aeuei?noth 03.00.05 — aioai?ea. — Aea?aeaaiee I?e?onueeee
aioai?/iee naae OAAI, sseoa, 1999.

I?aaenoaaeai? ?acoeueoaoe eiiieaeniiai aea/aiiy iniaeeainoae
ian?ii?aino? oaieieo ?io?iaeooeaio?a ia I?aaeaiiiio Noiae? Oe?a?ie:
oaii?eoi?ee, oa?i?i anooio aei aaia?aoeaii? oace, iniaeeainoae
ii?oiaaiaco, oa?i?i?a caieeaiiy oa yeino? ieeeo. Anoaiiaeai?
a?aeiia?aei? ?aaeoe?? aaia?aoeaiiai ?icaeoeo ?ineei ia ci?iai? oiiae
c?inoaiiy. Ia i?aenoaa? iaea?aeaieo ?acoeueoao?a aeaia ioe?iea
?ai?iaeoeoe?eii? caeaoiino? ?yaeo ?io?iaeoeiaaieo aeae?a oaieieo.
?ic?iaeai? iaoiaee/i? ?aeiiaiaeaoe?? ii ae?iuoaaiith iinaaeeiaiai
iaoa??aeo c ian?iiy i?noeaai? ?ai?iaeoeoe??.

Eeth/ia? neiaa: ?io?iaeoeoe?y, I?aaeaiiee No?ae Oe?a?ie, aaia?aoeaiee
?icaeoie, oaii?eoi?ea, ii?oiaaiac, i?e?inii?iaaiac, ieeie,
oa?oeeuei?noue, aia??iaaiac.

Noneiaa A.I. Aeiyeieiae/aneea iniaaiiinoe naiaiioaiey oaieiuo
eio?iaeooeaioia ia thai-ainoiea Oe?aeiu. — ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.05 — aioaieea. — Ainoaea?noaaiiue
Ieeeoneee aioaie/aneee naae OAAI, sseoa, 1999.

Aeenna?oaoeey iinayuaia aii?inai aaia?aoeaiiai ?acaeoey oaieiuo,
eio?iaeooee?iaaiiuo ia thai-ainoie Oe?aeiu. Ianiio?y ia aeanoeea
eani?anoeoaeueiua oneiaey ?aaeiia, o?iaaiue ieiaei?iaeey ciiaeueiuo
ii/a, aunieee oaieiaie aaeain e oaeiaeaoai?eoaeueiay
aeaaiiaania/aiiinoue iicaieytho eoeueoeae?iaaoue ia thai-ainoiea Oe?aeiu
?yae eio?iaeooee?iaaiiuo oaieiuo. Eco/aiea eo ?ai?iaeoeoeaiiai oeeeea
niinianoaoao ?aoaieth aii?inia iieo/aiey iieiioeaiiuo naiyi n oeaeueth
?ac?aaioee yooaeoeaiuo iaoiaeia eoeueoeae?iaaiey ?anoaiee naiaiiui
iooai.

Iauaeoaie enneaaeiaaiee auee oaieiua, i?aaenoaaeaiiua a eieeaeoeeiiiii
oiiaea Aeiiaoeeiai aioaie/aneiai naaea IAI Oe?aeiu: Chamaecyparis
lawsoniana (A. Mirr.) Parl., Juniperus virginiana L., Platycladus
orientalis (L.) Franco, Thuja occidentalis L.; Pinaceae — Larix
dahurica Turcz. ex Trautv., Picea pungens Engelm., Pinus pallasiana
D.Don, Pseudotsuga menziesi (Mirb.) Franco; Taxaceae — Taxus
canadensis Marsch.

I?e ioeaiea ioaaoiie ?aaeoeee ?anoaiee ia oaeoi?u aiaoiae n?aaeu eco/aee
eo naciiiia ?acaeoea. Onoaiiaeaii eciaiaiea n?ieia ia/aea e
i?iaeieaeeoaeueiinoe oaiioac o anao aeaeia a ?acee/iua aiaeu, /oi
iaoneiaeaii eciai/eainoueth iiaiaeiuo oneiaee. Ana aeaeu
oa?aeoa?ecothony naiaa?aiaiiui i?ae?auaieai ?inoa oaie e iiaaaia ei
a?aiaie ianooieaiey iioieiaeaiey. Eco/aiiua aeaeu ceiinoieee,
ia?aceiiauaatho aac iia?aaeaeaiee, ioeaiaiu I e II aaeeaie ceiinoieeinoe
(eneeth/aiea ninoaaee Chamaecyparis lawsoniana e Platycladus
orientalis). Aicaeaenoaeth eaoieo canoo iaeaieaa iiaeaa?aeaiu
Chamaecyparis lawsoniana e Taxus canadensis. Inoaeueiua aeaeu aiieia
canooionoie/eau.

Onoaiiaeaii, /oi a oaco ?ai?iaeoeoeee oaieiua anooiatho a ?aciii
aic?anoa ioiineoaeueii eoeueieiaoeee oaeouaai i?e?inoa a aunioo e
?acaeaeaiu ia o?e a?oiiu. Ia?ooaiey iaaeaeo aaaaoaoeaiui e aaia?aoeaiui
?acaeoeai ioia/aiu o Pinus pallasiana e Taxus canadensis, o eioi?uo
ia?aay caeeaaeea aaia?aoeaiuo ii/ae i?ienoiaeeo ?aiueoa, /ai ?anoaiea
aeinoeaaao aic?anoa eoeueieiaoeee oaeouaai i?e?inoa a aunioo. Iiaiaeiua
oneiaey aeeytho ia iaeeea no?iaeeia: ceiu 1994 e 1996 aa. n ii?icaie aei
-22,5iN e -21,5iN aucaaee aeaaeue cia/eoaeueiiai eiee/anoaa no?iaeeia. O
anao aeaeia i?aiaeaaeaao “ioaeneia oeaaoaiea”, /oi iaoneiaeaii
aeieiae/aneeie iniaaiiinoyie aeaeia.

I?e eco/aiee ii?oiaaiaca aaia?aoeaiuo ii/ae ioia/aii niauaiea n?ieia
ia/aea e i?iaeieaeeoaeueiinoe oac ioiineoaeueii i?e?iaeiuo ianoiiaeoaiee
eio?iaeooee?iaaiiuo aeaeia. Aeaiiue oaeo yaeyaony naeaeaoaeuenoaii
i?eniiniaeoaeueiie ?aaeoeee ?anoaiee ia eeeiaoe/aneea oneiaey ?aaeiia
eio?iaeoeoeee. Nii?iaaiiay oeaiue a i?ioeanna iee?inii?iaaiaca o
eco/aiiuo aeaeia ?acaeaaaony oae aea, eae e a i?e?iaeiuo a?aaeao; c?aeay
iueueoea o Pseudotsuga menziesi, Chamaecyparis lawsoniana, Juniperus
virginiana, Platycladus orientalis, Thuja occidentalis, Taxus canadensis
i?aaenoaaeaia iee?inii?aie, Picea pungens e Pinus pallasiana —
aeaooeeaoiaiui aaiaoioeoii. Ai a?aiy i?ioiaeaeaiey iaeica i?ienoiaeyo
ia?ooaiey, yaeythueany neaaenoaeai io?eoeaoaeueiiai aicaeaenoaey ia
?anoaiey i?e?iaeii-eeeiaoe/aneeo yeieiae/aneeo oneiaee. Iaia?oaeaiu
aacuyaea?iua iee?inii?u, aeeaaea, o?eaaeu, iee?inii?u n ioioaaeoae io
iaiei/ee oeeoiieaciie. Iaeaieueoaa eiee/anoai ia?ooaiee onoaiiaeaii o
aeaeia, iaeic eioi?uo i?ienoiaeeo ?aiiae aaniie i?e iaonoie/eaie
iiaiaea (Pseudotsuga menziesi, Platycladus orientalis, Thuja
occidentalis). Ana aeaeu oa?aeoa?ecothony oi?ie?iaaieai aunieiai
i?ioeaioa oa?oeeueiie iueueoeu (io 84% o Platycladus orientalis aei 92%
o Juniperus virginiana).

Ia thai-ainoiea Oe?aeiu o eio?iaeooee?iaaiiuo aeaeia oaieiuo
oi?ie?othony iaaano?iaeeu n aunieei i?ioeaioii oa?oeeueiuo eiiieaenia
/aooe n naiyii/eaie, /oi naeaeaoaeuenoaoao i ii?iaeueiii i?ioiaeaeaiee
iaaanii?iaaiaca.

I?e eco/aiee aeeiaieee nic?aaaiey naiyi onoaiiaeaii, /oi a oiaea
yia?eiaaiaca o anao aeaeia oaaee/eaaaony eiee/anoai ionouo naiyi.
I?e/eiie yoiai iiaeao auoue aeaenoaea eaoaeueiuo aaiia e ianiioaaonoaea
iaaeaeo ?acaeaathueiny ca?iaeuoai e aeaineei aaiaoioeoii. I?e aeaaee
ca?iaeuoae ia ?aiieo noaaeeyo “yiaeinia?i” aeaaaia?e?oao e oi?ie?othony
ionoua naiaia (?seudotsuga menziesi -70-80%, Chamaecyparis lawsoniana
89-92%); ia iicaeieo — “yiaeinia?i” ia aeaaaia?e?oao, a ia?acothony
auiieiaiiua naiaia aac ca?iaeuoae (Thuja occidentalis — 28-90%,
Juniperus virginiana — 70-98%, Taxus canadensis — 90-95%).

Aiaeec ea/anoaaiiuo iieacaoaeae naiyi iieacae, /oi ia thai-ainoiea
Oe?aeiu o eio?iaeooee?iaaiiuo aeaeia oaieiuo oi?ie?othony naiaia,
anoiaeanoue eioi?uo ieaea, /ai a i?e?iaeiuo a?aaeao: Larix
dahurica — a Aeiiaoeeii aioaie/aneii naaeo IAI Oe?aeiu 1-2%, a
i?e?iaeiii a?aaea — 55%; Juniperus virginiana — 30% e 87%
niioaaonoaaiii; Chamaecyparis lawsoniana — ia ?iaeeia — 57%, a
Aeiiaoeeii aioaie/aneii naaeo IAI Oe?aeiu — ia anoiaeyo; Pinus
?allasiana ia thai-ainoiea Oe?aeiu — 46-56% e 30-80% a E?uio.

Ia iniiaa iieo/aiiuo aeaiiuo aeaia ioeaiea ?ai?iaeoeoeaiie niiniaiinoe
?yaea eio?iaeooee?iaaiiuo aeaeia oaieiuo. ?ac?aaioaiu iaoiaee/aneea
?aeiiaiaeaoeee ii au?aueaaieth iinaaei/iiai iaoa?eaea ec naiyi ianoiie
?ai?iaeoeoeee.

Eeth/aaua neiaa: eio?iaeoeoeey, thai-ainoie Oe?aeiu, aaia?aoeaiia
?acaeoea, oaii?eoieea, ii?oiaaiac, iee?inii?iaaiac, iueueoea,
oa?oeeueiinoue, yia?eiaaiac.

Suslova E.P. Bioecological peculiarities of Coniferous introduced plants
seminification in the south-east of Ukraine. — Manuscript.

Thesis for conferring on a degree of a candidate of biological sciences,
specification 03.00.05 — botany. — The State Nikita Botanical Gardens,
Ukr. Agr. Acad. Sci., Jalta, 1999.

The results of the complex study of peculiarities of conifers
seminification in the south-east of Ukraine are presented:
phenorythmics, terms of entering a generative phase, peculiarities of
morphogenesis of generative buds and microsporogenesis, terms of
pollination and quality of pollen. The response of generative
development of plants on the changed conditions of growth has been
defermined. On the basis of the data obtained the assessment of the
reproductive ability of a number of introduced species of conifers has
been given. The methodic guidelines on the growing of plant material of
seeds of local reproduction have been developed.

The key words: introduction, the south-east of Ukraine, generative
development, morphogenesis, microsporogenesis, pollen, fertility,
embriogenesis.

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