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Вплив вірусів поліомієліту та їх бентонітових варіантів на пухлинні процеси в експерименті: Автореф. дис… канд. біол. наук / Н.В. Ахрамєєва, Нац. ме

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I?i?noa?noai ioi?iie caei?ia’y Oe?a?ie

Iaoe?iiaeueiee iaaee/iee oi?aa?neoao ?iai? I.I.Aiaiiieueoey

Ao?ai??aa Iaoae?y Aieiaeeie??aia

OAeE 578.835.15:578.72:616-006-092.4

AIEEA A??ON?A IIE?II??E?OOOA ?O AAIOII?OIAEO AA??AIO?A IA IOOEEII?
I?IOeANE A AENIA?EIAIO?

03.00.06 – a??onieia?y

Aaoi?aoa?ao

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa

a?ieia?/ieo iaoe

Ee?a – 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia a Iaoe?iiaeueiiio iaaee/iiio oi?aa?neoao?

?iai? I.I.Aiaiiieueoey IIC Oe?a?ie.

Iaoeiaee ea??aiee: Oe?iaieia A.I., ae.i.i., i?ioani?, /eai-ei?. IAI oa
AII Oe?a?ie, caneoaeaiee ae?y/ iaoee ? oaoi?ee Oe?a?ie, caa?aeoth/ee
eaoaae?ith i?e?ia?ieia??, a??onieia?? oa ?ioiieia?? IIO.

Io?oe?ei? iiiiaioe: aeieoi? iaaee/ieo iaoe, i?ioani? ?eaaeei Na?oeaia
Eaiioue?aia, noa?oee iaoeiaee ni?a?ia?oiee, a. i. caa. eaai?aoi???th
i?e?ia?ieia?/iiai eiio?ieth yeino? ?ioiia?ieia?/ieo i?aia?ao?a
Ee?anueeiai iaoeiai-aeine?aeiiai ?inoeoooo ai?aeai?ieia?? oa
?ioaeoe?eieo oai?ia ?i. E.A.A?iiaoaanueeiai

aeieoi? iaaee/ieo iaoe, i?ioani? A?eiaae/ TH??e Aeeiiae/, caneoaeaiee
ae?y/ iaoee, caa?aeoth/ee aenia?eiaioaeueii-eaai?aoi?iei a?aeae?eii
?inoeoooo iieieia?? oa ?aae?ieia??

I?ia?aeia onoaiiaa: Ee?anueea iaaee/ia aeaaeai?y i?neyaeeieiiii? ina?oe
?i. I.Ae.Ooieea IIC Oe?a?ie, i.Ee?a.

Caoeno a?aeaoaeaoueny “ 20 “ o?aaiy 1999 ?. i 13.30 ia can?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae.26.003.01 i?e Iaoe?iiaeueiiio
iaaee/iiio oi?aa?neoao? ?i. I.I.Aiaiiieueoey ca aae?anith: 252057,
Ee?a-57, i?iniaeo Ia?aiiae, 34, iaaeeei-i?io?eaeoe/iee ei?ion,
aoaeeoi??y ?2.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? Iaoe?iiaeueiiai
iaaee/iiai oi?aa?neoaoo ?i. I.I.Aiaiiieueoey ca aae?anith: Ee?a-57,
aoe.Ciieia?/ia,3.

Aaoi?aoa?ao ?ic?neaiee “ 20 ” ea?oiy 1999 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee

aeieoi? iaaee/ieo iaoe, i?ioani?
I.I. ssai?ianueeee

Aeooaeuei?noue ?iaioe

Iieieia?/i? caoai?thaaiiy yaeythoueny iaaecae/aeii aaaeeeaith iaaee/iith
oa nioe?aeueiith i?iaeaiith. Caaaeath/e ia oea, aenia?eiaioaeueia
iaa?oiooaaiiy iiaeo iaoiae?a oa?ai?? iooeei eeoa?oueny aeooaeueiei
caaaeaiiyi. A oaia??oi?e /an i?iaiaeeoueny ?ioaineaiee iiooe iiaeo
i?aeoiae?a aei e?eoaaiiy iieieia?/ieo caoai?thaaiue (A.A. Oeeiiaiei n
niaao., 1998; V Bertone et al.,1997; T Schunck et al, 1998). Iaeiei ?c
oaeeo iaoiae?a ? a??ioa?ai?y. Ieoaiiy i?i canoinoaaiiy a??on?a a
iieioa?ai?? aeine?aeaeo?oueny ? aeeneooo?oueny aaea i?ioyaii aeanyoe??/
– c iiiaioo a?aee?eooy yaeua a??oniiai iieio?ii?cio oa iieie?co.

Aea/aiiy aca?iiae?? a??on?a oa iooeei, ye iiaee iaoeiaee iai?yiie, ii/aa
oi?ioaaoeny a 50-o? ?iee. Aeey a??ioa?ai?? iieicaoai?thaaiue
aei?iaiaoaaeenue a??one ??cieo oaeniiii?/ieo a?oi oa a??oni? aaeoeeie –
a?aia??one, aaeaiia??one, a??one aa?iano, a??on a?niiaaeoeeie,
aioe?aa?/ia aaeoeeia c ??cieie, /anii noia?a/eeaeie, iane?aeeaie. Aea
niaeo? a??on?a, ui iiaeooue aooe aeei?enoai? a i?aeoe/i?e a??ioa?ai??,
iaiaaeo?oueny aeanoeainoyie oeeo a??on?a. Aiie iathoue aooe aiaoiaaiieie
/e iaeiiaoiaaiieie aeey ethaeeie ? ni?e/eiyoe ae?aaeaio ooii??ia?aoth/o
ae?th. C oe??? oi/ee ci?o ia?niaeoeaieie ? aioa?ia??one (AA), ye?
a?aeiinyoueny aei ?iaeeie Picornaviridae, ?iaeo Enterovirus (J.L.
Melnick, 1996). Ee?i?/io ai?iaaoe?th oa i?aeoe/ia canoinoaaiiy
aioa?ia??on?a a iieieia?/i?e ee?i?oe? ?icii/aea I.E.Ai?ioeeiaa (I.E.
Ai?ioeeiaa, 1986), yea noai?eea ?yae aeeaeo aioa?ia??onieo aaeoeei, ui
aeei?enoiaoaaeeny c 1958 ?ieo aeey ?ioi?caoe?? ?aeiaeo oai?eo c iaaiei
iiceoeaiei ee?i?/iei aoaeoii. Ia?aeaeueii c 1960 ?ieo Iooeai??oea A.ss.
?c ni?aaaoi?aie (A.ss. Iooeaieaoea, 1969; A.ss. Iooeaieaoea, 1977; A.ss.
Iooeaieaoea n niaao., 1982) i?iaiaeeee aenia?eiaioaeuei? aeine?aeaeaiiy
a oeueiio iai?yieo, a ?acoeueoao? yeeo aoei aeyaeaii iieio?ii?ci
cia/iiai /enea aioa?ia??on?a aei aeayeeo iooeei ethaeeie ? oaa?ei.

Na?aae AA iniaeeai ?aaeueiei aeey a??ioa?ai?? ? canoinoaaiiy aaeoeeiieo
ooai?a a??on?a iie?ii??e?oo (AAII), oiio, ui aiie ? neeaaeiaeie
eiiiiiaioaie aeeai? iie?ii??e?oii? aaeoeeie Naa?ia, yea
aeei?enoiao?oueny aeey ianiai? ?ioi?caoe?? ethaeae (oiaoi yaeythoueny
aacia/ieie), ?, ia?yaeo c ?ioeie aioa?ia??onaie, i?iyaeythoue iieio?iii?
aeanoeaino? (I.E Ai?ioeeiaa, 1979). C iaeyaeo ia oa, ui aca?iiae?y
a??on?a oa iooeei ia? aeoaea ?iaeea?aeoaeueiee oa?aeoa?, ia?aae oei ye
?aeiiaiaeoaaoe iaai? a??one aeey aeine?aeaeaiiy a iieieia?/i?e ee?i?oe?
iaiao?aeii ??ciia?/ii aea/eoe iaoai?cie ?o aieeao ia ?icaeoie iooeei oa
ia i?aai?ci a oe?eiio.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. A ?iaio?
ocaaaeueiaii ?acoeueoaoe aeine?aeaeaiue, cae?eniaieo ca?aeii c
aea?aeaaieie ieaiiaeie oaiaie IIC Oe?a?ie, a?aioiaeie aeine?aeaeaiiyie,
ye? aoee i?iaaaeai? ia eaoaae?? i?e?ia?ieia??. a??onieia??, ?ioiieia??
IIO i?e o?iainoaaii? IIC Oe?a?ie, Aea?aeaaiiai eii?oaoo ii ai?iouea? ?c
NI?Aeii :

Oaia: “Aieea aeeai? iie?ii??e?oii? aaeoeeie ia ?ioiio nenoaio”

? aea?aea?no?aoe?? 0194V025378.

Oaia: “?ioa?oa?aioe?y i?ae aioa?ia??onaie oa ?ao?ia??onaie ye iiaeeeaee
oeyo i?io?eaeoeee NI?Aeo oa NI?Ae-anioe?eiaaieo ?ioaeoe?e” ?
aea?aea?no?aoe?? 0195V0145.

Iaoith ?iaioe aoei aenia?eiaioaeueia aea/aiiy ?icaeoeo iooeei ?
i?ioean?a, ui ?o noi?iaiaeaeothoue, ia oii? aaaaeaiiy aaeoeeiieo
iie?ia??on?a oa aeine?aeaeaiiy iiaeeeaeo iaoai?ci?a iieiiiaeoethth/i?
ae?? inoaii?o.

Caaaeaiiy aeine?aeaeaiiy:

1. Aea/eoe aieea aaeoeeiieo iie?ia??on?a ia ?icaeoie iooeei ia
i?eeeaae?:

na?eiie 180 (S 180)

ai?aea?ii?aeii? ea?oeeiiie eaaaiue Eueth?na (LL)

a??on?iaeoeiaaiiai eaeeico ?aooa?a (LR)

2. Aeine?aeeoe aieea ia ?icaeoie aenia?eiaioaeueieo iooeei aaioii?oiaeo
aa??aio?a iie?ia??on?a ia i?eeeaae? AAII II oeio.

3. Aecia/eoe aieea aaeoeeiieo a??on?a iie?ii??e?oo ia aeaye? oaeoi?e
ianiaoeeo?/ii? ?acenoaioiino? i?aai?cio ieoae:

i?iaeoeoe?th ?ioa?oa?iio

neioac oaeoi?o iae?ico iooeei

ooieoe?iiaeueio aeoeai?noue ii?iaeueieo e?ea??a

4. Aea/eoe oeueo?ano?oeoo?i? ci?ie a ee?oeiao e?noeiaiai iiceo o ieoae c
eaeeicii ?aooa?a oa a iooeeii?e oeaiei? oaa?ei c ai?aea?ii?aeiith
ea?oeeiiiith eaaaiue Eueth?na i?ae aieeaii aaeoeeiieo iie?ia??on?a

5. Ii??aiyeueii aeine?aeeoe ?ai?iaeoeoe?th aaeoeeiieo iie?ia??on?a a
i?aaiao oa iooeeiieo oeaieiao oaa?ei.

Iaoeiaa iiaecia ?iaioe.

A ?acoeueoao? i?iaaaeaieo aeine?aeaeaiue aeyaeaii i?ioaeoeaiee
iieiiiaeoethth/ee aoaeo ?ioi?caoe?? aaeoeeiieie iie?ia??onaie ia
i?aai?ci oaa?ei c iiaiooai?aiiyie. C’yniaaii, ui aaaaeaiiy AAII III oeio
iaeae?aci?oa aieeaa? ia ?icaeoie eaeeico ?aooa?a oa ea?oeeiiie Eueth?na,
iaoiiaethth/e ca?eueoaiiy na?aaeiuei? o?eaaeino? aeeooy (NOAE) ieoae oa
ciaioaiiy na?aaeiuei? iane iooeei (NII) ? na?aaeiuei? e?eueeino?
iaoanoac?a. AAII II oeio iiceoeaii aieeaa? ia ieoae-iin??a na?eiie
S-180, iaoiiaeth? ca?eueoaiiy ?o NOAE oa ciaioaiiy iane iooeei.

Aia?oa i?iaaaeaii ii??aiyeueia aea/aiiy aieeao AAII ia ??ai? aeo?aeieo
iiioeyoe?e oa ?o aaioii?oiaeo aa??aio?a a?aeiinii ia?aa?ao aeanoiiieo
i?ioean?a. Ia i?eeeaae? AAII II oeio iieacaii, ui i?ioaeoeaiee aoaeo
?ioi?caoe?? aa??aioii Aaaio+ oaa?ei c S 180 oa LL aeuee, i?ae i?e
aeei?enoaii? a??on?a aeo?aeii? iiioeyoe??. Oea iiaea neoaeeoe
iaa?oiooaaiiyi noai?aiiy i?eioeeiiai iiaeo i?aia?ao?a ia iniia?
aaioii?oiaiai aa??aioo Aaaio+ iie?ia??on?a aeey canoinoaaiiy a oa?ai??
iooeei.

I?iaaaeaii eiiieaenio ioe?ieo ae?? AAII ia ?ioiieia?/i? iieacieee, ui
iia’ycai? c i?ioeiooeeiiei caoenoii. Anoaiiaeaii, ui aaeoeeii?
iie?ia??one i?aeaeuothoue ooieoe?iiaeueio aeoeai?noue ii?iaeueieo
e?ea??a, ?iaeoeothoue neioac ?ioa?oa?iio oa oaeoi?o iae?ico iooeei.

Oai?aoe/ia oa i?aeoe/ia cia/aiiy

I?iaaaeai? aeine?aeaeaiiy aeiiiaiththoue ? iiaeeaeththoue ?nioth/?
oyaeaiiy i?i aca?iiae?th a??on?a ? iooeei oiio ui:

eiiieaenia aea/aiiy aieeao AAII ia iooeeiiee i?ioean c aeei?enoaiiyi
a??onieia?/ieo, ?ioiieia?/ieo, aeaeo?iiiii?e?ineii?/ieo iaoiae?a oa
o?aaeeoe?eiiai i?aeoiaeo aei aecia/aiiy i?ioeiooeeiii? aeoeaiino?
a??onieo i?aia?ao?a iieacaei iayai?noue iieiiiaeoethth/eo aoaeo?a, i?iya
yeeo caeaaeeoue a?ae oeio ? aaioii?oiaiai aa??aioo a??on?a oa aeaeo
iooeeie;

iieiiiaeoethth/a ae?y AAII aeiaaaeaia ia eeoa ia ??ai? aeo?aeii?
a??onii? iiioeyoe??, a oaeiae ia ??ai? aaioii?oiaeo aa??aio?a – c
o?aooaaiiyi aeenioe?aoe?? aioa?ia??on?a ia aa??aioe Aaaio+ oa Aaaio
SYMBOL 45 \f “Symbol” \s 13.5 – a i?ioean? ?ai?iaeoeoe??. Ia i?eeeaae?
AAII II oeio aeiaaaeaii, ui aa??aio Aaaio+ (aeii?ioth/ee eiiiiiaio
iiioeyoe??) ia? a?eueo ae?aaeaiee i?ioaeoeaiee aoaeo, i?ae a??one
aeo?aeii? iiioeyoe??. Aa??aio Aaaio SYMBOL 45 \f “Symbol” \s 13.5 –
(i?ii?iee eiiiiiaio iiioeyoe?? aaeoeeiieo ooai?a iie?ia??on?a), iaaiaee,
coiiaeth? a?eueo ?ioaineaiee ??no iooeei.

?ic?iaeaia noaia eiiieaenii? aenia?eiaioaeueii? ioe?iee aieeao a??on?a
ia ?icaeoie iooeei, yeo iiaeia ?aeiiaiaeoaaoe aeey i?iaaaeaiiy
aiaeia?/ieo aeine?aeaeaiue a ee?i?/ieo oiiaao.

Io?eiai? ?acoeueoaoe aeiaiaeyoue, ui AAII yaeythoueny ia?niaeoeaieie
aeey iiaeaeueoiai aea/aiiy ?o oa?aiaaoe/ieo iiaeeeainoae a iieieia?/i?e
ee?i?oe?.

Iniaenoee aianie caeiaoaa/a. Aenia?eiaioaeueia aea/aiiy aieeao
iie?ia??on?a oa ?o aaioii?oiaeo aa??aio?a ia ?icaeoie iooeei,
aeine?aeaeaiiy oaeoi??a ianiaoeeo?/ii-ai caoenoo oaa?ei ia oii?
?ioi?caoe?? oa oaa?ei c iooeeiaie, aeyaeaiiy ?iciiiaeaiiy iie?ia??on?a a
oeaieiao oa i?aaiao ieoae c aeei?enoaiiyi iaoiaeo oioinainea?e?caoe??
a??on?a cae?eniai? aaciina?aaeiuei aaoi?ii ?iaioe ia eaoaae??
i?e?ia?ieia??, a??onieia?? oa ?ioiieia?? IIO. Aeine?aeaeaiiy
oeueo?ano?oeoo?e iooeeiieo ee?oei oa ee?oei e?no-eiaiai iiceo i?iaaaeai?
a eaai?aoi??? aeaeo?iiii? i?e?ineii?? OeIAeE IIO, eiinoeue-oaoe?? oa
aeiiiiiao iaaeaaaea i?io. Noa/aiei E.I., ca ui aeneiaeth?ii ?e ue?o
iiaeyeo.

Ai?iaaoe?y ?iaioe.

Iniiai? iieiaeaiiy ?iaioe aoee aeeeaaeai? ia 10-io i?aeia?iaeiiio
eiia?an? c a??onieia?? (Jerusalem, Israel, 1996), i?aeia?iaei?e iaoeia?e
eiioa?aioe??, i?enay/ai?e 150-??//th c aeiy ia?iaeaeaiiy Ia/ieeiaa (i.
Oa?e?a, 1995), i?aeia?iaei?e iaoeia?e eiioa?aioe?? “Ii?oieia?y –
i?aeoe/i?e aaoa?eia??? oa iaaeeoeei?” (i. A?ea Oea?eaa, 1998),
eiioa?aioe??, i?enay/ai?e 100-??//th a?ae aeiy ia?iaeaeaiiy N.N.Aey/aiea
(i. Ee?a, 1998).

Ioae?eaoe??. Ca oaiith aeena?oaoe?? iioae?eiaaii 13 ?ia?o, a oiio /ene?
7 noaoae, i?aaenoaaeaieo o aeaeaiiyo, ?aeiiaiaeiaaieo AAE Oe?a?ie.

No?oeoo?a oa ianya ?iaioe. Iniiaiee iaoa??ae aeena?oaoe?? aeeeaaeaii ia
169 noi??ieao iaoeiiieniiai oaenoo ? neeaaea?oueny ?c anooio, iaeyaeo
e?oa?aoo?e, 4 ?icae?e?a aeanieo aeine?aeaeaiue, aeniiae?a, nieneo
aeei?enoaieo aeaea?ae (186 a?o/eciyieo oa 96 ?iicaiieo). Aeena?oaoe?eia
?iaioa ?ethno?iaaia 16 oaaeeoeyie oa 12 iaethieaie.

Iniiaiee ci?no aeena?oaoe??.

Iaoa??aee oa iaoiaee aeine?aeaeaiue.

Aeey i?iaaaeaiiy aeine?aeaeaiue aoee aeei?enoai? ae?iaie/? aaeoeeii?
ooaie a??on?a iie?ii??e?oo ? – LSn2ab, ?? – P-712,ch 2ab oa ??? – Leon12
ab oei?a, io?eiai? c ?inoeoooo iie?ii??e?oo oa a??onieo aioeaoae?o?a AII
N?N? a 1983 ?ioe?.

Aea/aiiy iieiiiaeoethth/i? ae?? oeeo a??on?a i?iaiaeeee ia oaeeo
aenia?eiaioaeueieo iooeeiieo iiaeaeyo: na?eiia 180, ai?aea?ii?aeia
ea?oeeiiia eaaaiue Eueth?na oa a??on?iaeoeiaaiee eaeeic ?aooa?a, ye?
aoee etha’ycii iaaeai? no.i.n. E.?. Caai?oeei oa i?io. I.?. Iaoiaiei
(?inoeooo ?aae?ieia?? oa aenia?eiaioaeueii? iieieia?? ?i.
?.Eaaaoeueeiai). Ia?auaieaiiy iooeei oa aea/aiiy aieeao a??on?a ia
?icaeoie aeanoii i?iaiaeeee ca eeane/ieie iaoiaeeeaie, ui
aeei?enoiaothoueny aeey aeine?aeaeaiiy i?ioeiooeeiieo i?aia?ao?a
(A.Aooane n niaao., 1980). Aenia?eiaioe aeey aea/aiiy aieeao
iie?ia??on?a ia ?icaeoie iooeei i?iaiaeeee ca oaeith caaaeueiith
noaiith: cae?enithaaee iiia?aaeith ?ioi?caoe?th oaa?ei a??onai?niei
iaoa??aeii, /a?ac 2 aeiae i?euaiethaaee iooeeie, a i?ioean? ?icaeoeo
aeanoiie i?iaiaeeee iiaoi?io ?ioi?caoe?th a?aeiia?aeieie a??onaie.
?acoeueoaoe ioe?ithaaee ca na?aaeiueith ianith iooeei (aeey oeueiai
ieoae caaeaaee oa aeeo/aee iooeeie) oa na?aaeiueith o?eaae?noth aeeooy
oaa?ei. Ia iiaeae? ea?oeeiiie Eueth?na aecia/aee e?euee?noue iaoanoac?a.

Aeey aeine?aeaeaiiy iieiiiaeoethth/iai aoaeoo a??on?a iie?ii??e?oo ia
??ai? iiioeyoe?? iiia?aaeiuei io?eioaaee aaioii?oia? aa??aioe Aaaio+ oa
Aaaio SYMBOL 45 \f “Symbol” \s 13.5 – , ia ye? aeenioe?ththoue
aioa?ia??one a i?ioean? ?ai?iaeoeoe??. Aeey naeaeoe?? aaioii?oiaeo
aa??aio?a ?c iiioeyoe?e a??on?a aoei canoiniaaii niaoe?aeueiee iaoiae
(A.I. Oe?iaieia, 1977). Eiai i?eioeei iieyaa? a oiio, ui aeii?ioth/a
/anoeia iiioeyoe?? iie?ia??on?a aaeni?ao?oueny ia aaioii?o? i?e
iaeo?aeueieo aai neaaieoaeieo ?I, oiio oeae aa??aio iaea?aeaa neiaie
Aaaio +. I?ii?ia /anoeia iiioeyoe?? – oea a??one, ye? ia aaeni?aothoueny
ia aaioii?o? i?e aeaieo ?I. Aiie io?eiaee iacao aa??aio Aaaio SYMBOL 45
\f “Symbol” \s 13.5 – . Aeey naeaeoe?? aeei?enoiaoaaee 0,5% noniaic?th
aaeth aaioii?oo, yeee aeiaeaaaee aei a??onai?niiai iaoa??aeo (?I
7,2-7,6). Aai-oii?o c aaeni?aiaaieie a??onaie iaaeae? aeei?enoiaoaaee ye
aeaea?aei aa??aioo Aaaio+, a noia?iaoaio i?aeaeaaaany ua 6 aeniaaeaiiyi
aaeai aaioii?oo ? i?noea aa??aio Aaaio SYMBOL 45 \f “Symbol” \s 13.5 – .
Iio?i i?iaiaeeee ianaae? a??on?a iaio aa??aio?a ia ee?oeiieo eoeueoo?ao.
Io?eiaiee a?-?onai?niee iaoa??ae iiaoi?ii ia?iaeyee aaioii?oii i?e ?I
7,2-7,6 ? eiioeaio?oaaee.

Iiaeeeaeie iaoai?ciaie aieeao a??on?a ia ia?aa?a aeanoiiieo i?ioean?a ?
i?yiee iieie?c, iaoiiaeaiee iieio?ii?ciii ? ?iciiiaeaiiyi a??on?a a
iooeeii?e oeaiei?, a oaeiae a??on?iaeoeiaaia ?ioiiiiaeoeyoe?y.

C iaoith iaea?aeaiiy aeieac?a ?ai?iaeoeoe?? iie?ia??on?a a iooeeiao oa
?ioeo i?aa-iao oaa?ei aoa aeei?enoaiee oaiiiai oioinainea?e?caoe??
aioa?ia??on?a i?ioeaa?iii (A.A. Ei?ieaaa, 1980). Aeneiaeth?ii ue?o
iiaeyeo i?io. A.A.Ei?ieueia?e ca eiinoeue-oaoe?? oa iaaeaiiy i?aia?aoo
i?ioeaa?io. Ia?iaeai? i?ioeaa?iii a??one iaaoaathoue aea-noea?noue
?iaeoeaoaaoeny i?ae ae??th na?oea. Oea ia aaiaoe/iee oaiiiai, oiio ia
a??onia iioiinoai a?i ia ?iciianthaeaeo?oueny. Iiioeyoe?th a??on?a
ia?aae ?ioi?caoe??th ia?iaeythoue i?ioeaa?iii. A i?ioean? ?ai?iaeoeoe??
a i?aai?ci? oaa?ei iauaaeee ia?iaeaieo i?ioeaa?iii a??on?a ao?a/athoue
oioi/ooeea?noue. I?ney aeae?eaiiy c i?aai?a /anoeio iaoa??aeo ia?iaeyee
na?oeii ? i?iaiaeeee ii??aiyeueia oeo?oaaiiy. Aeyaeaiiy ?ioaeoe?eieo
a??on?a a /anoei? AAAI, ui aoea ia?iaeaia na?oeii, na?ae/eoue i?i
?ai?iaeoeoe?th aioa?ia??on?a a i?aaiao oaa?ei. Aeyaeaiiy ?ioaeoe?eieo
a??on?a o?eueee a /anoei? iaoa??aeo, ia ia?iaeai?e na?oeii, na?ae/eoue
i?i oa, ui oea a??one aeo?aeii? iiioeyoe??, oiaoi a??one, ye? eeoa
i?iieeaee a oeaieie, aea ia ?iciiiaeoaaeeny a ieo. Aeey i?iaaaeaiiy
aenia?eiaioo oaa?eiai-iooeeiiiin?yi aaiaeeee ia?iaeai? i?ioeaa?iii
a??one. Aeae?eaiiy a??on?a i?iaiaeeee /a?ac 48 aiaeei i?ney ?ioi?caoe??.

C iaeyaeo ia oa, ui iniiaio ?ieue a i?ioeiooeeiiiio caoeno?
a?ae?a?athoue ianiaoeeo?/i? oaeoi?e, aoei aeine?aeaeaii aieea
?ioi?caoe?? aaeoeeiieie iie?ia??onaie ?ioaeoieo ieoae oa
oaa?ei-iooeeiiiin??a ia oae? iieacieee ye ooieoe?iiaeueia aeoeai?noue
ii?iaeueieo e?ea??a (IE), neioac oaeoi?o iae?ico iooeei (OII) oa
?ioa?oa?iio (?OI). Aeey oeueiai ?ioaeoiei oaa?eiai aaiaeeee a??one
a?aeiia?aeiiai oeio a e?eueeino? 1o107 OOeAe50. *a?ac 1, 4, 7 oa 14 ae?a
i?ney ?ioi?caoe?? oaa?ei caaeaaee, a?aee e?ia oa naeac?iee aeey
iiaeaeueoeo aeine?aeaeaiue. C iaoith aea/aiiy aieeao iie?ia??on?a ia
?aaeoeai?noue iooeeiiiin??a oaa?ei ?ioi?coaaee, /a?ac 24 aiaeeie ?i
i?euaiethaaee iooeeie. Acyooy iaoa??aeo aeey aeine?aeaeaiue
cae?enithaaee /a?ac 14 ae?a i?ney ?iieoeyoe?? iooeeiieo ee?oei (LL oa S
180) /e /a?ac 22 aeiae i?ney aaaaeaiiy a??ono eaeeico ?aooa?a (VLR).

Aecia/aiiy aeoeaiino? ii?iaeueieo e?ea??a i?iaiaeeee eiei?eiao?e/iei
iaoiaeii (I.A. Eociaeiaa, 1991). Aeey oeueiai aeei?enoiaoaaee
nieaiioeeoe ieoae, ye? i?iyaeythoue aeoeai?noue ii?iaeueieo e?ea??a.
Oeth aeoeai?noue aeyaeyee ii oeeoioiene/i?e ae?? ia /ooeea? ee?oeie
L-929. Ioe?ieo oeeoie?co i?iaiaeeee c aeei?enoaiiyi a?oaeueiiai
aa?aieea, yeee aiineee a eoeueoo?o ee?oei-i?oaiae aei ia?iaee ?o IE.
I?ney ?oeioaaiiy /anoeie ee?oei i?ae ae??th NK c ieo aea?eueiy?oueny
aa?aiee. Iaoiaeii niaeo?ioioiiao??? aecia/aee ?ioaineai?noue caaa?aeaiiy
eoeueoo?aeueii? ??aeeie aeine?aeieo oa eiio?ieueieo i?ia (ia ia?iaeaieo
nieaiioeeoaie) ? ioe?ithaaee oeeoioiene/iee ?iaeaen.

Aecia/aiiy ??aiy OII a ieaci? e?ia? iniiaaia ia aeyaeaii? oeeoie?oe/ii?
ae?? OII ii a?aeiioaiith aei /ooeeaeo ee?oei L-929. Aeey oeueiai
ee?oeiio noniaic?th ?iceeaaee a 96-eoieia? ieaioaoe ? i?ney 24 aiaeei
?ieoaaoe?? aeiaeaaaee aeoeiii?oeei oa aeine?aei? ne?iaaoee. *a?ac 24
aiaeeie ?ieoaaoe?? a NI2-?ieoaaoi??, ceeaaee na?aaeiaeua, ui i?noeei
c?oeiiaai? i?ae ae??th OII ee?oeie, ? aeiaeaaaee 0,2% ?ic/ei
e??noae-a?ieaoo aeey caaa?aeaiiy ee?oei, ui caeeoeeeny a eoieao. Ia
niaeo?ioioiiao?? aecia/aee ?ioaineai?noue caaa?aeaiiy oa i?ae?aoiaoaaee
oeeoioiene/iee ?iaeaen.

Aeey aecia/aiiy ?ioa?oa?iio aoei aeei?enoaii 2 i?aeoiaee: aeyaeaiiy
ne?iaao-eiaiai ?OI, ?iaeoeiaaiiai iie?ia??onaie oa iooeeiaie ? aeyaeaiiy
?OI, neioaciaaii-ai nieaiioeeoaie i?ae ae??th ?iaeoeoi??a (eiieaiaaae?io
A – SYMBOL 103 \f “Symbol” \s 14 g -?OI oa a??ono oai?iae Iueth-eanea –
SYMBOL 97 \f “Symbol” \s 14 a -?OI). I?eioeei iaoiaeo iieyaa? a oiio,
ui ?OI ni?aaey? i?ioaeoeaio ae?th ii a?aeiioaiith aei ee?oei,
?io?eiaaieo a??onaie. Oeae aoaeo ioe?iththoue ii iaeo?ae?caoe?? OeIA
i?i?ioi ia 50%. Aeey aecia/aiiy ?OI ieoae canoiniaoaaee
caaaeueiii?eeiyoo nenoaio: eoeueoo?a ee?oei L-929 oa a??on aac?eoey?iiai
noiiaoeoo. Ue?i aeyeo?ii a. i.n. eaa. i?e?ia?ieia?? oa a??onieia?? IAe?
ioiei??iaieia?? ?i.I.?.Eieiiee/aiea, E?eaioaonuee?e E.Ae. ca etha’ycia
iaaeaiiy iaiao?aeieo ?aaeoea?a oa eiinoeueoaoeaio aeiiiiiao.

Aeey aeaeo?iiii-i?e?ineii?/ieo aeine?aeaeaiue oeueo?ano?oeoo?e iooeeiieo
ee?oei ieoae aeei?enoiaoaaee eeane/io iaoiaeeeo i?eaiooaaiiy
aeaeo?iiiii?e?ineii?/ieo i?aia?ao?a. Iaoa??ae o?enoaaee o ?ic/ei? OsO4
ii Eieo?eueaeo. C?acee ciaaiaeithaaee a aoaiie? c?inoath/i?
eiioeaio?aoe??, ii/eiath/e c 700 nie?oo, aoeaoiio oa caeeaaee o noi?o
aiiio oa a?aeae?oo ca caaaeueiia?aeiiith noaiith (A.ss.Ea?oio, 1984).
C??ce aeaioiaeyee ia oeueo?aoiiao LKB-O oa Reichert ? eiio?anooaaee 2%
?ic/eiii o?ai?eaoeaoaoo o 70% aoaiie? oa oeeo?aoii naeioey oa aea/aee a
aeaeo?iiieo i?e?ineiiao AIA-100A, AIA-100A?. Ioe?ieo ci?i
ii?oiooieoe?iiaeueii? i?aai?caoe?? ee?oei i?iaiaeeee ca?aeii iaoiaeai
(A.I.Ni?ieei, 1989; R.Duke et al., 1996; S.Bergese et al., 1997).

Io?eiai? aenia?eiaioaeuei? aeai? ia?iaeyee iaoiaeii aa??aoe?eii?
noaoenoeee c aecia/aiiyi e?eoa??th No’thaeaioa (Ieioea? I.I., 1982).

?acoeueoaoe aeine?aeaeaiiy oa ?o iaaiai?aiiy.

I?e aea/aii? iieiiiaeoethth/i? aeoeaiino? aaeoeeiieo a??on?a
iie?ii??e?oo anoaiiaeaii, ui an? 3 oeie a??on?a caeaoi? aieeaaoe ia
?icaeoie aenia?eiaioaeueieo iooeei. Oa?aeoa? oa ?ioaineai?noue ae??
caeaaeeoue ye a?ae oeio a??ono, oae ? a?ae aeaeo iooeeie.

Aeine?aeaeaiiy, i?iaaaeai? ia iiaeae? na?eiie 180 a?aeia?aaeai? a oaae.
1.

Oaaeeoey 1.

Aieea aaeoeeiieo iie?ia??on?a ia ?icaeoie na?eiie 180 o ieoae.

A??one Na?aaeiy iana iooeeie Na?aaeiy o?eaae?noue aeeooy

aaniethoia

(M SYMBOL 177 \f “Times New Roman” \s 14 ± m, a) a?aeiinii eiio?ieth

(a %)

aaniethoia

(a aeiaao) a?aeiinii eiio?ieth

(a %)

aac ?ioi?caoe??

(n=10) 4,42 SYMBOL 177 \f “Symbol” \s 14 ± 0,53 100 17,79 SYMBOL 177
\f “Symbol” \s 14 ± 0,11 100

I oei (n=10) 5,57 SYMBOL 177 \f “Symbol” \s 14 ± 0,56

? SYMBOL 62 \f “Symbol” \s 13.5 > 0,1 126 18,48 SYMBOL 177 \f “Symbol”
\s 14 ± 0,07

? SYMBOL 62 \f “Symbol” \s 13.5 > 0,1 103

II oei, (n=10) 2,21 SYMBOL 177 \f “Symbol” \s 14 ± 0,35

? SYMBOL 60 \f “Symbol” \s 13.5 0,1 100,7

I?ei?oea: P- ??aaiue cia/ouino? ii a?aeiioaiith aei eiio?ieueii? a?oie
(ia?ioi?ciaai? oaa?eie).

Iaea?eueoee aieea ia ?icaeoie iooeeie ni?e/eiy? AAII II oeio,
?ioi?caoe?y yeei aeeeeea? cao?eieo ?inoo na?eiie, i?iaeiaaeo?
o?eaae?noue aeeooy oaa?ei ia 44% ? ciaioo? iano iooeei a na?aaeiueiio a
2 ?ace ii??aiyii c eiio?ieueiith a?oiith. AAII III oeio ni?ey?
ciaioaiith na?aaeiuei? iane iooeei a 1,65 ?ace, aea ia aieeaa? ia NOAE.
Ae?y AAII I oeio a?ae??ciy?oueny oei, ui a?i ia eeoa ia aieeaa? ia
o?eaae?noue aeeooy ieoae, a iaa?oue ni?ey? i?enei?aiith ?inoo iooeeie oa
iaoiiaeth? ca?eueoaiiy ?? iane ia 26 % ii??aiyii c eiio?ieai.

Aieea aaeoeeiieo a??on?a iie?ii??e?oo ia ia?aa?a ea?oeeiiie Eueth?na
i?aaenoaaeaiee a oaae. 2.

Oaaeeoey 2.

Aieea aaeoeeiieo iie?ia??on?a ia ?icaeoie ea?oeeiiie Eueth?na o ieoae

A??one Na?aaeiy iana

iooeei Na?aaeiy e?euee?noue iaoanoac?a Na?aaeiy o?eaae?noue aeeooy

aaniethoia

(M SYMBOL 177 \f “Symbol” \s 14 ± m, a) a?aeiinii eiio?ieth

(a %) aaniethoia a?aeiinii eiio?ieth

(a %) aaniethoia

(a aeiaao) a?aeiinii eiio?ieth

(a %)

aac ?ioi?caoe??

(n=10) 2,03 SYMBOL 177 \f “Symbol” \s 14 ± 0,18 100 1,88 SYMBOL 177 \f
“Symbol” \s 14 ± 0,49 100 29,1 SYMBOL 177 \f “Symbol” \s 14 ± 0,92
100

? oei

(n=10) 1,66 SYMBOL 177 \f “Symbol” \s 14 ± 0,31

? SYMBOL 62 \f “Symbol” \s 13.5 > 0,1 81,8 2,88 SYMBOL 177 \f “Symbol”
\s 14 ± 0,62

? SYMBOL 60 \f “Symbol” \s 13.5 0,1 93,5

?? oei

(n=10) 1,36 SYMBOL 177 \f “Symbol” \s 14 ± 0,24

? SYMBOL 60 \f “Symbol” \s 13.5 0,1

II oei

(n=21) 240,0 SYMBOL 177 \f “Symbol” \s 13.5 ± 25,76

?0,1 12,1 SYMBOL 177 \f “Symbol” \s 13.5 ± 1,13

?

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