OA?E?ANUeEEE AeA?AEAAIEE OI?AA?NEOAO

EA??I? IAAeOA

OAeE 616.89-008.441.13:612.821.8:615.214

AIEEA IAAeEO?IIAEO IA OOIEOe?IIAEUeIEE NOAI IICEIAI? NENOAIE IICEOEAIIAI
I?AeE??IEAIIss O UO??A C AOI?EIII IAAOOEIE IIOEAAOe?ssIE
(AENIA?EIAIOAEUeIEI AEEIAIE?CIII)

03.00.13 – o?c?ieia?y ethaeeie oa oaa?ei

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Oa?e?a – 1999

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia ia eaoaae?? o?c?ieia?? ethaeeie oa oaa?ei Oa?e?anueeiai
aea?aeaaiiai oi?aa?neoaoo I?i?noa?noaa ina?oe Oe?a?ie ? o eaai?aoi???
iae?io?c?ieia?? oa ?ioiieia?? Oe?a?inueeiai IAe? ee?i?/ii? oa
aenia?eiaioaeueii? iaa?ieia?? oa ineo?ao??? IIC Oe?a?ie

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, i?ioani?

Ai?iaeiaa Oaia?a Ieoaee?aia,

caa?aeoaa/ eaai?aoi???th iae?io?c?ieia?? oa ?ioiieia??

Oe?a?inueeiai IAe? ee?i?/ii? oa aenia?eiaioaeueii? iaa?ieia?? oa
ineo?ao??? IIC Oe?a?ie.

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani?

Aeaaeeiaa Aeea ?aai?aia,

caa?aeoaa/ a?aeae?eii ?ai?iaeoeoeaii? aiaeie?eiieia?? Oe?a?inueeiai IAe?
oa?iaeieia?? aiaeie?eiieo caoai?thaaiue IIC Oe?a?ie.

aeieoi? iaaee/ieo iaoe, i?ioani?

Nin?i ?aai Eocueie/,

caa?aeoaa/ eaoaae?ith ia?eieia?? Oa?e?anueei? iaaee/ii? aeaaeai??
i?neyaeeieiiii? ina?oe IIC Oe?a?ie.

I?ia?aeia onoaiiaa: ?inoeooo o?c?ieia?? ?i. aeaae. I.I. Aiaiiieueoey IAI
Oe?a?ie, a?aeae?e o?c?ieia?? aieiaiiai iiceo.

Caoeno a?aeaoaeaoueny «24 “ eenoiiaaea 1999 ?. i 1515 aiaeei?

ia can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee E 64.051.17 i?e
Oa?e?anueeiio aea?aeaaiiio oi?aa?neoao? I?i?noa?noaa ina?oe Oe?a?ie ca
aae?anith: 310077, i. Oa?e?a, i. Naiaiaee, 4, aoaeeoi??y III-15.

C aeena?oaoe??th iiaeia iciaeiieoeny o Oeaio?aeuei?e iaoeia?e
a?ae?ioaoe? Oa?e?anueeiai aea?aeaaiiai oi?aa?neoaoo I?i?noa?noaa ina?oe
Oe?a?ie ca aae?anith: 310077, i. Oa?e?a, i. Naiaiaee, 4.

Aaoi?aoa?ao ?ic?neaiee «21.10. 1999 ?.”

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee

eaiaeeaeao a?ieia?/ieo iaoe,

noa?oee iaoeiaee ni?a?ia?oiee Iaaeaeei A.?.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeueiinoue oaie. O /enei aaaeeeaeo i?iaeai no/anii? iae?iaiieiai? ?
i?iaeaia aoi?eiii iaaooeo iioeaaoeie. ?? aeooaeueiinoue coiiaeaia
iiaeaeuaiiyi ?iciianthaeaeaiinoi a aaaaoueio e?a?iao oaeeo ooo/ieo
iioeaaoeie, ye iioya aei aeeiaieth, ii?oiio, ineoiaeoeaieo niieo/aiue
(Iyoieoeeay E.I., 1994; Aieioei I.A., 1997; Neieoeeee A.I., 1998;
*oi?eeia A.I., 1998; Aa?aociaa N.I., 1998).

Aiaeiii, ui aoi?eiii iaaooi iioeaaoei? oaeiae, ye i i?e?iaeii,
aiieiai/ii, iathoue oie aea iae?iaiieiai/iee noano?ao, oi ae
iiioeithth/i iiceiai iaoaiicie iioaa?aoei? oeeo iioyaia. Iieeaaeath/enue
ia aeiiiioth/i o eioa?aoo?i eiioeaioei? i?i i?e?iaeo ii?iaeueieo
aiieiai/ieo iioeaaoeie (Noaeaeia E.A., 1971; Neiiiia I.A., 1981; Ieieia
A.A., 1997; Stein, 1971), iiaeia c oeo aea iiceoeie ?icaeyaeaoe i
aoi?eiii iaaooi iioyae, yei aiae?iciythoueny ia nai?th nooiinoth, a
iai?e?iaeiith ni?yiiaaiinoth, yea i?ecaiaeeoue aei iaaaeaeaaoii?
iiaaaeiiee.

Caiaeii eioa?aoo?ieo aeaieo, iioya aei aeeiaieth oi?io?oueny ia iniiai
no?oeoo?ii-ooieoeiiiaeueii? i?aaiicaoei? iiceiai? nenoaie iiceoeaiiai
iiaee?iieaiiy, yea a iaaieo oiiaao neoaeeoue aiieiai/iith iao?eoeath
iiaoaeiae ooieoeiiiaeueii? nenoaie iioyao aei aeeiaieth ye oaeoi?o
iiaee?iieaiiy (Ai?iaueaaa O.I., 1977; Aa?/aiei I.A., 1992; Ai?iaueaaa
O.I., Aieioei I.A., Iaeeiaa E.I. e niaao., 1993).

Cia/ia iinoea a iaoaiiciao oi?ioaaiiy aeeiaieueii? iioeaaoei? caeia?
ii?ooaiiy iaoaaieicio aiiaaiieo iiiiaiiiia — eaoaoieaiiiia (Aiioeia
E.I., 1981; Aieei O.I., 1984, 1986), na?ioiiiio (A?iiiaa A.A., 1984;
Oeoeiaa A.I., 1984; Aiyiiae/ A.A., 1984; Naeeiia ?.I. n niaao., 1995;
Aeaeeineay E.A., 1998; Penn ?.A et al., 1978; Mc Bride et al., 1994;
Fahlke N. et al., 1994), a oaeiae iae?iiaioeaeia ( Aiioeia E.I., 1988;
Aoia?ei E.I., 1990; Oaaoe I.I., 1996; Noaeaeia N.E., 1996; Volpicelli
I.R et al., 1991; Sinclair I.D. et al., 1994).

Iniothoue ia?aeiieeai aeaii i?i oa, ui iiceoeaii aiioeiiaaiii no?oeoo?e
iiceo, aea oe?iei i?aaenoaaeaii cacia/aii aeua iae?iiaaeiaoi?e, ia?oeie
?aaaothoue ia i?eeiaiiy aoaiieo oa oiii/ii aaaioe, ui aeeeeeathoue
aaaeaiiy ?o iiaoi?iiai aaeeaaiiy (Ai?iaueaaa O.I., 1976, 1995;
Oaiaiaoaeee I.I. n niaao., 1992; Caa?oao Y.Y., 1993; Iyoieoeeay E.I.,
1994; Carlson, R.H., Lydic, 1976; Hunter B.E. et al., 1978; Dyr W. et
al., 1993), oiaoi aeayei i?aia?aoe, yei ao?o/athoueny o iaoaaieici
iiiiaiiiia oa iae?iiaioeaeia, naii aeeeeeathoue aoi?eiii i?eno?anoi,
oi?ioaaiiy iiaeo aai caieueoaiiy iaaooeo iioeaaoeie.

Iieacaii, ui oi?ioaaiiy iioyao aei aoaiieo caeaaeeoue i aiae eiai
aeaniiai iaoaaieicio a i?aaiicii ethaeeie i oaa?ei. Aoaiie — i?e?iaeiee
iaoaaieio i eiai i?enei?aiee iaiii iiaea noaoe iaeiei c oaeoi?ia
iiaeneeaiiy eiai aeciaaiiiai aaeeaaiiy (Ino?ianeee TH.I., 1981; Aiioeia
E.I., 1981; Iyoieoeeay E.I., 1984). Oei oyaeaiiy eyaee a iniiao iiooeo
iiaeo oiii/ieo niieo/aiue, ni?yiiaaieo nai?th aei?th ia iaoaaieici
aoaiieo. O /enei oaeeo niieo/aiue i?eaa?oa? oaaao iaaeeo?iiae, ui aoa
neioaciaaiee a Oe?a?ii. Oea aaaaoieiiiiiaioiee i?aia?ao —
aetheiciaeioeeioi?iiao, yeee ia? aeanoeainoi ca’ycoaaiiy
aoeaoaeueaeaaiaeo — iaeaieueo oiene/iiai iaoaaieioo aoaiieo.
Iiaeoeaoth/e aoeaoaeueaeaaiae, iaaeeo?iiae i?eaii/o? iaeei c iaoaiiciia
iioeaaoei? aei iueiai (Aoeue I.O. n niaao., 1990, 1998; Neieoeeee A.I.,
1990, 1996). Ii?yae c oeei, oeae i?aia?ao ni?ey? neioaco aiiaaiieo
iiiiaiiiia (Neieoeeee A.I., 1990), ?aaoeth? aiino oa aeoeaiinoue
na?ioiiiio (Ia?eiciaa E.I., 1997; Aiaeei A.O. n niaao., 1997).

?acii c oeei, iiaeeaeaii aeineiaeaeaiiy aieeaia iaaeeo?iiaeo ia
iae?ioiciieiai/ii iaoaiicie ?icaeoeo iioyao aei aeeiaieth a eioa?aoo?i
aiaenooii. O oie aea /an ?o ciaiiy oyaey?oueny iaiaoiaeiei ia oieueee o
oai?aoe/iiio aniaeoi i?iaeaie iaaooeo iioeaaoeie, eiie?aoii, iioyao aei
aeeiaieth, aea i aeey ae?iaeaiiy iioeiaeueieo aa?iaioia ?o eoiithaaiiy
oa i?eaii/aiiy.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. Aeineiaeaeaiiy
i?iaaaeaii a eaai?aoi?i? iae?ioiciieiai? oa iioiieiai? Oe?a?inueeiai
iaoeiai-aeineiaeiiai iinoeoooo eeiii/ii? oa aenia?eiaioaeueii?
iaa?ieiai? oa ineoiao?i? a ?aieao eiiieaenieo IAe?: oeo? 10.37.98
«?ic?iaeoe oaoiieiaith ioeiiee ?eceeo aaeeeoeaii? iiaaaeiiee i
noai?aiiy i?ia?ai ia?aeiii? oa aoi?eiii? i?ioieaeoeee» (? 0198V007888),
Iaoeiiiaeueii? i?ia?aie «Iaaeeei-aiieiai/ii oa nioeiaeueii-ineoieiai/ii
aniaeoe ia?eiiaiie» (oeo? 01.20.02/005E-95), ?icaeie «Aea/eoe
iae?iaiieiai/ii iniaeeainoi no/anieo oi?i ia?eiiaiie oa aeeiaieicio i
?ic?iaeyoe a aenia?eiaioi iiaeoiaee aei ?o eoiithaaiiy».

Iaoa i caaea/i aeineiaeaeaiiy. Iaoith aeineiaeaeaiiy ? c’ynoaaiiy
iae?ioiciieiai/ieo iaoaiiciia aieeao iaaeeo?iiaeo ia aiioeieio
iiaaaeiieo oa aeaeo?e/io aeoeaiinoue iiceo o iioaeoieo oaa?ei oa c
iiaeaethaaiiyi ooo/ii iaaooiai iioyao aei aeeiaieth.

O aiaeiiaiaeiinoi c iaoith aeineiaeaeaiiy ae?iooaaeeny ianooiii caaea/i:

1) aea/aiiy aieeaia iaaeeo?iiaeo ia ?aaeoeith naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono oa iiaaaeiieo iioaeoieo oaa?ei;

2) aea/aiiy aieeaia iaaeeo?iiaeo ia ?aaeoeith naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono oa iiaaaeiieo uo?ia c 14-oe aeaiiith
aeeiaieicaoei?th;

3) aea/aiiy aieeaia iaaeeo?iiaeo ia ?aaeoeith naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono o uo?ia c 6-oe iiny/iith aeeiaieicaoei?th;

4) aea/aiiy aieeaia iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo
iioaeoieo oaa?ei;

5) aea/aiiy aieeaia iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo uo?ia c
14-oe aeaiiith aeeiaieicaoei?th;

6) aea/aiiy aieeaia iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo uo?ia c
6-oe iiny/iith aeeiaieicaoei?th;

7) aea/aiiy aieeaia iaaeeo?iiaeo ia na?oeaai-noaeeiio nenoaio o
iioaeoieo i aeeiaieiciaaieo uo?ia;

8) i?iaaaeaiiy ii?iaiyeueiiai aiaeico oa ocaaaeueiaiiy ?acoeueoaoia
aieeao iaaeeo?iiaeo ia aeyaeaii iieacieee o iioaeoieo oa aeeiaieiciaaieo
uo?ia.

Iaoeiaa iiaecia i oai?aoe/ia cia/aiiy ?iaioe. Aia?oa iieacaii, ui iiaee
i?aia?ao iaaeeo?iiae (aetheiciaeioeeioi?iiao) ia? noeioethth/ee aieea ia
ooieoeiiiaeueiee noai iiceiai? nenoaie iiceoeaiiai iiaee?iieaiiy (ui
aeyaey?oueny o iiaeaeuaiii /anoioe naiinoeioethaaiiy eaoa?aeueiiai
aiiioaeaiono), ia?aaoaeiao? aeaeo?iaaiac iiceo o aie oaeaeeeo ?eoiia oa
i?eaii/o? ?o ia?ieneciaeueii i?iyae.

C’yniaaii, ui aeeiaieicaoeiy uo?ia i?ioyaii 14-oe aeiia i?eaiaeeoue aei
oi?ioaaiiy aeeiaieueii? aeiiiiaioe o iiaaaeiioei. Iicaaaeaiiy /a?aiai?
aeice aoaiieo aeoeao? o oeeo uo?ia i?ioean naiinoeioethaaiiy iiceo,
aeeeeea? aeaneio?iiicaoeith aiiiioaioeiaeia o iaiei?oaeni oa noaeiiio
aeoeaiinoue o eiiai/ieo aiioeiiaaiieo no?oeoo?ao iiceo c iiioeiaoei?th
?? a aiiioaeaioni oa n.accumbens septi. Aia?oa anoaiiaeaii, ui o oeeo
uo?ia iaaeeo?iiae i?eaii/o? aeeiaieueio iioeaaoeith /a?ac ?iceo
aecaeueoaoeith /anoioe ?aaeoei? naiinoeioethaaiiy, neio?iiicaoeith
aiiiioaioeiaeia iiceo (oaoa- i aeueoa-), ?iciiaeie i cieeiaiiy noaeiiii?
aeoeaiinoi, ?? caiiuaiiy ia?ieneciaeueiith a n.accumbens septi. Iiae
eiai aieeaii aeieeathoue iiai no?oeoo?ii-ooieoeiiiaeueii «?iaioie/?
ainaiaei», ui caaacia/othoue ?icaeoie aeaeo?ia?aoi/iiai oa
iiaaaeiieiaiai nio c aiaeiiaith uo?ia aiae aeeiaieth.

O uo?ia c o?iii/iith 6-oe iiny/iith aeeiaieicaoei?th iaaeeo?iiae
ia?aaoaeiao? i?ioean naiinoeioethaaiiy c aeoeaiei oeieani?yiiaaiei
iiooeii, iaeeaeaiiyi oiiaii-?aoeaeoi?iiai iaaeeo naiinoeioethaaiiy i
aeaneio?iiicaoei?th aiiiioaioeiaeia o iaiei?oaeni, ei?aeyoei?th ii
aaoa-?eoio a aiioeiiaaiieo no?oeoo?ao, i?eaii/o? noaeiiio aeoeaiinoue.

Aia?oa iieacaii, ui iaaeeo?iiae iaaea? ii?iaeicoth/i? aei? ia
na?oeaai-noaeeiio nenoaio aeeiaieiciaaieo uo?ia c iiiiaeaiiyi
niiaaiaeiioaiue ?? neiiaoe/ii? oa ia?aneiiaoe/ii? ?aaoeyoeie.

Aeieacaii, ui iaaeeo?iiae i?eaii/o? iioya aei aeeiaieth /a?ac
aeoeaoth/ee aieea ia iiceiaee noano?ao iiceoeaiiai aiioeieiiai
iiaee?iieaiiy oa neiaeiai i?ioeane, aioeaiieaioiaaiio i ianiaoeeoi/io
aeoeaoth/i nenoaie iiceo.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoaoia. ?acoeueoaoe ?iaioe iathoue
aaciina?aaeith eeiii/io ni?yiiaaiinoue, ?ice?eaathoue iaoaiicie
oa?aiaaoe/ieo aoaeoia aieeaia iaaeeo?iiaeo, ui ni?ey? eeiiioeenoai o
aeai?i oaeoeee eieoaaiiy oai?eo ia aeeiaieici.

Iniaenoee aianie caeiaoaa/a. ?iaioa c eioa?aoo?ieie aeaea?aeaie,
aenia?eiaioe c aeei?enoaiiyi noa?aioaene/ii? oaoiiee, iaoiaeia
naiinoeioethaaiiy iiceo, ?a?no?aoei? aeaeo?e/ii? aeoeaiinoi iiceo oa
aeaeo?ioiciieiai/ieo iieacieeia, a?oa?iaeueiiai oeneo caeieniaii
naiinoieii. Aiaeic ?o ?acoeueoaoia, o oiio /enei iaoaiaoe/iee,
i?iaaaeaii naiinoieii. Iioa?i?aoaoeiy ?acoeueoaoia i?iaaaeaia ca o/anoth
ea?iaieea. Iaienaiiy aeena?oaoei? naiinoieia.

Ai?iaaoeiy ?acoeueoaoia aeena?oaoei?. ?acoeueoaoe oa iniiaii iieiaeaiiy
?iaioe aeeeaaeaii ia Iiaeia?iaeiiio neiiicioii ii ea?aeeieiai? (Eiii,
1997), I-io Iaoeiiiaeueiiio eiia?ani iaa?iiaoieiaia i ineoiao?ia (1997),
THaieaeiie eiioa?aioei? «200 ?ieia Naao?iaie Aea/i» (1997), I-io
Iaoeiiiaeueiiio eiia?ani oiciieiaia Oe?a?ie (Aeiiaoeuee, 1998),
eiioa?aioei? Oe?a?inueeiai IAeI eeiii/ii? oa aenia?eiaioaeueii?
iaa?ieiai? oa ineoiao?i? «Iiai iiaeoiaee aei eoiithaaiiy aaeeeoeaieo
noaiia» (Oa?eia, 1999).

Ioaeieaoei?. Iniiaii iieiaeaiiy ?iaioe aiaeia?aaeaii a 6-oe
ioaeieaoeiyo, c ieo 4 noaooi o oaoiaeo aeo?iaeao.

No?oeoo?a oa ianya aeena?oaoei?. Aeena?oaoeieia ?iaioa neeaaea?oueny c
anooio i ?icaeieia: iaeyae eioa?aoo?e, ianya oa iaoiaee aeineiaeaeaiue,
?acoeueoaoe aeanieo aeineiaeaeaiue, caeeth/aiiy oa aeniiaeia, nienea
aeei?enoaieo aeaea?ae (215 iaeiaioaaiue). ?iaioa aeeeaaeaia ia 134-o
noi?iieao iaoeiiieno, iinoeoue 11 oaaeeoeue oa 15 ?enoieia.

IAOA?IAEE OA IAOIAeE AeINEIAeAEAIIss

Aeineiaeaeaiiy aeeiiaii ia 62-o iaeiiieieo uo?ao-naioeyo aaaith 250-270
a, aieii 8-9 iinyoeia o o?iii/iiio aenia?eiaioi.

Aeei?enoaii iaoiaee:

1. Noa?aioaene/ia oaoiiea aenia?eiaioo. Oaa?eiai c aeei?enoaiiyi
noa?aioaene/iiai aia?aoo oa aoeano iiceo uo?a (Ao?ao ss., Iao?aiue I.,
Caoa? I., 1962) iiae oiiiaioaeiaei ia?eicii, ui aaiaeeee
aioo?ioiuei/a?aaii, iiieaiooaaee iio?iiiai aeaeo?iaee o neeyiie
icieyoei? (aeiaiao? aeaeo?iaeo 80-100 iie?iiia) o eaoa?aeueiee
aiiioaeaion, ?aoeeoey?io oi?iaoeith na?aaeiueiai iiceo, i?ici?o
ia?aoeieo (n.accumbens septi), aiiieaii. O eiaii-oii’yio ei?o aeaeo?iaee
iiieaiooaaee aiiaeo?aeueii oaeiai naiiai aeiaiao?o o aeaeyaei nie?aei
(3-4 aeoea), nie?aeue aoea a aeiaiao?i 0,5 ii. Iiaeeoa?aioiee aeaeo?iae
?icoaoiaoaaany a einooei eiaii? iacooe (7-8 aeoeia).

2. Iaoiae naiinoeioethaaiiy iiceo ii Ieaeno oa Iieia?o (Olds J., Milner
P., 1954). Oeae iaoiae aeei?enoaiee ye iiaeaeue iiceoeaii? aiioeieii?
iiaaaeiiee. Nooue iaoiaeo iieyaa? o ae?iaeaiii iino?oiaioaeueiiai iaaeeo
naiinoeioethaaiiy aiioeiiaaiieo cii iiceo, cie?aia eaoa?aeueiiai
aiiioaeaiono. O?eaaeinoue iaeiiai iineeaiiy — 0,5 naeoiae, /anoioa — 50
Aoe, neea — 20-100 ieA. *anoioo ?aaeoei? naiinoeioethaaiiy ?a?no?oaaee c
aeei?enoaiiyi aaoiiaoe/iiai ei/eeueieea ca eiaeii 5 oaeeei i?ioyaii
aiaeeie.

3. Aeaeo?e/io aeoeaiinoue iiceo ?a?no?oaaee iiii- aai aiiiey?ii c
aeei?enoaiiyi 8-ie eaiaeueiiai aeaeo?iaioeaoaeia?aoo EEG-8 oi?ie
«Iaaeiei?» (Aaia?iy) c aaaaeaiiyi iioi?iaoei? a AII o ?aaeueiiio ?aaeeii
/ano.

4. Nenoiei/iee a?oa?iaeueiee oene ?a?no?oaaee ia oainoiaie a?oa?i? uo?ia
aeaeo?iieaoeciia?aoi/iei iaoiaeii c inoeeeia?aoi/iith iiaeeeaoei?th
(Ieiaeen A.A. n niaao., 1982).

5. ?a?no?aoeiy aeaeo?iea?aeeia?aoi/ieo iieacieeia i?iaiaeeeanue ia
aeaeo?iea?aeeia?aoi AE 2O-02 o o?ueio noaiaea?oieo aiaeaaaeaiiyo.
A?aoiaoaaee aiie?ooaei? oa /anoioi? ia?aiao?e, nenoie?/iee oa
?ioaa?aeueiee iieacieee.

6. Aoieiai/iee iaoiae aeeth/aa niinoa?aaeaiiy ca oaa?eiaie ye o
eiio?iei, oae i ia onio aoaiao aenia?eiaioo a oiiaao icieueiaaiiai
ia?aaoaaiiy oa a a?oiiaie iiaaaeiioei. 1 ie 20% aoaiieo uo?e io?eioaaee
i?eioniai, a iioii o aieueiiio aeai?i. Iaaeeo?iiae a aeici 1,4 a/ea
aaiaeeee ia?i?aeueii. Aeey noaoenoe/ii? ia?iaee aeaieo aoa aeei?enoaiee
iaia?aiao?e/iee e?eoa?ie Aieeieniia i O-e?eoa?ie Nouethaeaioa (Aoaea?
A.A., Aaieei A.A., 1973). Aa?ioieaoeith iiieaioaoei? aeaeo?iaeia
caeienithaaee aeaeo?ieioe/iei niiniaii.

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?enoiie 1. Aieea ia?i?aeueiiai aaaaeaiiy iaaeeo?iiaeo ia /anoioo
?aaeoei? naiinoeioethaaiiy eaoa?aeueiiai aiiioaeaiono uo?a ? 33.

A — a ii?ii; A — iiney 7-ai aaaaeaiiy; n — /anoioa iineeaiue
naiinoeioethaaiiy; t-/an (oa.).

Iioiai oa?aeoa?o ciiie oei/aniai? i?aaiicaoei? naiinoeioethaaiiy aoee
aeyaeaii o uo?ia c iiia?aaeiueith 14-oe aeaiiith aeeiaieicaoei?th. O?aaa
neacaoe, ui naia aeeiaieicaoeiy o oei?? a?oie uo?ia o noaii iicaaaeaiiy
/a?aiai? aeice ?icei iiaeaeuoaaea /anoioo naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono c iiaeneeaiiyi iiooeiai? iiaaaeiiee, ui
ni?yiiaaia ia io?eiaiiy iiceoeaii? aiioei? i?e naiinoeioethaaiii. Aeey
iethno?aoei? i?eaaaeaia i?inoi?iai-oei/aniaa oa?aeoa?enoeea i?ioeano
naiinoeioethaaiiy o uo?a ? 9 (?en.2).

Ii/aoeiaa aaaaeaiiy oeei uo?ai iaaeeo?iiaeo aeeeeeaei ?iceo
aecaeueoaoeith /anoioe ?aaeoei? naiinoeioethaaiiy aai ?? niaae. Ianooiii
aaaaeaiiy iaaeeo?iiaeo oeie a?oii oaa?ei aeeeeeaee iinooiiaa aai ?icea
cieaeaiiy /anoioe naiinoeioethaaiiy, i?e oeueiio ia aoei ?? ?iceeaeo
iaaeiei na?aaeiuei?, caieueooaaeeny iaoce aiaeii/eieo iiney na?ie
iineeaiue naiinoeioethaaiue. O 75% uo?ia aiaecia/aeanue iiaia aiaeiiaa
aiae i?eeiaiiy aeeiaieiciaaii? ?aei iaaioue a oiiaao aieiaeii?
iioeaaoei?.

A o?aoie iinoaiiaoei aenia?eiaioo 3-o iiny/ii uo?e uiaeaiii io?eioaaee
i?ioyaii 6-oe iinyoeia aoaiie i oaeei /eiii ia iiiaio aeineiaeaeaiiy
oei/aniai? i?aaiicaoei? /anoioe naiinoeioethaaiiy eaoa?aeueiiai
aiiioaeaiono aoee 9-oe iiny/iiai aieo. ?o iiaaaeiiea aiae?iciyeany
aenieith ?ooiaith aeoeaiinoth, niiioaiith aa?aneaiinoth, ia?iaeei
noi?iaiaeaeoaaeanue noaeiiieie ?aaeoeiyie o aeaeyaei i?aeueieo
aiia?eiiacia, caaaeueiei o?aioiiiyi.

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?enoiie 2. Aieea 14-oe aeaiii? aeeiaieicaoei? ia aeeiaiieo /anoioe
naiinoeioethaaiiy eaoa?aeueiiai aiiioaeaiono uo?a ? 9.

A — a ii?ii; A — iiney 14-oe aeaiii? aeeiaieicaoei? (a noaii
aanoeiaioei?); A — iiney 1-ai aaaaeaiiy; A — iiney 7-ai aaaaeaiiy. n —
/anoioa iineeaiue naiinoeioethaaiiy; t — /an (oa.).

Ia aiaeiiio aiae uo?ia c 14-oe aeaiiith aeeiaieicaoei?th, oey a?oia
uo?ia aaaeei iaaoaaea iino?oiaioaeueiiai iaaeeo naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono. O 88% uo?ia iiae /an naiinoeioethaaiiy
aiaecia/aany ae?aciee ?ooiaee ianiieie, yeee ia aoa ni?yiiaaiee ia
iaaeaeue, iiae /an naiiiiae?aciaiiy oaa?eie ?aaee oa a?ecee iaaeaeue, ca
aeoeaiee ia?iiae naiinoeioethaaiiy aeaaaee aaeeeo /anoioo, iaeiae
noia?ia /anoioa ca 1 aiaeeio aoea iaaenieith. Oei aeaii naiae/aoue i?i
oa, ui o uo?ia c 6-oe iiny/iith aeeiaieicaoei?th no?aaeaeathoue
iianoe/ii i?ioeane, iaaoaaiiy iaaeeo, a aiioeieii ?aaeoei? iathoue
aacoiiaiee oa?aeoa?, aac ei?aeoei? iaiei?oeeaeueieo no?oeoo? iiceo. Aeey
iethno?aoei? i?eaaaeaia i?inoi?iai-oei/aniaa oa?aeoa?enoeea i?ioeano
naiinoeioethaaiiy o uo?a ? 29 (?en.3).

Aaea ia?oi aaaaeaiiy oeie a?oii iaaeeo?iiaeo ia?aaiaeeei i?ioean
naiinoeioethaaiiy ia yeinii iiaee ?iaaiue o 75% uo?ia: aaeeeee ?iceeae
/anoioe ?aaeoei? iaaeiei na?aaeiuei? ca 5 oaeeei ii?aeioaaany c
iiaeaeuaiiyi eieueeinoi iineeaiue naiinoeioethaaiue ca 60 oaeeei, a
ii?iaiyiii c ii/aoeiaith.

?ooiaa aeoeaiinoue iaea oeieani?yiiaaiee oa?aeoa?, uo?e naii ciaoiaeeee
iaaeaeue, aac i?eioniaiai «iaaaaeoaaiiy». Ianooiii aaaaeaiiy
iaaeeo?iiaeo aeeeeeaee oi ae ciiie, ui i ia?oa.

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ffff030000000000

?enoiie 3. Oa?aeoa?enoeea /anoioe ?aaeoei? naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono o uo?ia c 6-oe iiny/iith aeeiaieicaoei?th
iiney aaaaeaiiy 1,4 a/ea iaaeeo?iiaeo uo?a ? 29.

A — iiney 6-oe iiny/ii? aeeiaieicaoei? (a aanoeiaioei?); A — iiney 1-ai
aaaaeaiiy iaaeeo?iiaeo ia oiii aanoeiaioei?; A — iiney 7-ai aaaaeaiiy
iaaeeo?iiaeo ia oiii 6-oe iiny/ii? aeeiaieicaoei?. n — /anoioa iineeaiue
naiinoeioethaaiiy; t — /an (oa.).

*anoioa i oa?aeoa? ?aaeoei? naiinoeioethaaiiy naiae/eee i?i oa, ui oeae
i?aia?ao iie?auo? iai’youe, aeoeao? nenoaio iiceoeaiiai iiaee?iieaiiy c
oeieani?yiiaaiith iiooeiaith iiaaaeiieith. O 25% uo?ia aaaaeaiiy
iaaeeo?iiaeo cieaeoaaei /anoioo naiinoeioethaaiiy. Aeey oeeo uo?ia aoea
oa?aeoa?iith aeniea aiioeieiinoue, iiooeiaa aeoeaiinoue ye aei-, oae i
iiney i?eeiaiiy yoaiieo, a iioii i iiney i?eeiaiiy iaaeeo?iiaeo c
noaeiiieie i?iyaaie a iiaaaeiioei.

Oaei ?icieoei a aoaeoao iaaeeo?iiaeo iiaeia iiynieoe oaeoaie, yei
iniothoue a eioa?aoo?i, i?i ii/aoeiao aaoa?iaaiiinoue aeeiaieueii?
iioeaaoei?, ui coiiaeaia ?iciiiaiioiith aeoeaiinoth
aeeiaieueaeaaiae?iaaiace, ?icieie nooiaiyie o/anoi aoeaoaeueaeaaiaeo a
ciiithaaiii aeoeaiinoi eaoaeace, iaoaaieicio aeioaiiio (Eaioieeiaa I.I.,
Naoaiianeay A.E., 1992; Heap L., Monica R., Pyatot et al., 1995).

Aieea iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo iioaeoieo uo?ia oa
i?e aeeiaieicaoei?. Ia?oa aaaaeaiiy iaaeeo?iiaeo o iioaeoieo uo?ia
aeeeeeaei oaciai ciiie: ai?iaeiaae 30-oe oaeeei aiaecia/aeanue
aeoeaaoeiy aeaeo?iaaiaco, iioii noiaeinoue ii/aoeiai?, a a ae?oaie oaci
/anoioiee niaeo? aiiiioaioeiaeia aoa oa?aeoa?iei aeey ?icaeoeo
aeaeo?ia?aoi/ii? ea?oeie nio. Iiney 5-6-oe aeaiiiai aaaaeaiiy
iaaeeo?iiaeo o aiaenoaaeaiiio ia?iiaei (aei aaaaeaiiy /a?aiai? aeice) o
63% uo?ia ciiithaaiiy AAA aoee iiaeiaieie aei oeo, ui iienaii i?e
?aaeoei? i/ieoaaiiy: ia?ieneciaeueii i?iyae oaoa- i aeueoa-, oeaiee
aaoa-aa?aoai o aiiioaeaioni, aiiieaiii, n.accumbens septi. Nueiia
aaaaeaiiy iaaeeo?iiaeo ia?aaoaeiaoaaei aeaeo?iaaiac o aie oaeaeeeo
?eoiia, i?eaii/oaaei ia?ieneciaeueii i?iyae aiiiioaioeiaeia, a /a?ac
40-50 oaeeei ?icaeaaeanue oa ia?aaaaeaea iiaieueia aeoeaiinoue,
ia?aaaaeii a eiiai/ieo no?oeoo?ao iiceo. sse iieacaa ei?aeyoeieiee
aiaeic, iaeiaioi ca’ycee ii ?eoiai oeeo no?oeoo? aoee c iiaith ei?ith.
Oei niiaaiaeiioaiiy ia ocaiaeaeothoueny c aeaieie i?i ooieoeiiiaeueiee
noai iiceoeaieo aiioeiiaaiieo no?oeoo?, yei iiae aieeaii iaaeeo?iiaeo
i?iyaeyee aenieo aeoeaiinoue.

Iioiai oa?aeoa?o ciiie aeaeo?e/ii? aeoeaiinoi iiceo aoee aecia/aii i?e
7-ie aeaiiiio aaaaeaiii iaaeeo?iiaeo uo?ai c 14-oe aeaiiith
aeeiaieicaoei?th. O?aaa neacaoe, ui aaaaeaiiy aoaiieo i?ioyaii 14-oe
aeiia ciiithaaei aeaeo?e/io aeoeaiinoue: o iiaie ei?i ia?aaaaeaea
aeaneio?iiicaoeiy aiiiioaioeiaeia oa ?o ia?ieneciaeueii i?iyae o
eiiai/ieo no?oeoo?ao iiceo.

Ii/aoeiaa aaaaeaiiy oeei uo?ai iaaeeo?iiaeo ia?aaoaeiaoaaei /anoioiee
niaeo? o aie neio?iiicaoei? aiiiioaioeiaeia, i?eaii/oaaei noaeiiii
?ic?yaee, i?ioa a aiiioaeaioni oa n.accumbens septi ?a?no?oaaeanue
ia?ieneciaeueia aeoeaiinoue. Aaea aei 6-7-ai aeiia aaaaeaiiy
iaaeeo?iiaeo aiaecia/aeanue oaiaeaioeiy aei aiaeiiaeaiiy aeaeo?e/ii?
aeoeaiinoi iiceo, ui aoea aeanoeaa eiaeiie c cacia/aieo no?oeoo? — iiaie
ei?i, aiiieaiio, aiiioaeaiono, n.accumbens septi oa ?aoeeoey?iie
oi?iaoei? na?aaeiueiai iiceo. Aeeiaiiea ia?aaoaeiae /anoioiiai niaeo?o
AAA aoea aeyaeaia aieueo iiaeeaeaii ca aeiiiiiaith ei?aeyoeieiiai
niaeo?o, aoee iieacaii ai?iaiaeii ca’ycee iiae ie?aieie aiiiioaioeiaeaie
o ?icieo no?oeoo?ao iiceo. Oae, i?e aeeiaieueiie aanoeiaioei? (noaii
iicaaaeaiiy aoaiieo) ca’ycee ?icieo iaoa?iia AAA iaee aeiaaeueiee
oa?aeoa?, aiaeia?aaeaee niaith iaaeioeieueiinoue ?iciiiaiioieo
eiioiieaoeie ye ii /anoioai, oae i ii no?oeoo?ai. O oie aea /an iiney
aaaaeaiiy iaaeeo?iiaeo ei?aeyoeieii ca’ycee ?icieo /anoio niaeo?o AAA ii
no?oeoo?ai aiaeia?aaeaee aaeaioeaiinoue aiaeiiaiaei iiceo: ciaiooaaeenue
ei?aeyoeieii ca’ycee, aiaeaoaaany ?iciiaeie iioaa?aoei? iioyao aei
aeeiaieth oa oi?ioaaiiy iiaeo «?ia?oie/eo ainaiaeia» a caeieniaiii
aaeaeaaoiiai ?iaiy ianiaiiy oa niieith, ?icaeoeo iiaaaeiieiaiai oa
aeaeo?ia?aoi/iiai nio.

O uo?ia o?aouei? a?oie c 6-oe iiny/iith aeeiaieicaoei?th a aeaeo?e/iie
aeoeaiinoi iiai? ei?e oa aiiieaiio ia?aaaaeaea iiaieueia aeoeaiinoue, a
?aoeeoey?iie oi?iaoei? na?aaeiueiai iiceo, aiiioaeaioni oa n.accumbens
septi — ia?ieneciaeueia. Iiney i?eeiaiiy aoaiieo o 60% iineethaaeenue
aai aeieeaee noaeiiii i?iyae (aaoa-aa?aoaia, aeaooiaa aeoeaiinoue,
aieaaeaey?iee ?eoi), oiaoi iaea iinoea caieueoaia ii nooiaith
aeeiaieueia aanoeiaioeiy, ui aoea noeioeueiaaia aoaiieii o iaaeinoaoiie
aeici, uia eoiithaaoe ii/aoeiao. Ii o iaeii?? oaa?eie ia aoei aiaeiiae
aiae io?eiaiiy aeiaeaoeiai? aeice aoaiieo.

Aaaaeaiiy iaaeeo?iiaeo noeioethaaee o 75% aeaeo?iaaiac o aie oaeaeeeo
?eoiia o iiaie ei?i, a eiiai/ieo no?oeoo?ao iiceo ia?aaaaeaea
ia?ieneciaeueia aeoeaiinoue. Iniaeeai oey ia?aaoaeiaa aeaeo?iaaiaco
i?inoaaeoaaeanue i?e ei?aeyoeieiiio aiaeici: aoee aeyaeaii oinii
ei?aeyoeieii ca’ycee /anoioiiai niaeo?o iiai? ei?e oa aiiieaiio,
aiiioaeaiono oa n.accumbens septi ci ca’yceii onio aeineiaeaeaieo
no?oeoo? c iiaith ei?ith; o?eaa?ii no?oeoo?e oi?ioaaiiy aeeiaieueii?
iioeaaoei? (aiiioaeaion i n.accumbens septi) ei?aethaaee ii oaoa- i
aaoa-?eoiai, niaeo? /anoio iiai? ei?e ei?aethaaa ii anii ?eoiai o anio
no?oeoo?ao, ui aeaco? ia iniaeeao ?ieue iaiei?oaeno a iaoieiai/iie
iioaa?aoei? iioyao aei aeeiaieth. O 25% uo?ia, aeey yeeo aoei
oa?aeoa?iei cieaeaiiy /anoioe naiinoeioethaaiiy, aiaecia/aeanue aeniea
noaeiiia aeoeaiinoue, ia?aaaaeii o iiaie ei?i oa aiiieaiii. Aaaaeaiiy
iaaeeo?iiaeo ?? eoiithaaee, a aeaeo?e/iie aeoeaiinoi ?iciiiaiioieo
no?oeoo? ia?aaaaeaee /anoioe, ui aoee oa?aeoa?ieie aeey noaio niieieiiai
ianiaiiy (aeueoa-, oaoa-, a iiaie ei?i e aaoa-?eoiia).

Aieea iaaeeo?iiaeo ia na?oeaai-noaeeiio nenoaio. I?e aiaeici aoaeoia
7-ie aeaiiiai aaaaeaiiy iaaeeo?iiaeo eiio?ieueiei uo?ai aoei aeyaeaii
ianooiia: iiaeaeuaiiy aiieiooaee coaoey O, caieueoaiiy aieueoaaeo QRS,
ciaioaiiy aiieiooaee coaoey ?, cieaeoaaeanue /anoioa na?oeaaeo
nei?i/aiue oa aaee/eia iioaa?aeueiiai iieacieea (oaae.1).

Ciiie nenoiei/iiai oeneo aoee ia nooo?ai. ?acii c oeei, i?e aeyaeaiii o
eiio?iei iciae ii?ooaiiy na?oeaaiai ?eoio (iiiioiiieo oeoieiaeo
aeno?anenoie) oa i?iaiaeeiinoi aecia/aany ea?aeeio?iiiee aoaeo
i?aia?aoo: iiiiaethaaany i?aaeeueiee (neioniaee) na?oeaaee ?eoi. Oea ie
iia’yco?ii c aiaeiiaeaiiyi iaoaaiei/ieo i?ioeania iiiea?aeo, ui
i?eaiaeeoue aei ii?iaeicaoei? aaoiiaoecio eeioei i?iaiaeii? nenoaie
na?oey, a iayaiinoue iiceoeaii? ei?aeyoei? iiae iioaa?aeueiei iieacieeii
oa iieacieeaie a?oa?iaeueiiai oeneo naiae/eee i?i ia?aneiiaoe/iee
iai?yiie aieeaia iaaeeo?iiaeo ia na?oeaai ooieoei?.

O uo?ia c 6-oe iiny/iith aeeiaieicaoei?th aaaaeaiiy iaaeeo?iiaeo oaeiae
aeyaeyee iiceoeaii aieeae ia iieacieee ooieoeiiiaeueiiai noaio
na?oeaai-noaeeiii? nenoaie, yea aoea ii?ooaia aeeiaieicaoei?th. Oae, o
oeeo uo?ia o ii/aoeiaiio oiii a ?acoeueoaoi o?eaaei? aeeiaieicaoei?
?a?no?oaaeinue ciiuaiiy naaiaioo SO c iayaiinoth aeeaieiai coaoey S oa
aenieiai iiceoeaiiai coaoey O (oaae.2).

Oaaeeoey 1

Ciiie iniiaieo AEA-iieacieeia o iioaeoieo uo?ia i?e aaaaeaiii
iaaeeo?iiaeo (Im, n=6).

AEA – iieacieeeEiio?ieue*a?ac 7 aei?a aaaaeaiiy iaaeeo?iiaeoO
iA0,0580,0260,0830,026*? iA0,0310,0230,0250,011QRS
iA0,3470,1680,5210,258*NI %54,838,5756,724,09EI
%54,1719,2330,4212,40***NN oae/i337,7416,43323,2617,55*I?eiioee:

1. O ii?iaiyiii c eiio?ieai: * — p<0.1, ** - p<0.05; 2. O, ? - aiieiooaea coaoeia AEA; 3. QRS - aiieiooaea coaoeia eiiieaeno AEA; 4. NI - nenoiei/iee iieaciee; 5. II - iioaa?aeueiee iieaciee; 6. *NN - /anoioa na?oeaaeo nei?i/aiue; 7. IO - iaaeeo?iiae. O aieueoinoi uo?ia aiaecia/aeinue ia?aaaioaaeaiiy ia?aaena?aeue aai ii?ooaiiy aioo?ioiueina?oeaai? i?iaiaeeiinoi, iiaa?niy coaoey O. *anoioa na?oeaaeo nei?i/aiue oa?aeoa?ecoaaeanue aaeeeei ?iceeaeii o aie iiaeaeuaiiy, ui aeacoaaei ia ?icaeoie oaoiea?aei?, iiaeaeuoaaany nenoiei/iee a?oa?iaeueiee oene, ui ei?aethaaei ci caieueoaiiyi iioaa?aeueiiai iieacieea oa yaeyei niaith ia?aaaaeaiiy a ?aaoeyoei? na?oeaai? aeiyeueiinoi neiiaoe/ii? ia?aiai? nenoaie. Oaeei /eiii, aoaiie i?eaii/oaaa ia?aneiiaoe/io oa aeoeaoaaa neiiaoe/io eaiee na?oeaai? ?aaoeyoei? (oaae.2). Naieaeaiia aaaaeaiiy aeeiaieiciaaiei uo?ai iaaeeo?iiaeo aeeeeeaei ciiie a na?oeaai-noaeeiiie aeiyeueiinoi, yei iiaeia aoei ioeiithaaoe a na?aaeiueiio ii a?oii ye a aeayeiio ?iaei ii?iaeicoth/i: ciaiooaaeanue aiieiooaea coaoey O, iiiiaethaaany aei ii?iaeueieo aaee/ei iioaa?aeueiee iieaciee, noaaieicoaaeanue /anoioa na?oeaaeo nei?i/aiue. Iaeiae aieueoaae eiiieaeno QRS oa nenoiei/iee iieaciee nooo?ai ia ciiithaaeenue, ui ie iia'ycoaaee c iiaeiaaeaiiyi aeaeo?e/ii? nenoiee oeoii/eia (Q-O), ia aoei cia/ieo ciii coaoey ?, naaiaio O iiaa?oaany aei icieiii?, aiaeiiaethaaany neioniaee na?oeaaee ?eoi. Oaaeeoey 2 Ciiie iniiaieo AEA-iieacieeia o uo?ia i?e aaaaeaiii iaaeeo?iiaeo iiney 6-oe iiny/ii? aeeiaieicaoei? (Im, n=10). AEA – iieacieeeO?ii?/ia aeeiaie?caoe?y*a?ac 7 aei?a aaaaeaiiy iaaeeo?iiaeoO iA0,0810,0290,0570,011**? iA0,0330,0090,0310,007QRS iA0,4060,2120,3970,148NI %46,848,6648,893,85EI %46,8615,8655,4214,11*NN oae/i342,5318,92328,1818,64* I?eiioee: 1. O ii?iaiyiii c o?. aeeiaieicaoei?th: * - p<0.1, ** - p<0.05; 2. O, ? - aiieiooaea coaoeia AEA; 3. QRS - aiieiooaea coaoeia eiiieaeno AEA; 4. NI - nenoiei/iee iieaciee; 5. II - iioaa?aeueiee iieaciee; 6. *NN - /anoioa na?oeaaeo nei?i/aiue; 7. IO - iaaeeo?iiae. I/aaeaeii, ui oaeee ii?iaeicoth/ee aeiyeueiinoue na?oeaai-noaeeiii? nenoaie aieea iaaeeo?iiaeo aiaeaoaaany /a?ac iiiiaeaiiy ooieoeie aioo?ioiueiia?aaena?aeieo iaoaiiciia, i?ioeano ?aiiey?ecaoei? iiiea?aeo, a oaeiae niiaaiaeiioaiue ?aaoeyoeie ia?aneiiaoe/ii? oa neiiaoe/ii? ia?aiaeo nenoai. Ea?aeeio?iiia aeiy iaaeeo?iiaeo i?e aeeiaieicaoei? iiaeoaa?aeaeoaaeanue i iiiiaeaiiyi iaae ii?iaeueiiai a?oa?iaeueiiai oeneo o uo?ia. AENIIAEE 1. Iaaeeo?iiae (aetheiciaeioeeioi?iiao) i?e ia?i?aeueiiio aaaaeaiii a aeici 1,4 a/ea iaeii?aciai oa ai?iaeiaae 7-ie aeiia ia? noeioethth/ee aieea ia ?aaeoeith naiinoeioethaaiiy eaoa?aeueiiai aiiioaeaiono oa neiaeiai i?ioeane (iai'youe ia iiceoeaio aiioeith), ciiith? aeaeo?iaaiac o aie oaeaeeeo ?eoiia, i?eaii/o? ia?ieneciaeueii i?iyae aeaeo?e/ii? aeoeaiinoi aieiaiiai iiceo o uo?ia. 2. Aei aaaaeaiiy /a?aiai? aeice iaaeeo?iiaeo (5-6-e aeii) o 63% uo?ia aeyaeaii iiaeaeuaiiy ?iaiy aiioeieiiai ?aaaoaaiiy (?aaeoeie naiinoeioethaaiiy, no?aoo i caoenii-iai?iiii? iiaaaeiiee), aeieeiaiiy ia?ieneciaeueii? aeoeaiinoi ia AAA, ui aoee oa?aeoa?ieie aeey noaio iicaaaeaiiy ineoiaeoeaieo niieo/aiue. Aaaaeaiiy ia oeueiio oiii iaaeeo?iiaeo i?eaii/o? ia?ieneciaeueio aeoeaiinoue a aiioeiiaaiieo no?oeoo?ao iiceo c ?icaeoeii o 25% uo?ia aeaeo?ia?aoi/iiai oa iiaaaeiieiaiai nio, noeioeth? iiceiao nenoaio iiceoeaiiai iiaee?iieaiiy, ui ciaoiaeeoue aiaeaeooy a aecaeueoaoei? /anoioe ?aaeoei? naiinoeioethaaiiy. 3. Aeeiaieicaoeiy uo?ia i?ioyaii 14-oe aeiia i?eaiaeeoue aei oi?ioaaiiy aeeiaieueii? aeiiiiaioe o iiaaaeiioei, iicaaaeaiiy /a?aiai? aeice aoaiieo aeoeao? i?ioean naiinoeioethaaiiy, aeeeeea? aeaneio?iiicaoeith aiiiioaioeiaeia o iaiei?oaeni oa noaeiiio aeoeaiinoue a eiiai/ieo aiioeiiaaiieo no?oeoo?ao iiceo c iiioeiaoei?th a aiiioaeaioni oa n.accumbens septi. 4. Aaaaeaiiy iaaeeo?iiaeo uo?ai c 14-oe aeaiiith aeeiaieicaoei?th i?eaii/o? aeeiaieueio iioeaaoeith: aiaeaoaa?oueny ?icea aecaeueoaoeiy /anoioe ?aaeoei? naiinoeioethaaiiy aai ?? cieaeaiiy. I?e aenieiio ?iaii naiinoeioethaaiiy ianooiii eiai aaaaeaiiy cieaeothoue /anoioo aeaii? ?aaeoei?. 87% uo?ia aiaeiiaeyeeny aiae i?eeiaiiy aoaiieo, ui ii?aeio?oueny c neio?iiicaoei?th aiiiioaioeiaeia (oaoa-, aeueoa-), ?icaeaaiiyi oa cieeiaiiyi a aiioeiiaaiieo no?oeoo?ao noaeiiii? aeoeaiinoi oa caiiuaiiyi ?? ia?ieneciaeueiith a n.accumbens septi; aiaeaoaa?oueny oi?ioaaiiy iiaeo "?ia?oie/eo ainaiaeia" a caeieniaiii aiioeieiiai ?aaaoaaiiy oa ianiaiiy c ianooiiei ?icaeoeii aeaeo?ia?aoi/iiai i iiaaaeiieiaiai nio. 5. Aeey uo?ia c 6-oe iiny/iith aeeiaieicaoei?th aoee oa?aeoa?ieie iaoeieani?yiiaaia ?ooiaa aeoeaiinoue, niiioaia aa?aneaiinoue, yei noi?iaiaeaeoaaeenue ia ?ac noaeiiieie ?aaeoeiyie, aaaeeei iaaoaaiiyi iino?oiaioaeueiiai iaaeeo naiinoeioethaaiiy; o 88% aeiaaeeia aoea aeyaeaia iecueea /anoioa naiinoeioethaaiiy, ii?yae c ia?aaaaeaiiyi iiaieueieo eieeaaiue (iie-aeaeueoa, iie-oaoa), ui aoee ?aaoey?ieie iiney i?eeiaiiy aeeiaieth, oiaoi iaei iinoea iineeaiiy aanoeiaioei?, ui noeioethaaeany aaaaeaiiyi iaaeinoaoiuei? aeice aeeiaieth. 6. Ia?oa aaaaeaiiy iaaeeo?iiaeo, ye i ianooiii, uo?ai c 6-oe iiny/iith aeeiaieicaoei?th o noaii iicaaaeaiiy o 75% oaa?ei ia?aaiaeeoue i?ioean naiinoeioethaaiiy ia yeinii iiaee ?iaaiue ooieoeiiioaaiiy: /anoioa naiinoeioethaaiiy iaea aenieee ?iaaiue oa aaeeeee ?iceeae ca 5-oe oaeeeiii iioa?aaee c aeoeaiei iiooeii iaaeaei aac iaaaaeoaaiiy, c ia?aaoaeiaith aeaeo?iaaiaco o aie oaeaeeeo ?eoiia o iaiei?oaeni oa ei?aeyoei?th ii aaoa- a aiioeiiaaiieo no?oeoo?ao (aiiieaiii, aiiioaeaioni, n.accumbens septi), a oaeiae ?aoeeoey?iie oi?iaoei? na?aaeiueiai iiceo. Oea naiae/eoue i?i iniaeeao ?ieue iaiei?oaeno o oi?ioaaiii iioaa?aoei? iioyao aei aeeiaieth oa ?? aeai?/i? noeeueiinoi aei aieeao iaaeeo?iiaeo. O 25% uo?ia iaaeeo?iiae cieaeo? /anoioo naiinoeioethaaiiy, ii?yae c i?eaii/aiiyi noaeiiii? aeoeaiinoi oa ia?aaaaeaiiyi niaeo?o /anoio, ui aoa oa?aeoa?iei aeey noaio niieieiiai ianiaiiy. 7. Iaaeeo?iiae iaaea? ii?iaeicoth/i? aei? ia na?oeaai-noaeeiio nenoaio aeeiaieiciaaieo uo?ia /a?ac iiiiaeaiiy i?ioeano ?aiiey?ecaoei? iiiea?aeo, niiaaiaeiioaiue neiiaoe/ii? oa ia?aneiiaoe/ii? ?aaoeyoeie na?oeaai-noaeeiii? aeiyeueiinoi. Iaaeeo?iiae aeyaey? ea?aeeio?iiio aeith ia oieueee o aeeiaieiciaaieo uo?ia, aea e o iioaeoieo i?e iayaiinoi o ieo ooieoeiiiaeueieo ciii na?oeaai-noaeeiii? nenoaie. 8. I?eaii/aiiy aeeiaieueii? iioeaaoei? o uo?ia iaaeeo?iiaeii caeienith?oueny /a?ac eiai noeioethth/ee aieea ia iiceiai iaoaiicie iiceoeaiiai aiioeieiiai iiaee?iieaiiy oa neiaeiai i?ioeane, aioeaiieaioiaaiio i ianiaoeeoi/io aeoeaoth/i nenoaie iiceo. NIENIE I?AOeUe, UI AIAeIA?AAEATHOUe INIIAII IIEIAEAIIss AeENA?OAOeI? 1. Ai?iaueaaa O.I., Aa?/aiei I.A., Ea?eie I. Aeeyiea iiaiai i?aia?aoa – iaaeeo?iiaea ia yeaeo?e/aneoth aeoeaiinoue iicaa e yiioeeiiaeueiia iiaaaeaiea e?un n yenia?eiaioaeueiui aeeiaieeciii // Oe?. a?niee ineoiiaa?ieia??. – Oa?e?a, 1995. – O.3, aei.2(6). – N.235-236. 2. Ea?eie I. Aeeyiea iaaeeo?iiaea ia yiioeeiiaeueiia iiaaaeaiea e?un n aeeiaieueiie iioeaaoeeae // Aeie. aanoiee OAO. – 1998. - O.2, ? 1. – N.65-69. 3. Ea??i? I. Iae?io?c?ieia?/i? aoaeoe iaae?o?iiaeo o uo??a c aenia?eiaioaeueiei aeeiaie?ciii // I-ee Iaoeiiiaeueiee eiia?an oiciieiaia Oe?a?ie, Aeiiaoeuee, 1998 // O?c?ie. aeo?iae. – 1998. – O.44, ? 3. – N.34-35. 4. Ea?eie I. Aeeyiea iaaeeo?iiaea ia yeaeo?e/aneoth aeoeaiinoue iicaa e?un n aeeiaieueiie iioeaaoeeae // A?niee i?iaeai a?ie. ? iaae.. – Iieoaaa, 1999. - ? 11. - N.118-124. 5. Ea?eie I. Iiuo i?eiaiaiey iiaiai ioa/anoaaiiiai i?aia?aoa iaaeeo?iiaea a ea/aiee yenia?eiaioaeueiiai aeeiaieecia // Na. “Enoi?ey Naao?iaie aea/e. Oniaoe ineoeao?ee, iaa?ieiaee, iae?ioe?o?aee e ia?eieiaee”. – Oa?ueeia, 1996. – O.3. – N.499-501. 6. Ea?eie I., Ai?iaueaaa O.I., Aa?/aiei I.A. Aeeiaieea aeeiaieueiie ea?aeeiieiiaoee iiae aeeyieai iaaeeo?iiaea // Iaaeaeoia?. neiiiceoi “N?aaieoaeueiay yeaeo?iea?aeeieiaey”. – Nueouaea?, 1997. – N.109-110. AIIIOAOeEss Ea?eie Iaaeoa. Aeeyiea iaaeeo?iiaea ia ooieoeeiiaeueiia ninoiyiea iicaiaie nenoaiu iiceoeaiiai iiaee?aieaiey o e?un n aoi?e/ii i?eia?aoaiiuie iioeaaoeeyie (yenia?eiaioaeueiui aeeiaieeciii). - ?oeiienue. Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo iaoe ii niaoeeaeueiinoe 03.00.13 - oeceieiaey /aeiaaea e aeeaioiuo, Oa?ueeianeee ainoaea?noaaiiue oieaa?neoao, Oa?ueeia, 1999. Ia?i?aeueiia aaaaeaiea eiio?ieueiie a?oiia (1) 1,4 a/ea iaaeeo?iiaea iiauoaao /anoioo naiinoeioeyoeee aeiioaeaiona, eciaiyao yeaeo?iaaiac iicaa a noi?iio auno?uo ?eoiia. O e?un n 14-oe aeiaaiie aeeiaieecaoeeae (2) iaaeeo?iiae nieaeaao /anoioo naiinoeioeyoeee, neio?iiece?oao oaoa- e aeueoa-, iiaeaaeyao noaei?iaeioth aeoeaiinoue, ainnoaiaaeeaaao niioiioaiea neiiaoe/aneie e ia?aneiiaoe/aneie ?aaoeyoeee aaiiaeeiaieee. O 87% e?un ioia/ai "ioeac" io i?eiyoey yoaiiea. O 75% e?un n 6-oe iany/iie aeeiaieecaoeeae (3) iaaeeo?iiae aieaa /ai a 2,5 ?aca iiauoaao /anoioo naiinoeioeyoeee, ia?ano?aeaaao yeaeo?iaaiac iicaa a noi?iio auno?uo ?eoiia. O 25% e?un nieaeaao /anoioo aeaiiie ?aaeoeee n oaiaoaieai noaei?iaeiie aeoeaiinoe e i?aiaeaaeaieai aeueoa-?eoia. O 50% e?un ioia/ai "ioeac" io yoaiiea, o 50% - aiaeaaeaioiia ioiioaiea. Ioia/ai ea?aeeio?iiiue yooaeo iaaeeo?iiaea. Iiaeaaeaiea aeeiaieueiie iioeaaoeee iaaeeo?iiaeii iiin?aaeoaony /a?ac iicaiaoth nenoaio iiceoeaiiai iiaee?aieaiey. Eeth/aaua neiaa: iaaeeo?iiae, yoaiie, naiinoeioeyoeey, aeiioaeaion, aeeiaieueiay iioeaaoeey, YYA, YEA, nenoiee/aneia aeaaeaiea. AIIOAOeIss Ea?iii Iaaeoa. Aieea iaaeeo?iiaeo ia ooieoeiiiaeueiee noai iiceiai? nenoaie iiceoeaiiai iiaee?iieaiiy o uo?ia c aoi?eiii iaaooeie iioeaaoeiyie (aenia?eiaioaeueiei aeeiaieiciii). - ?oeiien. Aeena?oaoeiy ia caeiaoooy a/aiiai nooiaiy eaiaeeaeaoa aiieiai/ieo iaoe ca niaoeiaeueiinoth 03.00.13 - oiciieiaiy ethaeeie oa oaa?ei, Oa?eianueeee aea?aeaaiee oiiaa?neoao, Oa?eia, 1999. Ia?i?aeueia aaaaeaiiy eiio?ieueiie a?oii (1) 1,4 a/ea iaaeeo?iiaeo iiaeaeuo? /anoioo naiinoeioethaaiue a?iioaeaiono, ciiith? aeaeo?iaaiac iiceo o aie oaeaeeeo ?eoiia. O uo?ia c 14-oe aeaiiith aeeiaieicaoei?th (2) iaaeeo?iiae cieaeo? /anoioo naiinoeioethaaiue, neio?iiico? oaoa- i aeueoa-, i?eaii/o? noaeiiio aeoeaiinoue, aiaeiiaeth? niiaaiaeiioaiiy neiiaoe/ii? oa ia?aneiiaoe/ii? ?aaoeyoeie aaiiaeeiai?ee. O 87% uo?ia aeyaeaii "aiaeiiao" aiae i?eeiaiiy aoaiieo. O 75% uo?ia c 6-oe iiny/iith aeeiaieicaoei?th (3) iaaeeo?iiae aieueo iiae o 2,5 ?ace iiaeaeuo? /anoioo naiinoeioethaaiiy, ia?aaoaeiao? aeaeo?iaaiac iiceo o aie oaeaeeeo ?eoiia. O 25% uo?ia cieaeo? /anoioo aeaii? ?aaeoei? c i?eaii/aiiyi noaeiiii? aeoeaiinoi oa iayaiinoth aeueoa-?eoio. O 50% uo?ia aecia/aii "aiaeiiao" aiae aoaiieo, o 50% - aiaiaaeaioia aiaeiioaiiy. Aeyaeaii ea?aeeio?iiiee aoaeo iaaeeo?iiaeo. I?eaii/aiiy aeeiaieueii? iioeaaoei? iaaeeo?iiaeii iiina?aaeeiao?oueny /a?ac iiceiao nenoaio iiceoeaiiai iiaee?iieaiiy. Eeth/iai neiaa: iaaeeo?iiae, aoaiie, naiinoeioethaaiiy, aiiioaeaion, aeeiaieueia iioeaaoeiy, AAA, AEA, nenoiee/iee oene. SUMMARY Karimi Padsha. Influence of Medichronal on the functional state of the positive refreshment brain system of rats with motivations attained for the second time (experimental alcoholism). - Manuscript. Dissertation aimed at obtaining the Ph.D. scientific degree in biological sciences under speciality 03.00.13 - physiology of humans and animals, Kharkov State University, Kharkov, 1999. The Dissertation is dedicated to the study of the influence of the new medicine - Medichronal (glucose-oglicinumformiat) on the functional state of the positive refreshment brain system of rats with modelling of motivation attained for the second time - alcohol addiction. The study has been carried out on 62 laboratory rats - non-linear male rats. As a model, reaction of self-stimulation of lateral hypothalamus positive-emotiogenic zones was used. Models of alcoholic motivation was created by means of daily alcoholisation of rats (in the course of 14 days at a time within 6 months' period), which was achieved step by step: at first, by forced intake of ethanol, then in the state of hunger motivation with possibility of free choice of alcoholised or any other (dainty) food. Medichronal in the dose of 1.4 g/kg was given to rats for 7 days perorally in three groups: control (1), following the 14 days' (2) and following the 6 months' alcoholisation (3). Methods used: stereotaxic technique of electrodes implantation into the brain, brain self-stimulation after Oldes and Miller (1954), electro-encephalography, electrocardiography, method of registration of cystolic pressure registration, ethological observation. Verification of electrodes localisation in brain was carried out by transmission of electric stream 1-4 mkA for 10-15 seconds. Statistic data processing was carried out with the use of Wilcockson non-parametric criterion and Student T-criterion. It was demonstrated that Medichronal in the single dose of 1.4 g/kg within the 7 days' period has stimulating influence of the lateral hypothalamus self-stimulation reaction and emotional memory, alters the brain electrogenesis toward fast rhythms, suppresses paroxysmal manifestations of the brain electric activity arising prior to the next introduction of Medichronal. Alcoholisation within 14 days leads to formation of the dominant in behaviour, deprivation of the next dose of ethanol activates the self-stimulation process, leads to un-synchronisation of biopotentials in neocortex and spasmodic activity in limbical emotiogenic brain structures with initiation in hypothalamus and n.accumbens septi. Introduction of Medichronal into these animals suppresses alcoholic motivation: exultation of the frequency of self-stimulation reaction or its reduction is observed. Under high self-stimulation level its subsequent introductions reduce frequency of this reaction. 87% of rats "refused" to take ethanol which combined with synchronisation of teta-rhythm, shattering and disappearing of spasmodic activity in limbical emotiogenic structures and its replacement with paroxysmal activity in n.accumbens septi. Behaviour of rats with 6 months' alcoholisation has deviant character: chaotic movement activity, often accompanied with spontaneous aggressiveness and spasmodic reactions - these rats are hard to train the instrumental skill of self-stimulation; 88% of rats showed low frequency of self-stimulation, slow vibrations prevail on EEG, delta- and teta- peak is regular following the intake of ethanol, i.e. abstinence increases, being stimulated by the introduction of insufficient dose of alcohol. Introduction of Medichronal into this group of rats in the state of ethanol deprivation in 75% of animals leads to the increase in the self-stimulation reaction frequency, accompanied by the active search for the pedal without any "reminding", restructurisation of brain electrogenesis toward fast rhythms in neocortex and beta-correlation in hyppocampus, hypothalamus, n.accumbens septi, reticular formation of middle cerebrum. 25% of rats showed reduction in self-stimulation reaction frequency along with disappearance of spasmodic activity and prevailing EEG frequencies characteristic for the state of calm vigilance. It was detected that Medichronal has normalising effect on the cardio-vascular system of rats with alcoholic motivation through restoration of myocardium depolarisation processes and correlation of sympathetic and parasympathetic regulation of cardio-vascular activity. Suppression of alcoholic motivation in rats is carried out through activation of brain mechanisms of positive emotional refreshment and trace processes, antiepileptogenic and non-specific activising brain systems. Key words: Medichronal, ethanol, self-stimulation reaction, lateral hypothalamus, alcoholic motivation, EEG, cardiogram, systolic pressure. I?aeienaii aei ae?oeo 13.10.99 ?. Cai. N 543 Iaeeaae – 100 i?ei. Oi. ae?oe. a?e. 1,0. Oi?iao 60x84 1/16. OIA “Ciaiiy EOAe”, i. Oa?e?a, aoe. A?oaia 32.

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