9

EE?ANUeEEE OI?AA?NEOAO

?IAI? OA?ANA OAA*AIEA

?AIAOTHE

IEAEN?E AIAOIE?EIAE*

OAeE 574.55:574.64

CAEAAEI?NOUe I?IAeOEOe?EII-

AIA?AAOE*IEO IIEACIEE?A A?INENOAI

A?Ae ??AIss CAA?OAeIAIIss

NA?AAeIAEUA AAAEEEIE IAOAEAIE

03.00.16. AEIEIA?ss

AAOI?AOA?AO

AeENA?OAOe?? IA CAeIAOOOss IAOEIAIAI NOOIAIss

EAIAeEAeAOA A?IEIA?*IEO IAOE

EE?A-1999

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia ia eaoaae?? o?c?ieia?? oa aeieia?? ?ineei

Ee?anueeiai oi?aa?neoaoo ?iai? Oa?ana Oaa/aiea

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, i?ioani?

Ion??iei Ieeiea Ieeieaeiae/,

Ee?anueeee oi?aa?neoao ?iai? Oa?ana

Oaa/aiea, caa?aeoaa/ eaoaae?ith

o?c?ieia?? oa aeieia?? ?ineei.

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani?

A?aa?inueeee Eaa Iaaeiae/,

“Aeae?iaeieiae/aneee aeo?iae”,

canooiiee aieiaiiai ?aaeaeoi?a c

iaoeiaeo ieoaiue;

aeieoi? a?ieia?/ieo iaoe, i?ioani?

Ao/aoeueeee Eaii?ae Iao?iae/,

Ee?anueeee oi?aa?neoao ?iai? Oa?ana

Oaa/aiea, i?ia?aeiee iaoeiaee

ni?a?ia?oiee eaai?aoi??? aeieia?? ?

oieneeieia?? a?ieia?/iiai oaeoeueoaoo.

I?ia?aeia onoaiiaa: Oeaio?aeueiee aioai?/iee naae IAI

Oe?a?ie ?iai? I.I.A?eoea.

Caoeno aeena?oaoe?? a?aeaoaeaoueny “14” aa?aniy 1999 ?. i 12-00

aiaeei? ia can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee Ae 26.001.24

a?ieia?/iiai oaeoeueoaoo Ee?anueeiai oi?aa?neoaoo ?iai? Oa?ana

Oaa/aiea ca aae?anith: Ee?a-127, i?iniaeo aeaaeai?ea Aeooeiaa,

2, a?ieia?/iee o-o, aoae. 215.

Iiooiaa aae?ana: 252033, Ee?a-33, aoe. Aieiaeeie?nueea,64.

C aeena?oaoe??th iiaeia iciaeiieoeny a a?ae?ioaoe? oi?aa?neoaoo:

aoe. Aieiaeeie?nueea, 58.

Aaoi?aoa?ao ?ic?neaii “25” /a?aiy 1999 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee A?aeii I.A.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. Ci?ia noaio aeiae?eey oa caa?oaeiaiiy a?inoa?e
c?inoa? c eiaeiei ?ieii, niinoa??aa?oueny ??cea iia??oaiiy oiia
?nioaaiiy a?eueoino? aeae?a ?ineei ? oaa?ei, aeiaaeathoue eaiee
o?io?/ieo eaioetha?a, ii?ooothoueny ca`ycee a aeinenoaiao, iaaea?
i?iaeoeoeai?noue a?ioeaiic?a. Iaeiei c /eiiee?a i?ioeano iia??oaiiy ?
oiene/i? caa?oaeieee, caeaa?eueoiai aio?iiiaaiiiai iioiaeaeaiiy. Na?aae
ieo i?ia?aeia i?noea iaeaaeeoue aaaeeei iaoaeai.

Aaeaeaaoia ioe?iea noaio aeiae?eey ? aaaeeeaei ieoaiiyi, iia`ycaiei ?c
eiai caa?oaeiaiiyi. O?aaeeoe?eiee niin?a ioe?iee caa?oaeiaiiy —
ii??aiyiiy eiioeaio?aoe?? oieneeaioo a na?aaeiaeu? c aaee/eiaie a?aie/ii
i?eionoeieo eiioeaio?aoe?e (AIE) oieneeaioo a no/anieo oiiaao ia
caaaeaee aea? a?aeiia?aeio ea?oeio aca?iei i?aai?ci?a c aeiae?eeyi. Ca
oaeeo ianoaaei ioe?iea yeino? na?aaeiaeua naia ca noaiii a?inenoai
iiaea aeyaeoenue a?eueo aaeaeith [A?aa?inueeee,95]. Iaei??th c
aeanoeainoae, ui oa?aeoa?eco? noai aoaeue-yeeo a?inenoai, ? caeaoi?noue
aei noai?aiiy a?ii?iaeoeoe?? — i?iaeoeoeai?noue. I?iaeoeoea?noue ?
aeanoea?noth aeeai? iaoa???, ui i?aaenoaaeaia ia an?o ??aiyo ??
i?aai?caoe??, e??i oiai, aiia oa?aeoa?ia ye aeey aaoio?ioieo, oae ? aeey
aaoa?io?ioieo i?aai?ci?a. ?noioii, ui i?iaeoeoeai?noue ? oaeiae
?ioaa?aeueiith oa?aeoa?enoeeith aeeaeo ia`?eo?a, e?ioeaaith ooieoe??th
?o aeeoo?ae?yeueiino?, yea a?aoiao? aanue eiiieaen ? aaeaioeaieo, ?
eiiiainaoi?ieo a?aeiinei c on?ia oaeoi?aie na?aaeiaeua.

A oe?eiio iieacaii, ui caaaeueiith ?aaeoe??th aeinenoaie ia oiene/iee
i?an ? cieaeaiiy ?? i?iaeoeoeaiino? ? cnoa i?iaeoeoe?eii-aeano?oeoe?eieo
i?ioean?a o a?e ca?eueoaiiy aio?ii?? [A?aaeineee,88]. Aea io?eiai?
?acoeueoaoe ua ia aaeaeiny iia`ycaoe c iaea?eueo aaaeeeaeie
i?iaeoeoe?eii-aia?aaoe/ieie iieacieeaie, oaeeie ye /enoa oa aaeiaa
ieoiia i?iaeoeoe?y, aoaeoeai?noue (/enoa oa aaeiaa) eiiaa?ooaaiiy ei?io,
aoaeoeai?noue aeei?enoaiiy oioineioaoe/ii aeoeaii? ?aae?aoe??, ui ia
aeicaiey? aeeoe ia e?euee?niee ??aaiue ioe?iee aeioieneeieia?/ii?
neooaoe?? ca i?iaeoeoe?eieie ia?aiao?aie a?inenoai ??ciiai ??aiy.

Ca`ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. Aeine?aeaeaiiy,
i?iaaaeai? ii oai? aeena?oaoe?eii? ?iaioe, ? /anoeiith o?aoueiai aoaio
iaoeiai-aeine?aeii? ?iaioe “Aea/aiiy aieeao aeciaaiieo /eiiee?a ia
i?iaeoeoe?eii-aia?aaoe/i? ia?aiao?e aenia?eiaioaeueii? iiaeae?
?ineeiiiai oa?oiiaaiiy”, ui aoiaeeoue aei eiiieaenii? iaoeiai? i?ia?aie
“Aa?ii?iaeeiiieaen” ye aea?aeathaeaeaoia iaoeiai-aeine?aeia ?iaioa ca
oaiith “Aeciaaiia ?iaeoeoe?y aaeaioeaieo ?aaeoe?e aeey noaa?e?caoe??
i?iaeoeoeaiino? ca?iiaeo eoeueoo? a iani?eyoeeaeo aeieia?/ieo oiiaao”,
OC IAe? ?97/090, aea?aeaaiee ?a?no?aoe?eiee ?0197U003136, ui
aeeiio?oueny a eaai?aoi??? o?c?ieia?/ieo iniia i?iaeoeoeaiino? ?ineei
eaoaae?e o?c?ieia?? oa aeieia?? ?ineei a?ieia?/iiai oaeoeueoaoo
Ee?anueeiai oi?aa?neoaoo ?iai? Oa?ana Oaa/aiea; oa oaie I?i?noa?noaa c
ieoaiue iaoee ? oaoiieia?e Oe?a?ie “I?iaeoeoeai?noue aeinenoai ??ciiai
nooiaiy caa?oaeiaiiy oa e?euee?ni? ?ioaa?aeuei? e?eoa??? ioe?iee yeino?
na?aaeiaeua ca i?iaeoeoe?eii-aia?aaoe/ieie ia?aiao?aie a?ieia?/ieo
nenoai” ?6.4/249, aea?aeaaiee ?a?no?aoe?eiee ?0197U017066.

Iaoa ? caaea/? aeine?aeaeaiiy. Iaoa aeaii? ?iaioe — aeyaeoe a
iiaeaeueieo aeine?aeao e?euee?ni? oa ye?ni? ci?ie iieaciee?a
a?ii?iaeoeoe?eiiai i?ioeano ca oiia caa?oaeiaiiy na?aaeiaeua
oieneeaioaie (ia i?eeeaae? aaaeeeo iaoae?a), iia`ycaoe iniiai?
i?iaeoeoe?eii-aia?aaoe/i? iieacieee a?inenoai oa ?o ci?io c ??aiai
caa?oaeiaiiy na?aaeiaeua.

Aeey aeinyaiaiiy oe??? iaoe iio??aii aoei ae??oeoe ianooii? caaea/?:

—i?iaanoe eiiieaeniee ii??aiyeueiee aiae?c ci?i
i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a aaoio?ioieo ? aaoa?io?ioieo
i?aai?ci?a ca oiia ??ciiai nooiaiy caa?oaeiaiino? na?aaeiaeua;

—aeyaeoe e?euee?nio caeaaei?noue i?ae i?iaeoeoe?eii-aia?aaoe/ieie
ia?aiao?aie oa ??aiai caa?oaeiaiiy na?aaeiaeua;

—ciaeoe /ooeea? ? ?ioi?iaoeai? i?iaeoeoe?eii-aia?aaoe/i?
iieacieee-?iaeeeaoi?e, ui iaea?eueo aaeaeaaoii oa?aeoa?ecothoue
aeioieneeieia?/io neooaoe?th;

—?aeiiaiaeoaaoe iaea?eueo /ooeea? ? ?ioi?iaoeai?
i?iaeoeoe?eii-aia?aaoe/i? ia?aiao?e a?inenoai aeey iiaeeeaiai
aeei?enoaiiy a aeioieneeieia?/ieo aeine?aeaeaiiyo oa canoinoaaiiy a
nenoai? aeieia?/iiai iii?oi?eiao.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Iaoeiaa iiaecia aeaii? ?iaioe
iieyaa? a oiio, ui aia?oa:

—i?iaaaeaii nenoaii? aeine?aeaeaiiy ci?i oe?eiai ?yaeo iniiaieo
i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a aaoio?ioieo oa aaoa?io?ioieo
i?aai?ci?a a caeaaeiino? a?ae ??aiy caa?oaeiaiiy na?aaeiaeua aaaeeeie
iaoaeaie ia i?eeeaae? ?ii?a naeioeth oa i?eaeth;

—i?inoaaeaii aeeiai?eo ci?ie i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a o
/an? ca oiia o?ii?/iiai cao?o?iiy;

—aeey ioe?iee ??aiy oiene/iino? canoiniaaii ?ioaa?aeuei? iieacieee
i?iaeoeoe?eiiai i?ioeano (?ioaa?aeueiee iieaciee i?iaeoeoeaiino? oa
iieaciee iioeiaeueiino?); cai?iiiiiaaii ?ioaa?aeueiee iieaciee
i?iaeoeoeaiino? (?II) aeey aaoio?ioieo i?aai?ci?a;

—cai?iiiiiaaii iaoaiaoe/io iiaeaeue ae?? oieneeaio?a ia
a?ii?iaeoeoeai?noue iieiae? ?ea, ui ia?aaoaathoue ca oiia caa?oaeiaiiy
(aiae?oe/iee iien ci?ie i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a a?ae
eiioeaio?aoe?? oa /ano ae?? oieneeaioy).

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. Io?eiai? aeai? uiaei ci?i
i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a ??cieo a?inenoai a caeaaeiino?
a?ae ??aiy caa?oaeiaiiy na?aaeiaeua aaaeeeie iaoaeaie aeathoue
iiaeeea?noue:

—iaa?oiooaaoe i?iaiicoaaiiy ci?i, ui caciathoue i?e?iaei? oa?oiiaaiiy
i?ae aieeaii caa?oaeiee?a;

—a?aooaaoe aieea ?ii?a naeioeth oa i?eaeth i?e ?ic?aooieao
aia?aaoe/iiai aaeaino aiaeiei, ye? caciaee oiene/iiai aieeao;

—ii/aoe noai?aiiy aaieo aeaieo ii ci?i? i?iaeoeoe?eii-aia?aaoe/ieo
iieaciee?a iniiaieo a?oi a?ae?ia?iio?a i?ae aieeaii oieneeaio?a a
nenoai? aeieia?/iiai iii?oi?eiao;

—aeei?enoaoe iae/ooeea?o? i?iaeoeoe?eii-aia?aaoe/i? iieacieee aeey
ae?aaiinooaaiiy aeioieneeieia?/ii? neooaoe?? o aiaeieiao.

Iniaenoee aianie caeiaoaa/a. Aaoi?ii ?iaioe inoaenoi aoee iinoaaeai? ?
i?iaaaeai? aeine?aee ii aea/aiith aieeao oieneeaio?a ia
i?iaeoeoe?eii-aia?aaoe/i? iieacieee ??cieo i?aai?ci?a; cai?iiiiiaaii
iaoaiaoe/io iiaeaeue ae?? oieneeaio?a ia aaoa?io?ioi? i?aai?cie;
?ioa?i?aoiaaii ?acoeueoaoe aieeao oieneeaio?a ia aaoio?ioi? i?aai?cie
oa cai?iiiiiaaii ?ioaa?aeueiee iieaciee i?iaeoeoeaiino? aeey aaoio?ioieo
i?aai?ci?a.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iniiai? iieiaeaiiy aeena?oaoe?eii?
?iaioe aeiiia?aeaeenue ia:

—??/ieo iaoeiaeo eiioa?aioe?yo i?ioani?nueei-aeeeaaeaoeueeiai neeaaeo
Ee?anueeiai oi?aa?neoaoo ?iai? Oa?ana Oaa/aiea (Ee?a, 1997, 1998);

—?? c`?cae? a?ae?iaeieia?/iiai oiaa?enoaa Oe?a?ie (Ee?a, 1997);

—Anaoe?a?inuee?e i?aeaoc?anuee?e iaoeiai-iaoiaee/i?e eiioa?aioe?? c
i?iaeai i?e?iaeie/eo iaoe “*aoaa?o? Ea?eoeinuee? /eoaiiy” (Iieoaaa,
1997);

—Anaoe?a?inuee?e iaoeiai-i?aeoe/i?e eiioa?aioe?? niaoe?ae?no?a oa
nooaeaionueei? iieiae? “No/ani? iiaeyaee c iiioey?ecaoe?? i?e?iaeie/eo
iaoe” (Iieoaaa, 1998).

Ioae?eaoe??. Ca iaoa??aeaie aeena?oaoe?eieo aeine?aeaeaiue iioae?eiaaii
9 ae?oeiaaieo ?ia?o, a oiio /ene? noaoo? a iaoeiaeo aeo?iaeao — 3,
noaoo? a ca??ieeao iaoeiaeo i?aoeue — 4, oace a iaoa??aeao eiioa?aioe?e
— 2.

No?oeoo?a ? ianya ?iaioe. Aeena?oaoe?eia ?iaioa neeaaea?oueny c?
anooio, iaeyaeo e?oa?aoo?e, ?icae?eo, ui oa?aeoa?eco? ia`?eoe oa iaoiaee
aeine?aeaeaiue, oa 2-o ?icae?e?a, ui i?noyoue aenia?eiaioaeuei?
?acoeueoaoe. Iiaiee ianya aeena?oaoe?eii? ?iaioe caeia? 121 noi??ieo
iaoeiiieno, aeeth/a? 9 oaaeeoeue, 10 ?enoie?a. Nienie aeei?enoaieo
aeaea?ae aeeth/a? 161 iaeiaioaaiiy, c ieo 50 aiaeiiiaieo.

IAOA??AE OA IAOIAeEEA AeINE?AeAEAIUe

Iiaeaeuei? oieneeaioe. sse iiaeaeuei? oieneeaioe a ?iaio? aoee
aeei?enoai? ?iie naeioeth oa i?eaeth. Aea?? oeeo aeaio aaaeeeo iaoae?a
aeey i?iaaaeaiiy i?iaeoeoe?eii-oieneeieia?/ieo aeine?aeaeaiue coiiaeaiee
oaeeie oaeoi?aie: aea/ai?noth, oiene/i?noth oa cia/aiiyi aeey
a?ieia?/ieo nenoai.

Ia`?eoe aeey aeine?aeaeaiue a?ii?iaeoeoe?eiiai i?ioeano. Iaea?eueo
o?aceeaei aei ae?? oieneeaio?a ? aiaeia na?aaeiaeua. Aiae?c e?oa?aoo?ieo
a?aeiiinoae c i?eaiaeo ae?? caa?oaeiee?a ia i?iaeoeoeai?noue ??cieo a?oi
a?ae?ia?iio?a iieacaa, ui iaeiaio aea/aieie a?oiaie ? aeu? aiaei?
?ineeie oa aiaei? o?aaaoi? (?eae).

A aeai?e ?iaio? ye aaoa?io?ioi? iiaeaeuei? ia`?eoe aoee aeei?enoai?
iieiaeue cieioi? ?eaee — Carasius auratus auratus (L.) oa aoi? Poecilia
reticulata Peters; ye aaoio?ioi? iiaeaeuei? ia`?eoe — a?ae?iiae?io?oe
Elodea canadensis Michx. oa aeaa aeaee ?ynie Lemna minor L. oa Lemna
trisulca L., ye? ? oeiiaeie i?aaenoaaieeaie oei?e i??nieo aiaeiei
Oe?a?ie.

Iaoiaeeea aecia/aiiy a?ii?iaeoeoeaiino?. I?aeaeine?aei? i?aai?cie
ae?iuoaaeenue a eaai?aoi?ieo oiiaao ca noaiaea?oieie iaoiaeeeaie
i?ioyaii 0.5-1.5 i?nyoey (a caeaaeiino? a?ae neee i?iyao oiene/iiai
aoaeoo). Oieneeaioe aeiaeaaaeeny o aiaeo o aeaeyae? ?ic/eio nieae
Pb(NO3)2 oa NiSO4 a?aeiia?aeii? eiioeaio?aoe??, ?ic?aooiie
eiioeaio?aoe?? a?any ia ?ii oieneeaioo. Ca ?acoeueoaoaie aenia?eiaioo
aecia/aeenue oae? i?iaeoeoe?eii-aia?aaoe/i? iieacieee:

Ieoiia oaeaee?noue ?inoo [Caeea,83] ca oi?ioeith:

g = ln(m2/m1)/(t2-t1)

aea m2 ? m1 — a?aeiia?aei? cia/aiiy a?iiane a iiiaioe /ano t2 ? t1.

Aaeiaa aoaeoeai?noue eiiaa?ooaaiiy ei?io (aeey aaoa?io?ioieo i?aai?ci?a)
[Caeea,83] ca oi?ioeith:

K1 =m/R

aea m — i?e??no a?iiane ca iaaiee /an, R — aaee/eia niiaeeoiai
?aoe?iio ca oie aea /an.

Iieaciee iioeiaeueiino? (aeey aaoa?io?ioieo i?aai?ci?a) [Aaiaecth?a, 93]
ca oi?ioeith:

II =gK1

aea g — ieoiia oaeaee?noue ?inoo ca iaaiee ia??iae, K1 — aaeiaa
aoaeoeai?noue eiiaa?ooaaiiy ei?io ca oie aea ia??iae.

Aaciiai?ii? iieacieee i?iaeoeoe?? oa aeano?oeoe??: ieoiia i?iaeoeoe?y
a?iiane NPP (/enoa) oa GPP (aaeiaa), iieaciee aeeoaiiy — RA. Iieacieee
aecia/aeenue a i?ioean? aaciiai?io ca a?aeiioaiiyi aeae?eaiiai aai
iiaeeiooiai eenith aei ne?i? a?iiane, ca i?eioeeiii oaiieo ? na?oeeo
neeyiie. Aeniiceoe?y o?eaaea 2 aiaeeie, ?ic?aooiie a?any ia 24 aiaeeie c
o?aooaaiiyi o?eaaeino? na?oeiai ? oaiiiai ia??iaeo. Eiioeaio?aoe?th
eenith o aiae? aecia/aee o?i?/ii ca iaoiaeii A?ieea?a.

Ai?no nooi? ?a/iaeie, aecia/aany i?ney aeaiaiaeeiiiai aenoooaaiiy i?e
oaiia?aoo?? +1050N.

Ai?no i?aai?/iiai aoaeaoeth a noo?e ?a/iaei? aecia/aany ca Oth??iei
(iie?a a?o?iiaoia ieeneaiiy), canoiniaoaaee iiaeeo?eaoe?th Inoaiaiy
(1965).

?ioaa?aeueiee iieaciee i?iaeoeoeaiino? (aeey aaoio?ioieo
i?aai?ci?a)[Ion??iei 98], ui aecia/aany ca oi?ioeith:

?II=gNPP

aea g — ieoiia oaeaee?noue i?e?inoo a?iiane ca iaaiee ia??iae, NPP —
/enoa ieoiia i?iaeoeoe?y, aecia/aia ca aaciiai?iii iai?ee?ioe? oeueiai
ia??iaeo.

Aeey ioe?iee aoaeoo ae?? oieneeaio?a ia iaai? i?iaeoeoe?eii-aia?aaoe/i?
iieacieee aoea canoiniaaia oi?ioea Aaaa?a (Oaaa?a) [Eiee/anoaaiiay
oieneeieiaey 73]:

E=Ct

aea E — oea aoaeo ae?? oieneeaioo, C — eiai eiioeaio?aoe?y, a t — /an
ae?? ia aeine?aeaeoaaiee ia`?eo.

Io?eiai? ?acoeueoaoe aoei ia?iaeaii noaoenoe/ii [Aaeee 62].
Aeinoia??i?noue a?aei?iiinoae ioe?ithaaeanue ca t-e?eoa???i Nouethaeaioa
i?e ??ai? cia/eiino? ?=0.05. An? io?eiai? ?acoeueoaoe ii?iiaaii
a?aeiinii eiio?ieth ? aeey c?o/iino? ii??aiyiiy ae?aaeaii o a?aenioeao.
Iiaoaeiaa a?ao?/ieo caeaaeiinoae oa ?o iaoaiaoe/iee iien
cae?enithaaeenue ia ia?niiaeueiiio ?AI noi?niiio eiii`thoa?? ca
aeiiiiiaith i?ia?aie MicroCal Origin: Version 3.0. I?eaeaoi?noue
iaoaiaoe/iiai iieno aieeao oieneeaio?a ioe?ithaaee ca eiao?oe??ioii
ia?ii? ei?aeyoe?? i?ae iaoaiaoe/ii i/?eoaaieie ? aenia?eiaioaeueii
cao?eniaaieie cia/aiiyie i?iaeoeoe?eieo iieaciee?a oa ca O2 e?eoa???i
I??niia.

CI?IA A?II?IAeOEOEAIINO? ?EA CA OIIA CAA?OAeIAIIAI NA?AAeIAEUA

Aieea eii?a naeioeth ia i?iaeoeoe?eii-aia?aaoe/i? ia?aiao?e iieiae? ?ea.
Aea/aiiy aieeao eii?a naeioeth ia a?ii?iaeoeoeai?noue ?ea i?iaiaeeeinue
ia iieiae? C. auratus a?eii i?aoi?a i?nyoe?. I?ioyaii o?ueio oeaei?a
i?aeaeine?aei? oaa?eie oo?eioaaeeny a aiae? c eiioeaio?aoe??th eii?a
naeioeth 0.2; 0.4; 0.8; 1.6; 3.2; 6.4; 12.8 ia/aei3.

Io?eiai? iaie ?acoeueoaoe iieacaee a oe?eiio iaaaoeaiee aieea ?ii?a
naeioeth ia i?iaeoeoe?eiee i?ioean o iieiae? ?ea. Ia?oee oeaeaeaiue
oiene/iiai aieeao eii?a naeioeth oa?aeoa?eco?oueny aeinoia??iei
iaae?iiyi ieoiii? oaeaeeino? ?inoo i?e eiioeaio?aoe?yo 0.2 ? 0.4
ia/aei3, aaeiai? aoaeoeaiino? eiiaa?oaoe?? ?ae? oa iieacieea
iioeiaeueiino? i?e 0.2 ia/aei3. A?eueo aenie? eiioeaio?aoe?? oieneeaioo
aeyaeythoue noeioethth/o ae?th ? aeeeeeathoue aeinoia??ia i?aeaeuaiiy
ieoiii? oaeaeeino? ?inoo i?e eiioeaio?aoe?yo 0.8; 1.6; 12.8 ia/aei3,
aaeiai? aoaeoeaiino? eiiaa?oaoe?? ?ae? oa iieacieea iioeiaeueiino? i?e
eiioeaio?aoe?yo 0.8; 1.6; 3.2; 6.4; 12.8 ia/aei3. I?e/iio noeioethth/ee
aoaeo, i?aeaeuaiiy iieacieea iioeiaeueiino? ia 26%, iaeyne?aa?oa
i?iyaey?oueny i?e iaea?eueo?e eiioeaio?aoe?? oieneeaioo. Noeioeyoe?y
i?iaeoeoeaiino?, yea aeineoue /?oei i?ine?aeeiaoaaeanue ia nueiio aeiao,
cieea? ia /ioe?iaaeoeyoo. Caeeoa?oueny aeinoia??iei cieaeaiiy on?o
o?ueio i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a i?e eiioeaio?aoe?yo 0.2 ?
0.4 ia/aei3 ? niinoa??aa?oueny iiaeaeueoa cieaeaiiy i?iaeoeoeaiino? i?e
a?eueo aenieeo eiioeaio?aoe?yo oieneeaioo. Ieoiia oaeaee?noue
?inoo aeinoia??ii cieaeo?oueny i?e 6.4 ? 12.8 ia/aei3, a iieaciee
iioeiaeueiino? i?e 1.6; 3.2; 6.4; 12.8 ia/aei3. Iaea?eueoa cieaeaiiy
i?iaeoeoeaiino? — ciaioaiiy ieoiii? oaeaeeino? ?inoo ? iieacieea
iioeiaeueiino? ia 76% ii??aiyii c eiio?ieai niinoa??aa?oueny ciiao ae
oaee i?e iaea?eueo?e eiioeaio?aoe?? eii?a naeioeth 12.8 ia/aei3.
Iaiao?aeii caa?iooe oaaao ia oa, ui aoaeoeai?noue eiiaa?oaoe?? ?ae? i?e
oe?e eiioeaio?aoe?? caeeoa?oueny aeineoue aenieith ? eeoa ia 1.6%
ieae/ith a?ae eiio?ieth. Oea iiynith?oueny ??ceei cieaeaiiyi iiaeeiaiiy
i?aeaeine?aeieie ?eaaie ei?iiaiai ?aoe?iio. O?ao?e oeaeaeaiue
aenia?eiaioo oa?aeoa?eco?oueny ae??aithaaiiyi iieaciee?a i?iaeoeoeaiino?
i?aeaeine?aeieo ?ea i?e eiioeaio?aoe?yo naeioeth aei 6.4 ia/aei3.
Aeinoia??ia cieaeaiiy ieoiii? oaeaeeino? ?inoo a?aei?/aii i?e
eiioeaio?aoe?yo 0.2; 1.6 oa 6.4 ia/aei3, a iieacieea iioeiaeueiino? eeoa
i?e 6.4 ia/aei3. Eiioeaio?aoe?y 12.8 ia/aei3 i?ecaaea aei ae?i?iaoe??
on?o i?aeaeine?aeieo ?ea c 16-? ii 19-o aeiao aenia?eiaioo.

I?ioeaioiee ai?no nooi? ?a/iaeie, ui aecia/aany iai?ee?ioe?
aenia?eiaioo, aeey an?o a?oi i?aeaeine?aeieo ?ea, ie??i eiioeaio?aoe??
12.8 ia/aei3, eieeaaany a?ae (19.2±0.2)% a eiio?ie? aei (20.4±0.2)% i?e
eiioeaio?aoe?? 3.2 ia/aei3.

Aiae?c aenia?eiaioaeueieo oi/ie, ui a?aeia?aaeathoue aeiciaee aoaeo ae??
caa?oaeieea ia ieoiio oaeaee?noue ?inoo oa iieaciee iioeiaeueiino?
i?aeaeine?aeieo ?ea, aoa i?iaaaeaiee ca oi?ioeith Aaaa?a. Ca aeiiiiiaith
AII ii aenia?eiaioaeueieo oi/eao aoea iiaoaeiaaia iie?iii?aeueia
?aa?an?y aeey ieoiii? oaeaeeino? ?inoo c ??aiyiiyi:

y=93.83+0.311x-0.004×2; R=0.99; O2 = 3.98

aeey iieacieea iioeiaeueiino? c ??aiyiiyi:

y=93.76+0.488x-0.005×2; R=0.98; O2 = 3.69

aea o — oea aeica, ui aecia/aeanue ye aeiaooie eiioeaio?aoe?? oieneeaioo
ia /an eiai ae??. A aeaiiio aeiaaeeo iiceoeai? eiao?oe??ioe a?ey o
oa?aeoa?ecothoue noeioethth/ee aieea, a iaaaoeai? — ?ia?aoth/ee. Aiae?c
??aiyiue iieaco?, ui noeioeyoe?y i?iaeoeoeaiino? c?inoa? e?i?eii
oiene/iiio iaaaioaaeaiith, a ?ia?aoaaiiy — eaaae?aoe/ii. Ine?eueee
eiao?oe??io noeioeyoe?? ca aaniethoiith aaee/eiith cia/ii ia?aaeuo?
eiao?oe??io ?ia?aoaaiiy, oi ?nio? i?ii?aeie eiioeaio?aoe?e eii?a
naeioeth oa /ano ?o ae??, aea noeioethth/ee aoaeo ia?ae?eaa? ?ia?aoth/ee
? a?aeaoaa?oueny ca?eueoaiiy i?iaeoeoeaiino?. Oaeei /eiii, oai?aoe/ii
iiynith?oueny iiyaa noeioethth/eo aoaeo?a.

Aieea eii?a i?eaeth ia i?iaeoeoe?eii-aia?aaoe/i? ia?aiao?e iieiae? ?ea.
Aea/aiiy aieeao eii?a i?eaeth ia a?ii?iaeoeoeai?noue ?ea i?iaiaeeeinue
ia iieiae? C.auratus oa P.reticulata a?eii i?aoi?a i?nyoe?.

I?ioyaii oanoe oeaei?a iieiaeue C.auratus oo?eioaaeanue o aiae? c
eiioeaio?aoe??th eii?a i?eaeth 0.01; 0.1; 1 ia/aei3. An? aeine?aeaeoaai?
eiioeaio?aoe?? eii?a i?eaeth aeeeeeathoue noeioeyoe?th an?o
aeine?aeaeoaaieo i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a i?ioyaii onueiai
aenia?eiaioo. Iaeneiaeueia noeioeyoe?y niinoa??aa?oueny i?e
eiioeaio?aoe?? 1 ia/aei3, aeey ieoiii? oaeaeeino? ?inoo ia 12.6%
iai?ee?ioe? ia?oiai oeaeiy, aeey aaeiai? aoaeoeaiino? eiiaa?ooaaiiy ?ae?
oa iieacieea iioeiaeueiino? ia 13.2% oa 27.2%, a?aeiia?aeii,
iai?ee?ioe? oinoiai oeaeiy. Iaea?eueo? a?aeoeeaiiy a?ae eiio?ieth aea?
iieaciee iioeiaeueiino?.

A?aenooi?noue ?ia?aoth/iai aoaeoo aeeeeeaea iaiao?aei?noue i?iaaaeaiiy
iiaeaeueoeo aeine?aeaeaiue c? ca?eueoaiiyi eiioeaio?aoe?? caa?oaeieea.
Iaeneiaeueia eiioeaio?aoe?y oieneeaioo aoea ca?eueoaia o aeanyoue ?ac?a.
Iieiaeue P.reticulata oo?eioaaeanue i?ioyaii i?nyoey i?e eiioeaio?aoe??
eii?a i?eaeth 0.1; 1; 10 ia/aei3. I?iaeoeoe?eii-aia?aaoe/i? iieacieee
(g, K1, II) o?enoaaeenue eiaei? aeanyoue ae?a. I?eai?/oaaiiy
i?iaeoeoeaiino? i?e eiioeaio?aoe?? 10 ia/aei3 ii/eia?oueny aaea c ia?oi?
aeaeaaee aenia?eiaioo ?, iineethth/enue iaaeae?, i?ecaiaeeoue aei
ae?i?iaoe?? 25% i?aeaeine?aeieo ?ea o a?oi? ia o?eaeoeyoo aeiao
aenia?eiaioo. A?eueo iecuee? eiioeaio?aoe?? oieneeaioo aeathoue
noeioeyoe?th i?iaeoeoeaiino?. Cia/ia noeioeyoe?y niinoa??aa?oueny i?e
eiioeaio?aoe?? 0.1 ia/aei3 c aeaaaeoeyoi? aeiae ? ia o?eaeoeyoo aeey
iieacieea iioeiaeueiino? aaea?/? ia?aaeuo? eiio?ieue. Eiioeaio?aoe?y 1
ia/aei3 aea? i?eai?/aiiy ieoiii? oaeaeeino? ?inoo ? iieacieea
iioeiaeueiino? ia aeaaaeoeyoo aeiao. Ia o?eaeoeyoo aeiao ??aaiue oeeo
iieaciee?a c?inoa? ? iaa?oue aeaui ia?aaeuo? eiio?ieue.

I?ioeaioiee ai?no nooi? ?a/iaeie, ui aecia/aany iai?ee?ioe?
aenia?eiaioo, iaaeaa c? ca?eueoaiiyi eiioeaio?aoe?? caa?oaeieea.

?acoeueoaoe iaio na??e aeine?ae?a ii aea/aiith i?eaeaaiai cao?o?iiy
iieacaee, ui, ye ? o aeiaaeeo naeioeaaiai cao?o?iiy, iae/ooeea?oei c
aeine?aeaeaieo i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a ? iieaciee
iioeiaeueiino?. A?i aea? iaea?eueo? a?aeoeeaiiy a?ae eiio?ieth ?
iaeae?aci?oa iieaco? aoaeo noeioeyoe?? aai ?ia?aoaaiiy i?iaeoeoe?eiiai
i?ioeano.

Ia aeaeue, a ca`yceo c iaaeinoaoiueith e?euee?noth aenia?eiaioaeueieo
oi/ie aeiciaa caeaaei?noue aeey iieacieea iioeiaeueiino? P.reticulata ia
i?aeaea?oueny iaoaiaoe/iiio iieno iie?iiiii. Aenia?eiaioaeueieo oi/ie,
ui oa?aeoa?ecothoue caeaaei?noue aeica — aoaeo aeey N.auratus, a?eueoa,
? ca aeiiiiiaith AII aiia aoea iienaia ??aiyiiyi:

y=111.98+0.87x-0.015×2; R=0.72; O2 = 3.09

Ia ?en. 1 cia?aaeaii aenia?eiaioaeuei? oi/ee, ui a?aeiia?aeathoue ci?i?
iieacieea iioeiaeueiino? aeey iaio aeae?a ?ea. Oaeei /eiii, c?iaeaia
ni?iaa aeaanoe ?aeeia ??aiyiiy aeey iieno ae?? iecueeeo eiioeaio?aoe?e
eii?a i?eaeth ia iaeaea? i?aeaeine?aei? a?oie. Iie?iii?aeueia ??aiyiiy:

y=111.05+0.81x-0.02×2; R=0.87; O2 = 26.98

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iieno? aenia?eiaioaeuei? cia/aiiy iieacieea iioeiaeueiino? iaio aeae?a
?ea. I?e ii??aiyii? /eneiaeo eiao?oe??io?a iaio ??aiyiue ?noioii?
??cieoe? i?ae ieie ia niinoa??aa?oueny. Ioaea, i?iaiae?coaaaoe io?eiai?
aenia?eiaioaeuei? aeai? oa ?o oai?aoe/io ia?iaeo, iiaeia noaa?aeaeoaaoe,
ui aieea ye ?ii?a naeioeth, oae ? ?ii?a i?eaeth ia
i?iaeoeoe?eii-aia?aaoe/i? iieacieee iieiae? ?ea ? ie?aiaeeiai?/iei. Iae?
eiioeaio?aoe?? caa?oaeieea noeioeththoue a?ii?iaeoeoea-i?noue, a?eueo
aenie? — ?ia?aothoue.

?en.1 Aeiciaa caeaaei?noue iieacieea iioeiaeueiino?

iaio aeae?a ?ea i?e i?eaeaaiio cao?o?ii?.

CI?IA A?II?IAeOEOEAIINO? A?Ae?IIAE?IO?O?A CA OIIA CAA?OAeIAIIAI
NA?AAeIAEUA

Aieea eii?a naeioeth ia i?iaeoeoe?eii-aia?aaoe/i? ia?aiao?e
a?ae?iiae?io?o?a. I?e aea/aii? aieeao eii?a naeioeth ia
a?ii?iaeoeoeai?noue aeueo aiaeieo ?ineei ia`?eoii aeine?aeaeaiue aoee
aeaa aeaee ?ynie Lemna minor L. oa Lemna trisulca L., ? aeiaeay —
Elodea canadensis Michx. ?ineeie ae?iuoaaeenue i?e eiioeaio?aoe??
naeioeth 0.1, 1, 10 ? 100 ia/aei3.

Aeeiai?ea ci?ie i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a aeiaea? a?ae
eiioeaio?aoe?? eii?a naeioeth aeyaeeanue ie?aiaeeiai?/iith. I?e
eiioeaio?aoe?? 0.1 ia/aei3 niinoa??aaeanue noeioeyoe?y ieoiii?
oaeaeeino? ?inoo ?ineei, yea eeoaeanue aeinoia??iith eeoa i?ioyaii
ia?oeo aeaio oeaei?a. A?eueo aenie? eiioeaio?aoe?? oieneeaioo aeaee
aeaye? eieeaaiiy ieoiii? oaeaeeino? ?inoo, aea an? ?o cia/aiiy
aeinoia??ii iaio? a?ae eiio?ieth. Aaee/eia noeioeyoe?? iieacieea
i?ioyaii aenia?eiaioo c /anii ciaioo?oueny, a aaee/eia ?ia?aoaaiiy
ca?eueoo?oueny. *enoa ieoiia i?iaeoeoe?y a?iiane aeiaea?, ui
aecia/aeanue ca aaciiai?iii, a ia?oee aeaiue cao?o?iiy aeaea eeane/io
ea?oeio caeaaeiino? a?ae eiioeaio?aoe?? oieneeaioo, i?eai?/aiiy
iieacieea a?aeaoaaeinue iiiiaeeiai?/ii. Iaaeae? /enoa ieoiia i?iaeoeoe?y
aeiaea? ci?ith?oueny aeineoue ia??aiii??ii ? i?e an?o eiioeaio?aoe?yo
eii?a naeioeth, e??i iaeneiaeueii?, aeaaaea cia/i? a?aeoeeaiiy a?ae
eiio?ieth, ye o a?e noeioeyoe??, oae ? o a?e ?ia?aoaaiiy, eeoa i?e
eiioeaio?aoe?? 100 ia/aei3 /enoa ieoiia i?iaeoeoe?y aanue /an
caeeoaeanue cia/ii iaioith ca eiio?ieue.

Canoinoaaaoe aeey ioe?iee aoaeoo ae?? naeioeth ia aeiaeath oi?ioeo
Aaaa?a ? ?icaeyioaoe oeae iieaciee a eii?aeeiaoao aeica-aoaeo, ie
io?eia?ii e?eao, cia?aaeaio ia ?en.2. Ea?oeia ci?ie /enoi? ieoiii?
i?iaeoeoe?? iaaaaeo? ia??iaee/i? aa?iii?ei? eieeaaiiy, ui caoooathoue.

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Oiio aeey ?ioa?i?eoaoe?? io?eiaieo ?acoeueoao?a ie i?iiiio?ii
a?iioaoe/io iiaeaeue “a?ieia?/iiai inoeeeyoi?a”. Aeey oeueiai
?icaeyiaii i?aai?ci c eiai caeaoi?noth noai?thaaoe i?iaeoeoe?th ye
nenoaio, ui ciaoiaeeoueny a noai? ??aiiaaae c ioi/oth/ei na?aaeiaeuai, a
caa?oaeiee, ui iio?aiey? a na?aaeiaeua, ye no?aniaee oaeoi?, ui ii?ooo?
oeth ??aiiaaao. Iiaeaeue ia?aaeaa/a?, ui i?aai?ci ia? iaaiee caian
i?oeiino? (aaeaioeaiee iioaioe?ae), ? iiaea

?en.2 Aeeiai?ea ci?ie /enoi? ieoiii? i?iaeoeoe??

aeiaea? i?e naeioeaaiio cao?o?ii?.

i?ioeae?yoe no?aniaiio oaeoi?o. sseui oieneeaio aoaea ae?yoe a iaaeao
ciie oiea?aioiino? i?aai?cio ? ia i?ecaaaea aei oaeaeeiai ?oeioaaiiy
a?inenoaie i?aai?ci — na?aaeiaeua, a eeoa aeaaaea ?? c? noaio
??aiiaaae, oi ie io?eia?ii eieeaaeueiee i?ioean. Eieeaaiiy i?eieiyoueny
oiae?, eiee cieeia ae?y no?aniaiai oaeoi?a, aai eiai cia/aiiy
ia?aaeuaoue iaae? ciie oiea?aioiino? ? c?oeiothoue nenoaio. Ca
cai?iiiiiaaiith iiaeaeeth i?ioean aoaea iiaoi?thaaoenue aei iiaiiai
aeniaaeaiiy nenoaie. Ia ?en.2 ie aa/eii, ui c? c?inoaiiyi aeice
iaeneioie eieeaaiue iieacieea i?iaeoeoeaiino? ciaioothoueny, i?i?ioie
caeeoathoueny a?aeiinii iino?eieie. I?aai?ci aeniaaeo?oueny,
i?iyaey?oueny oiene/iee aieea caa?oaeieea.

I?e ii??aiyii? ae?? eii?a naeioeth ia a?ii?iaeoeoeai?noue aeiaea? c
aeei?enoaiiyi ?II aeaeii, ui eiioeaio?aoe?y 0.1 ia/aei3 ae?? ia ?ineeie
noeioethth/a, a a?eueo aenie? eiioeaio?aoe?? i?iaeoeoeai?noue
?ia?aothoue.

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00000000

I?e aea/aii? aieeao eii?a naeioeth ia a?ii?iaeoeoeai?noue ?ynee iaie
aeine?aeaeoaaeanue oaeiae caeaaei?noue i?iaeoeoe?eii-aia?aaoe/ieo
iieaciee?a a?ae eiioeaio?aoe?? oieneeaioo i?e ??ci?e ?ioaineaiino?
ina?oeaiiy. Ciaioaiiy ?ioaineaiino? ina?oeaiiy a ii?aeiaii? c?
c?inoaiiyi oiene/iino? na?aaeiaeua i?ecaiaeeoue aei a?eueo ??ceiai
niaaeo ieoiii? oaeaeeino? ?inoo. Ia ?en. 3 iieacai? ci?ie ieoiii?
oaeaeeino? ?inoo a?iiane L.minor, ui i?ioyaii aeaio oeaei?a
ae?iuoaaeanue ca oiia caa?oaeiaiiy na?aaeiaeua naeioeai i?e
?ioaineaiino? ina?oeaiiy 3000 ee oa 2000 ee.

?acoeueoaoe a iaio aeiaaeeao ii?iiaai? a?aeiinii eiio?ieth c
a?aeiia?aeiei ??aiai ina?oeaiino?, ui aea? iiaeeea?noue iieacaoe naia
aoeeo ae?? oieneeaioo ? iicaaaeoeny “ii?iaeueiiai” cieaeaiiy
i?oaeoeoeaiino? ?ineei i?e ciaioaii? ??aiy ina?oeaiiy.

Cia/ia ca?eueoaiiy ??cieoe? a?ii?iaeoeoe?eieo ia?aiao??a i?e aenieeo
??aiyo caa?oaeiaiiy na?ae/eoue, c iaeiiai aieo, i?i c?inoaiiy
aia?aaoe/ieo aeo?ao i?aai?cio a oiiaao oiene/iiai na?aaeiaeua,
ci?io eiai

?en.3 Caeaaei?noue ieoiii? oaeaeeino? ?inoo ?ynee iaeaiueei?

a?ae eiioeaio?aoe?? naeioeth oa ?ioaineaiino? ina?oeaiiy.

aia?aaoe/iiai aaeaino a iai?yieo aeano?oeoe??, c ?ioiai — i?i aaeeeo
aia?ai?ii?noue i?ioeano aaeaioaoe?? ?ineei aei ?nioaaiiy ca oiia
caa?oaeiaiiai na?aaeiaeua. I?e ?icaeyae? i?iaeoeoe?eiiai i?ioeano
L.trisulca aeyaeeany cia/ii /ooeea?oith aei ae?? eii?a naeioeth
aeine?aeaeoaaieo eiioeaio?aoe?e, i?ae L.minor. Ieoiia oaeaee?noue ?inoo
L.trisulca i?e eiioeaio?aoe?? 0.1 ia/aei3 aea? ?noioia ca?eueoaiiy a
ia?oee oeaeaeaiue aenia?eiaioo. Ia ae?oaiio oeaei? noeioeyoe?y iieacieea
i?iaeoeoeaiino? cieea? ? eiai cia/aiiy iaeaea ia ia?aaeuo? eiio?ieue.
Iiaia i?eieiaiiy ?inoo ?ineei niinoa??aa?oueny i?e eiioeaio?aoe?yo
naeioeth 10 ? 100 ia/aei3. I?ioeaioiee ai?no nooi? ?a/iaeie a ?ineeiao,
ui caciaee oiene/iiai aieeao, aeaui c?inoa? i?e eiioeaio?aoe?? 0.1
ia/aei3, i?e a?eueo aenieeo eiioeaio?aoe?yo caa?oaeieea cieaeo?oueny.
Aeinoia??ieo a?aei?iiinoae ai?noo i?aai?/iiai aoaeaoeth a?ae
eiioeaio?aoe?? eii?a naeioeth ia aeyaeaii.

Aieea eii?a i?eaeth ia i?iaeoeoe?eii-aia?aaoe/i? ia?aiao?e
a?ae?iiae?io?o?a. Aea/aiiy aieeao eii?a i?eaeth ia a?ii?iaeoeoeai?noue
aeueo aiaeieo ?ineei i?iaiaeeeiny, ye ? i?e aea/aii? ae?? eii?a
naeioeth, ia aeaio aeaeao ?ynee — Lemna minor L. oa Lemna trisulca L.,
? Elodea canadensis Michx. Eiioeaio?aoe?y caa?oaeieea noaiiaeea c
?ic?aooieo ia eii Ni2+ — 0.1, 1.0, 10, 100 ia/aei3. Aeiaeaoeiai
aecia/aeeny iieaciee aaoio?ioiiai aeeoaiiy RA oa aaeiaa ieoiia
i?iaeoeoe?y GPP, iaeaeaa iieacieee aecia/aeeny aeey aeiaea? o i?ioean?
aaciiai?io ca aeae?eaiei eeniai.

sse iieacaee ?acoeueoaoe i?iaaaeaieo aeine?ae?a, i?iaeoeoeai?noue
aeiaea? eaiaaenueei? a oiiaao i?eaeuecaa?oaeiaiiai na?aaeiaeua oaeiae a
oe?eiio iaaea?, aea aeeiai?ea ci?ie i?iaeoeoe?eii-aia?aaoe/ieo
iieaciee?a a?ae??ciy?oueny a?ae aeeiai?ee i?e naeioeaaiio cao?o?ii?. I?e
iaeaeu?e eiioeaio?aoe?? 100 ia/aei3 ie niinoa??aa?ii iaea?eueoee
noeioethth/ee aoaeo ae?? ?ii?a i?eaeth, ye aeey i?ioean?a i?iaeoeoaaiiy
(/enoa ? aaeiaa ieoiia i?iaeoeoe?y), oae ? aeey i?ioean?a aeano?oeoe??
(iieaciee aeeoaiiy). I?e a?eueo iecueeeo eiioeaio?aoe?yo iaeaeaa
iieacieee i?iaeoeoe?? eieeaathoueny c iaaaeeeeie a?aeoeeaiiyie a?ae
eiio?ieth, a i?ioeane aeeoaiiy noeioeththoueny i?e eiioeaio?aoe?yo 0.1 ?
1 ia/aei3 oa ?ia?aothoueny i?e 10 ia/aei3.

Ia nueiio aeiao oiene/i?noue ?ii?a i?eaeth i?iyaey?oueny a?eueo iii?oii.
Niinoa??aa?oueny cia/ia iaae?iiy /enoi? ? aaeiai? ieoiii? i?iaeoeoe??.
Cieaeaiiy oeeo iieaciee?a aeia?a ai?ieneio?oueny i?yieie c
eiao?oe??ioaie e?i?eii? ?aa?an?? -22.53.9; R=-0.96; P=0.01 ? -26.34.3;
R=-0.96; P=0.01 aeey aaeiai? ? /enoi? ieoiii? i?iaeoeoe??, a?aeiia?aeii.
Oe? caeaaeiino? iieacai? ia ?en. 4. Iaiao?aeii cacia/eoe, ui ciaioaiiy
/enoi? ieoiii? i?iaeoeoe?? a?aeaoaa?oueny a?eueo ??cei, i?ae aaeiai?.
I/aaeaeii, oea iia’ycaii c? noeioeyoe??th i?ioean?a aeeoaiiy. Iieaciee
aeeoaiiy ci?ith?oueny ie?aiaeeiai?/ii, nii/aoeo c?inoa? (i?e
eiioeaio?aoe?? 0.1 ia/aei3 ia 30% ia?aaeuo? eiio?ieue), a iio?i
ciaioo?oueny aei 39% i?e eiioeaio?aoe?? 100 ia/aei3. Ieoiia oaeaee?noue
?inoo aeiaea? oaeiae ci?ith?oueny ie?aiaeeiai?/ii, i?ioa, aeeiai?ea ??
ci?ie aeaui a?ae??ciy?oueny. Caeeoath/enue cia/ii iaioith a
ii??aiyii? c eiio?ieai, aiia nii/aoeo ciaioo?oueny (i?i?ioi i?e
eiioeaio?aoe?? 10 ia/aei3), a iio?i aea? i?aeaeuaiiy. ?ioaa?aeueiee
iieaciee i?iaeoeoeaiino? iaee?aua oa?aeoa?eco? ??aaiue caa?oaeiaiiy,
aeath/e iaeiaio? cia/aiiy ii??aiyii c eiio?ieai i?e a?aeiia?aeieo
eiioeaio?aoe?yo. I?eai?/aiiy oeueiai iieacieea a?aeaoaa?oueny
iiiiaeeiai?/ii.

Ia 14 aeiao niinoa??aa?oueny ua a?eueo neeueia i?eai?/aiiy
i?iaeoeoe?eieo iieaciee?a aeiaea?, i?ae ia nueiio, i?e eiioeaio?aoe?yo
0.1 ? 1 ia/aei3, ? iiaia i?eai?/aiiy i?e 10 ? 100 ia/aei3. Iieaciee
aeeoaiiy i?e 100 ia/aei3 ciaioo?oueny aei 15%, a i?e iaioeo
eiioeaio?aoe?yo caeeoa?oueny a?eueoei ca eiio?ieue. Ai?no nooi? ?a/iaeie
a ?ineeiao, ui caciaee oiene/iiai aieeao, aeaui cieaeo?oueny ? eeoa i?e
100 ia/aei3 a iaaeao iioeaee ia?aaeuo? eiio?ieue. Aeinoia??ieo
a?aei?iiinoae ai?noo i?aai?/iiai aoaeaoeth ia aeyaeaii.

picscalex100010009000003f43400000f002c00000000001400000026060f001e00ffff
ffff040014000000576f72640e004d6963726f736f667420576f7264050000000b020000
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d6ff0000000000009001000000000440001254696d6573204e657720526f6d616e000000
040000002d01010005000000090200000000040000000201010007000000fc0201000000
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0300070000001b04f7030301b703a80007000000fc020000ffffff000000040000002d01
040009000000fa02000000000000000000002200040000002d010500030000001e000700
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3c04302e30001a000d001a00040000002e01000005000000140200000000040000002d01
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040000002d010300070000001b04f703dd01b7038201040000002d010400040000002d01
0500030000001e00070000001604f603dd01b70382010400000002010100040000002d01
0600050000000902ff000000050000001402b8038201040000002e01010010000000320a
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040000002d010400040000002d010500030000001e00070000001604f603b702b7035c02
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0100040000002d010600050000000902ff000000050000001402b803f703040000002e01
010013000000320ab803f70305000400000000009b043c043130302e30001a0019001a00
0d001a00040000002e0100000500000014023880b0a2040000002d010100050000000902
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070000001b049903a70058037500040000002d010400040000002d010500030000001e00
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040000002d010300070000001b04d401ae0094014400040000002d010400040000002d01
0500030000001e00070000001604d201ae00930144000400000002010100040000002d01
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060102000c00000024030400c400a803cb00a803cb008903c40089030400000006010100
040000002d010500040000002d0104000800000026060f000600ffffffff010010000000
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f301000009000000fa02000003000000ff0000002200040000002d010800040000002d01
020008000000250302005902ef019e02f001040000002d01050004000000f00108000400
00002d0104000800000026060f000600ffffffff01001000000026060f001600ffffffff
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040000002d010800040000002d01020008000000250302007b020a027c02150204000000
2d01050004000000f0010800040000002d0104000800000026060f000600ffffffff0100
1000000026060f001600ffffffff00005702000013020000a10200001802000009000000
fa02000003000000ff0000002200040000002d010800040000002d010200080000002503
0200590215029e021502040000002d01050004000000f0010800040000002d0104000800
000026060f000600ffffffff01001000000026060f001600ffffffff000054030000b702
000059030000c502000009000000fa02000003000000ff0000002200040000002d010800
040000002d01020008000000250302005603c2025603b902040000002d01050004000000
f0010800040000002d0104000800000026060f000600ffffffff01001000000026060f00
1600ffffffff000031030000b70200007b030000bc02000009000000fa02000003000000
ff0000002200040000002d010800040000002d01020008000000250302003303b9027803
b902040000002d01050004000000f0010800040000002d0104000800000026060f000600
ffffffff01001000000026060f001600ffffffff000054030000d202000059030000e002
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08000000250302005603d4025603dd02040000002d01050004000000f001080004000000
2d0104000800000026060f000600ffffffff01001000000026060f001600ffffffff0000
31030000db0200007b030000e002000009000000fa02000003000000ff00000022000400
00002d010800040000002d01020008000000250302003303dd027803dd02040000002d01
050004000000f0010800040000002d0104000800000026060f000600ffffffff01001000
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2e0436032e042d03040000002d01050004000000f0010800040000002d01040008000000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04000800000026060f000600ffffffff01001000000026060f001600ffffffff00007303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02000400000000009b043c0420200c000c00040000002e0100000500000014023880b0a2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Ae?iuoaaiiy iaio aeae?a ?ynee a na?aaeiaeu?, caa?oaeiaiiio ?iiaie
i?eaeth, iieacaei iaaaoeaiee aieea caa?oaeieea ia a?ii?iaeoeoeai?noue
?ineei. Eiioeaio?aoe?? iaoaeo 10 ? 100 ia/aei3 iiai?noth i?eai?/othoue
i?iaeoeoeai?noue iaio aeae?a ?ineei aaea ia nueiio aeiao aenia?eiaioo.
I?eai?/aiiy ieoiii? oaeaeeino? ?inoo L.trisulca a?aeaoaa?oueny
iiiiaeeiai?/ii c? ca?eueoaiiyi eiioeaio?aoe?? eii?a i?eaeth. Ci?ia
ieoiii? oaeaeeino? ?inoo L..minor i?ioiaeeoue ie?aiaeeiai?/ii ? i?ney
ciaioaiiy oeueiai iieacieea aei 43% i?e eiioeaio?aoe?? 0.1 ia/aei3
niinoa??aa?oueny eiai c?inoaiiy aei ??aiy eiio?ieth i?e 1 ia/aei3 oa
iiaeaeueoa iaae?iiy aei ioey i?e aeueo eiioeaio?aoe?yo i?eaeth. Ia
/ioe?iaaeoeyoo aeiao ie?aiaeeiai?/ia caeaaei?noue iaae?iiy ieoiii?
oaeaeeino? ?inoo L.minor a?ae eiioeaio?aoe?? caa?oaeieea noa? ia
aeinoia??iith (cia/aiiy oeueiai

?en.4 Caeaaei?noue a?ii?iaeoeoe?eieo iieaciee?a aeiaea?

a?ae eiioeaio?aoe?? i?eaeth ia nueiio aeiao cao?o?iiy.

iieacieea i?e eiioeaio?aoe?yo 0.1 ? 1 ia/aei3 a?ae??ciythoueny a iaaeao
iioeaee).

Aeey L.trisulca iiai?noth i?eai?/o?oueny i?iaeoeoeai?noue i?e
eiioeaio?aoe?? 1 ia/aei3.Aeinoia??ieo a?aei?iiinoae ai?noo i?aai?/iiai
aoaeaoeth a caeaaeiino? a?ae eiioeaio?aoe?? i?eaeth aeey iaio aeae?a
?ineei ia aeyaeaii. Aeey L.minor a?aei?/aii aeayea i?aeaeuaiiy ai?noo
nooi? ?a/iaeie i?e eiioeaio?aoe?yo 0.1 ? 10 ia/aei3. Ai?no nooi?
?a/iaeie a L.trisulca iiiiaeeiai?/ii ca?eueoo?oueny c? c?inoaiiyi
eiioeaio?aoe?? ?ii?a i?eaeth.

Ca ?acoeueoaoaie i?iaaaeaieo aeine?aeaeaiue iiaeia noaa?aeaeoaaoe, ui
iayai?noue ye ?ii?a naeioeth, oae ? ?ii?a i?eaeth a na?aaeiaeu?
iaaaoeaii aieeaa? ia a?ii?iaeoeoeai?noue a?ae?iiae?io?o?a. Aieea ?ii?a
naeioeth ia an? aeaee aeine?aeaeoaaieo ?ineei aeyaeany ie?aiaeeiai?/iei.
?iie i?eaeth, iaaiaee, iiiiaeeiai?/ii i?eai?/othoue a?ii?iaeoeoeai?noue.
A ?iaio? aeei?enoai? iaeiaeia? eiioeaio?aoe?? iaio iiaeaeueieo
oieneeaio?a. Caaeaaeue aai iiaia i?eai?/aiiy i?iaeoeoe?eieo iieaciee?a
on?o aeine?aeaeoaaieo ?ineei i?e ae?? ?ii?a i?eaeth niinoa??aaeenue
?ai?oa ? i?e a?eueo iecueeeo eiioeaio?aoe?yo, i?ae i?e ae?? ?ii?a
naeioeth. Oea na?ae/eoue, ui ?iie i?eaeth ? a?eueo oiene/ieie aeey
a?ae?iiae?io?o?a.

AENIIAEE

1. Iayai?noue ?ii?a naeioeth oa i?eaeth a na?aaeiaeu? iaaaoeaii aieeaa?
ia aeine?aeaeoaai? a?inenoaie, niinoa??aa?oueny ??cea cieaeaiiy
a?ii?iaeoeoeaiino? an?o ia`?eo?a i?e iaeneiaeueieo (a?eueoa 10 ia/aei3)
eiioeaio?aoe?yo caa?oaeiee?a.

2. Aeeiai?ea aieeao naeioeth ia i?iaeoeoe?eii-aia?aaoe/i? iieacieee an?o
aeine?aeaeoaaieo ia`?eo?a oa i?eaeth ia aaoa?io?ioi? i?aai?cie (iieiaeue
?ea) ? ie?aiaeeiai?/iith, i?e eiioeaio?aoe?yo aei 10 ia/aei3
i?iyaey?oueny noeioethth/ee (ai?iaoe/iee) aieea oieneeaio?a.

3. ?iie i?eaeth iiiiaeeiai?/ii i?eai?/othoue a?ii?iaeoeoeai?noue
a?ae?iiae?io?o?a.

4. Aiae?oe/ia caeaaei?noue i?ae ??aiai caa?oaeiaiiy na?aaeiaeua ?
i?iaeoeoe?eii-aia?aaoe/ieie iieacieeaie iieiae? ?ea iiaea aooe ae?aaeaia
??aiyiiyi aeaeo:

y = Ax — Bx2 + C

aea o — i?iaeoeoe?eii-aia?aaoe/iee iieaciee, o — aeica (aeiciaa
iaaaioaaeaiiy) oieneeaioo, A, A ? N — aeaeiniaoeeo?/i? eiinoaioe.

5. Iaea?eueo /ooeeaeie ? ?ioi?iaoeaieie c on?o aeine?aeaeaieo
i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a ? ?ioaa?aeuei? iieacieee —
iieaciee iioeiaeueiino? (II) aeey aaoa?io?ioieo i?aai?ci?a oa
?ioaa?aeueiee iieaciee i?iaeoeoeaiino? (?II) aeey aaoio?ioieo
i?aai?ci?a.

6. Aeey aeine?aeaeoaaieo aeae?a a?ae?iiae?io?o?a ?iie i?eaeth aeyaeeenue
a?eueo oiene/ieie, i?ae ?iie naeioeth, aeey iieiae? ?ea oiene/i?noue
iaio ?ii?a i?eaeecii iaeiaeiaa.

7. I?iya oiene/ii? ae?? caa?oaeiee?a ia aaoio?ioi? a?inenoaie
ca?eueoo?oueny c? ciaioaiiyi iioieo aeinooiii? aia?a??.

8. Iecuee? eiioeaio?aoe?? oieneeaio?a (iaioa 10 ia/aei3) ?ia?aothoue
i?iaeoeoe?ei? i?ioeane ? noeioeththoue aeano?oeoe?ei?, a?eueo aenie?
eiioeaio?aoe?? i?eai?/othoue iaeaeaa i?ioeane.

9. Iieacieee II oa ?II iaea?eueo aaeaeaaoii a?aeia?aaeathoue
oieneeieia?/io neooaoe?th ? iiaeooue aooe ?aeiiaiaeiaai? ye
a?ii?iaeoeoe?ei? e?eoa??? oiene/iino? i?e a?ioanooaaii? caa?oaeiaieo
aiae.

NIENIE ?IA?O, IIOAE?EIAAIEO CA OAIITH AeENA?OAOe??

1.Aaiaecth?a A.I., Eaiaothe A.A.Aeeyiea eiiia naeioea e aiiiiey ia
aeii?iaeoeoeeiiiua ia?aiao?u iieiaee ?ua//Aeae?iaeie. aeo?i.—
1998.—34,?1.— N.85-90.

2.Gandziura V.P., Ignatyuk A.A. The use of productive energetic
parameters for estimation of an ecotoxicological situation//Aeiiia?ae?
Iaoe?iiaeueii? aeaaeai?? iaoe Oe?a?ie.— 1998.— ?11.— N.182-184.

3.Ion??iei I.I., Aaiaecth?a A.I., ?aiaothe I.A. Aieea naeioeth ia
a?ii?iaeoeoe?ei? ia?aiao?e
a?ae?iiae?io?o?a//Oe?.aio.aeo?i.—1998.—?6.— N. 609-614.

4.?aiaothe I.A. Canoinoaaiiy iao?aaeeoe?eieo iiaeaeae a
aeioieneeieia?/ieo aeine?aeaeaiiyo//A?niee Ee?anueeiai oi?aa?neoaoo
?iai? Oa?ana Oaa/aiea. ?io?iaeoeoe?y oa caa?aaeaiiy ?ineeiiiai
??ciiiai?ooy.— Aei.2.— Ee?a, 1999.— N.69-70.

5.?aiaothe I.A. I?iya ia?aaeienaeueieo aoaeo?a a
i?iaeoeoe?eii-oieneeieia?/ieo aeine?aeaeaiiyo//Ca. iaoe. i?aoeue
“Aeieia?y.A?ieia?/i? iaoee”.— Aei.1.— Iieoaaa, 1999.— N.83-91.

6.Aaiaecth?a A.I., ?aiaothe I.A. A?ii?iaeoeoe?eiee i?aeo?ae aei
aeioieneeieia?/ieo aeine?aeaeaiue//*aoaa?o? Ea?eoeinuee? /eoaiiy
(ca??iee noaoae).— Iieoaaa, 1997.—O.1.—N.45-47.

7.?aiaothe I.A., Nei?ioiae I.A. I?iaeoeoeai?noue ieaenoio?o?a a oiiaao
oiene/iiai na?aaeiaeua//I`yo? Ea?eoeinuee? /eoaiiy (ca??iee noaoae).—
Iieoaaa, 1998.— N.108-110.

8.?aiaothe I.A. Ci?ia a?ii?iaeoeoe?eieo ia?aiao??a iieiae? ?ea a oiiaao
o?ii?/iiai oieneeico//Ae?oaee c`?cae a?ae?iaeieia?/iiai oiaa?enoaa
Oe?a?ie.Oace aeiiia?aeae.— Oii 2.— Ee?a, 1997.— N.83-84.

9.?aiaothe I.A. Ioe?iea noaio aeiae?eey ca aeiiiiiaith a?ii?iaeoeoe?eieo
iieaciee?a//Iaoa??aee eiio. iai`yo? Aa?iaaenueeiai.—Iieoaaa,
1998.—N.25.

AIIOAOe?ss

?aiaothe I.A. Caeaaei?noue i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a
a?inenoai a?ae ??aiy caa?oaeiaiiy na?aaeiaeua aaaeeeie iaoaeaie.-
?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
ca niaoe?aeuei?noth 03.00.16 aeieia?y. — Ee?anueeee oi?aa?neoao ?iai?
Oa?ana Oaa/aiea, Ee?a, 1999.

Aeena?oaoe?y i?enay/aia aea/aiith a o?ii?/ieo aeine?aeao ye?niiai e
e?euee?niiai ca`yceo i?ae ??aiai caa?oaeiaiiy na?aaeiaeua ?iiaie
naeioeth oa i?eaeth oa i?iaeoeoe?eieie iieacieeaie aaoi- ? aaoa?io?ioieo
a?inenoai ia i?eeeaae? eaai?aoi?ieo iiioeyoe?e aeueo aiaeieo ?ineei oa
?ea. Io?eiaii iia? ?acoeueoaoe ii ci?i? ?ioaineaiino? oa aoaeoeaiino?
a?ii?iaeoeoe?eiiai i?ioeano a oiene/iiio na?aaeiaeu?. Iieacaii
ie?aiaeeiai?/ia i?eai?/aiiy i?iaeoeoe?eii-aia?aaoe/ieo iieaciee?a
aeine?aeaeoaaieo ?ineei ? oaa?ei c? ca?eueoaiiyi eiioeaio?aoe??
caa?oaeiee?a. Anoaiiaeaii aiae?oe/iee ca`ycie i?ae aeicith oieneeaioo oa
a?ii?iaeoeoeai?noth iieiae? ?ea. C?iaeaii i?eiouaiiy, ui c? ca?eueoaiiyi
aeice noeioethth/ee aoaeo c?inoa? e?i?eii, a ?ia?aoth/ee eaaae?aoe/ii.
I?iaeaiiino?iaaii c?inoaiiy oiene/iiai aoaeoo c? ciaioaiiyi iioieo
aeinooiii? aia?a??. Ciaeaeaii iaea?eueo /ooeea? oa ?ioi?iaoeai?
i?iaeoeoe?eii-aia?aaoe/i? iieacieee. Iaa?oioiaaii iiaeeea?noue ?o
iiaeaeueoiai aeei?enoaiiy aeey ioe?iee noaio na?aaeiaeua ca ooieoe??th
a?aeaoeo a?inenoai.

Eeth/ia? neiaa: aaaee? iaoaee, ye?noue na?aaeiaeua, aeioieneeieia?y,
i?iaeoeoeai?noue, a?i?iaeeeaoe?y, i?iaeoeoe?eii-aia?aaoe/i? iieacieee.

AIIIOAOeEss

Eaiaothe A.A. Caaeneiinoue i?iaeoeoeeiiii-yia?aaoe/aneeo ia?aiao?ia
aeinenoai io o?iaiy caa?yciaiey n?aaeu oyae?euie iaoaeeaie.- ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/?iie noaiaie eaiaeeaeaoa aeieiae/aneeo iaoe
ii niaoeeaeueiinoe 03.00.16 yeieiaey. — Eeaaneee oieaa?neoao eiaie
Oa?ana Oaa/aiei, Eeaa, 1999.

Aeenna?oaoeey iinayuaia eco/aieth a o?iie/aneii yenia?eiaioa
ea/anoaaiiie e eiee/anoaaiiie nayce iaaeaeo o?iaiai caa?yciaiey n?aaeu
eiiaie naeioea e ieeaey e i?iaeoeoeeiiiuie iieacaoaeyie aaoi- e
aaoa?io?ioiuo aeinenoai ia i?eia?a eaai?aoi?iuo iiioeyoeee aunoeo
aiaeiuo ?anoaiee e ?ua. Iieo/aiu iiaua aeaiiua ii eciaiaieth
eioaineaiinoe e yooaeoeaiinoe aeii?iaeoeoeeiiiiai i?ioeanna a
oiene/aneie n?aaea. Enneaaeiaaia caaeneiinoue oaeaeueiie nei?inoe ?inoa,
aaeiaie yooaeoeaiinoe eiiaa?oe?iaaiey ei?ia, Iieacaoaey iioeiaeueiinoe
(ii?aaeaeyaony eae i?iecaaaeaiea aeaoo i?aaeuaeoueo ia?aiao?ia) aeey
aaoa?io?ioia e oaeaeueiie nei?inoe ?inoa, /enoie e aaeiaie oaeaeueiie
i?iaeoeoeee, iieacaoaey aeuoaiey, niaea?aeaiey nooiai aauanoaa e
i?aaie/aneiai oaea?iaea, Eioaa?aeueiiai iieacaoaey i?iaeoeoeaiinoe
(ii?aaeaeyaony eae i?iecaaaeaiea oaeaeueiie nei?inoe ?inoa ia
iieacaoaeue /enoie oaeaeueiie i?iaeoeoeee) aeey aaoio?ioia io
eiioeaio?aoeee eiiia naeioea e ieeaey a n?aaea. Oaiaoaiea
i?iaeoeoeeiiiuo iieacaoaeae anao enneaaeiaaiiuo iauaeoia iaaethaeaeinue
i?e iaeneiaeueiuo, aieaa 10 ia/aei3 eiioeaio?aoeeyo eiiia iaoaeeia.
Aieaa ieceea eiioeaio?aoeee iinoiyiii eee a?aiaiii aucuaaee noeioeyoeeth
(ai?iacen) oao eee eiuo i?iaeoeoeeiiii-yia?aaoe/aneeo iieacaoaeae.
Iaeiaei, noeioeyoeey ia?aiao?ia iiae aeaenoaeai caa?yciythueo ?aaaaioia
oaeaea naeaeaoaeuenoaoao i aicieeiiaaiee oiene/aneiai yooaeoa, /oi
naycaii n iaee/eai ?acee/iuo oac ?aaeoeee-ioaaoa aeinenoaiu ia aeaenoaea
oieneeaioa. Yoi aea?o i?aai ooaa?aeaeaoue, /oi iaee/ea eiiia naeioea e
ieeaey a n?aaea a oeaeii iaaaoeaii aeeyao ia aeii?iaeoeoeaiinoue iieiaee
?ua e aeae?iiae?ioeoia. Nieaeaiea i?iaeoeoeeiiiuo ia?aiao?ia aunoeo
aiaeiuo ?anoaiee i?e aic?anoaiee eiioeaio?aoeee ieeaey i?ienoiaeeo
iiiiaeeiaie/ii. Ieeaiaeeiaie/aneia oaiaoaiea
i?iaeoeoeeiiii-yia?aaoe/aneeo iieacaoaeae n oaaee/aieai eiioeaio?aoeee
caa?ycieoaeae ioia/aii aeey enneaaeiaaiiuo ?anoaiee a neo/aa
eniieueciaaiey eiiia naeioea, aeey aeeaioiuo — i?e eniieueciaaiee
eiiia e naeioea, e ieeaey. Aeey eco/aiey aeiciauo caaeneiinoae i?eiaiaia
oi?ioea Aaaa?a. Onoaiiaeaia aiaeeoe/aneay caaeneiinoue iaaeaeo aeiciaie
iaa?oceie (i?iecaaaeaiea eiioeaio?aoeee ia a?aiy aeaenoaey oieneeaioa) e
aeii?iaeoeoeaiinoueth iieiaee ?ua. Iaoaiaoe/anee yoa caaeneiinoue iiaeao
auoue iienaia o?aaiaieai aeaea:

y = Ax — Bx2 + C

aaea o — yoi i?iaeoeoeeiiiue ia?aiao?, o — aeica (aeiciaay iaa?ocea),
A, A, N — aeaeiniaoeeoe/aneea eiinoaiou. Ia iniiaaiee aiaeeca o?aaiaiey
naeaeaii i?aaeiieiaeaiea, /oi n oaaee/aieai aeicu noeioee?othuee yooaeo
aic?anoaao eeiaeii, a eiaeae?othuee eaaae?aoe/ii. Aeey eioa?i?aoaoeee
aeiciauo caaeneiinoae aaoio?ioiuo iauaeoia i?aaeeiaeaia aeiioaoe/aneay
iiaeaeue aa?iiie/aneiai aeieiae/aneiai inoeeeeyoi?a, eioi?ay
?anniao?eaaao i?aaieci (aeiiauaeo), iaoiaeyueeny a ninoiyiee ?aaiiaaney
ni n?aaeie, eae nenoaio, eiathuoth ii?aaeae?iiue caian i?i/iinoe, a
oieneeaio — eae no?anniaue oaeoi?, auaiaeyuee nenoaio ec ?aaiiaaney e
aucuaathuee eieaaaiey aei oao ii?, iiea aai aeaenoaea ia en/aciao eee
i?aauneo a?aieoeu ciiu oiea?aioiinoe iauaeoa ii ioiioaieth e aeaenoaeth
no?anniaiai oaeoi?a e ?ac?ooeo nenoaio. Ia aaoio?ioiuo iauaeoao
i?iaeaiiino?e?iaaii oaaee/aiea oiene/aneiai yooaeoa n oiaiueoaieai
iioiea aeinooiiie yia?aee. Iieacaii, /oi oaaee/aiea eiioeaio?aoeee eiiia
ieeaey aia/aea eiaeae?oao i?iaeoeoeeiiiua i?ioeannu e noeioee?oao
aeano?oeoeeiiiua, a caoai eiaeae?oao iaa i?ioeanna. Onoaiiaeaiu iaeaieaa
/oanoaeoaeueiua e eioi?iaoeaiua i?iaeoeoeeiiii-yia?aaoe/aneea iieacaoaee
— Iieacaoaeue iioeiaeueiinoe e Eioaa?aeueiue iieacaoaeue
i?iaeoeoeaiinoe. Iainiiaaia aiciiaeiinoue eo aeaeueiaeoaai i?eiaiaiey
aeey ioeaiee ea/anoaa aiaeiie n?aaeu ii ooieoeee ioeeeea aeinenoai.

Eeth/aaua neiaa: oyae?eua iaoaeeu, ea/anoai n?aaeu, yeioieneeieiaey,
i?iaeoeoeaiinoue, aeieiaeeeaoeey, i?iaeoeoeeiiii-yia?aaoe/aneea
ia?aiao?u.

SUMMARY

Ignatyuk A.A. The dependence of productive-energy parameters of
biosystems on the level of environment pollution by heavy metals.-
Manuscript.

The dissertation for the degree of Candidate of Biological Sciences on
speciality 03.00.16 — ecology. Taras Shevchenko Kyiv University.
Kyiv,1999.

The dissertation is devoted to study of the qualitative and quantitative
correlation between the level of an environment pollution by lead and
nickel ions and productive indeces of auto- and heterotrophic
biosystems by a chronic experiment within the laboratory populations of
the aquatic Higher Plants and Fishes. The new results have been
obtained on a change of intensity and efficiency of a bioproductive
process in the toxic sphere. The oligodynamic oppression of the
productive-energy indeces in the studied plants and animals results
from the increase of a toxicant concentration. The correlation between
a dose of toxicant and bioproductivity of young fishes has been
defined. According to our supposition, a stimulating effect increases
linearly and inhibiting effect increases squarely if to increase a dose
of pollution. An increasing of a toxicant effect correlates with the
decreasing of a flow of an accessible energy. The most sensible and
informative productive-energy indeces have been established. The
possibility of use of the above indeces for the estimation of the
aquatic sphere quality by the function of biosystems response has been
proved.

Key words: heavy metals, quality of environment, ecotoxicology,
productivity, bioindication, productive-energy indeces.

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