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Вплив медихроналу на функціональний стан мозкової системи позитивного підкріплення у щурів з вторинно набутими мотиваціями (експериментальним алкоголі

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OA?E?ANUeEEE AeA?AEAAIEE OI?AA?NEOAO

EA??I? IAAeOA

OAeE 616.89-008.441.13:612.821.8:615.214

AIEEA IAAeEO?IIAEO IA OOIEOe?IIAEUeIEE NOAI IICEIAI? NENOAIE IICEOEAIIAI
I?AeE??IEAIIss O UO??A C AOI?EIII IAAOOEIE IIOEAAOe?ssIE
(AENIA?EIAIOAEUeIEI AEEIAIE?CIII)

03.00.13 – o?c?ieia?y ethaeeie oa oaa?ei

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Oa?e?a – 1999

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia ia eaoaae?? o?c?ieia?? ethaeeie oa oaa?ei Oa?e?anueeiai
aea?aeaaiiai oi?aa?neoaoo I?i?noa?noaa ina?oe Oe?a?ie ? o eaai?aoi???
iae?io?c?ieia?? oa ?ioiieia?? Oe?a?inueeiai IAe? ee?i?/ii? oa
aenia?eiaioaeueii? iaa?ieia?? oa ineo?ao??? IIC Oe?a?ie

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, i?ioani?

Ai?iaeiaa Oaia?a Ieoaee?aia,

caa?aeoaa/ eaai?aoi???th iae?io?c?ieia?? oa ?ioiieia??

Oe?a?inueeiai IAe? ee?i?/ii? oa aenia?eiaioaeueii? iaa?ieia?? oa
ineo?ao??? IIC Oe?a?ie.

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani?

Aeaaeeiaa Aeea ?aai?aia,

caa?aeoaa/ a?aeae?eii ?ai?iaeoeoeaii? aiaeie?eiieia?? Oe?a?inueeiai IAe?
oa?iaeieia?? aiaeie?eiieo caoai?thaaiue IIC Oe?a?ie.

aeieoi? iaaee/ieo iaoe, i?ioani?

Nin?i ?aai Eocueie/,

caa?aeoaa/ eaoaae?ith ia?eieia?? Oa?e?anueei? iaaee/ii? aeaaeai??
i?neyaeeieiiii? ina?oe IIC Oe?a?ie.

I?ia?aeia onoaiiaa: ?inoeooo o?c?ieia?? ?i. aeaae. I.I. Aiaiiieueoey IAI
Oe?a?ie, a?aeae?e o?c?ieia?? aieiaiiai iiceo.

Caoeno a?aeaoaeaoueny “24 “ eenoiiaaea 1999 ?. i 1515 aiaeei?

ia can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee E 64.051.17 i?e
Oa?e?anueeiio aea?aeaaiiio oi?aa?neoao? I?i?noa?noaa ina?oe Oe?a?ie ca
aae?anith: 310077, i. Oa?e?a, i. Naiaiaee, 4, aoaeeoi??y III-15.

C aeena?oaoe??th iiaeia iciaeiieoeny o Oeaio?aeuei?e iaoeia?e
a?ae?ioaoe? Oa?e?anueeiai aea?aeaaiiai oi?aa?neoaoo I?i?noa?noaa ina?oe
Oe?a?ie ca aae?anith: 310077, i. Oa?e?a, i. Naiaiaee, 4.

Aaoi?aoa?ao ?ic?neaiee “21.10. 1999 ?.”

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee

eaiaeeaeao a?ieia?/ieo iaoe,

noa?oee iaoeiaee ni?a?ia?oiee Iaaeaeei A.?.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeueiinoue oaie. O /enei aaaeeeaeo i?iaeai no/anii? iae?iaiieiai? ?
i?iaeaia aoi?eiii iaaooeo iioeaaoeie. ?? aeooaeueiinoue coiiaeaia
iiaeaeuaiiyi ?iciianthaeaeaiinoi a aaaaoueio e?a?iao oaeeo ooo/ieo
iioeaaoeie, ye iioya aei aeeiaieth, ii?oiio, ineoiaeoeaieo niieo/aiue
(Iyoieoeeay E.I., 1994; Aieioei I.A., 1997; Neieoeeee A.I., 1998;
*oi?eeia A.I., 1998; Aa?aociaa N.I., 1998).

Aiaeiii, ui aoi?eiii iaaooi iioeaaoei? oaeiae, ye i i?e?iaeii,
aiieiai/ii, iathoue oie aea iae?iaiieiai/iee noano?ao, oi ae
iiioeithth/i iiceiai iaoaiicie iioaa?aoei? oeeo iioyaia. Iieeaaeath/enue
ia aeiiiioth/i o eioa?aoo?i eiioeaioei? i?i i?e?iaeo ii?iaeueieo
aiieiai/ieo iioeaaoeie (Noaeaeia E.A., 1971; Neiiiia I.A., 1981; Ieieia
A.A., 1997; Stein, 1971), iiaeia c oeo aea iiceoeie ?icaeyaeaoe i
aoi?eiii iaaooi iioyae, yei aiae?iciythoueny ia nai?th nooiinoth, a
iai?e?iaeiith ni?yiiaaiinoth, yea i?ecaiaeeoue aei iaaaeaeaaoii?
iiaaaeiiee.

Caiaeii eioa?aoo?ieo aeaieo, iioya aei aeeiaieth oi?io?oueny ia iniiai
no?oeoo?ii-ooieoeiiiaeueii? i?aaiicaoei? iiceiai? nenoaie iiceoeaiiai
iiaee?iieaiiy, yea a iaaieo oiiaao neoaeeoue aiieiai/iith iao?eoeath
iiaoaeiae ooieoeiiiaeueii? nenoaie iioyao aei aeeiaieth ye oaeoi?o
iiaee?iieaiiy (Ai?iaueaaa O.I., 1977; Aa?/aiei I.A., 1992; Ai?iaueaaa
O.I., Aieioei I.A., Iaeeiaa E.I. e niaao., 1993).

Cia/ia iinoea a iaoaiiciao oi?ioaaiiy aeeiaieueii? iioeaaoei? caeia?
ii?ooaiiy iaoaaieicio aiiaaiieo iiiiaiiiia – eaoaoieaiiiia (Aiioeia
E.I., 1981; Aieei O.I., 1984, 1986), na?ioiiiio (A?iiiaa A.A., 1984;
Oeoeiaa A.I., 1984; Aiyiiae/ A.A., 1984; Naeeiia ?.I. n niaao., 1995;
Aeaeeineay E.A., 1998; Penn ?.A et al., 1978; Mc Bride et al., 1994;
Fahlke N. et al., 1994), a oaeiae iae?iiaioeaeia ( Aiioeia E.I., 1988;
Aoia?ei E.I., 1990; Oaaoe I.I., 1996; Noaeaeia N.E., 1996; Volpicelli
I.R et al., 1991; Sinclair I.D. et al., 1994).

Iniothoue ia?aeiieeai aeaii i?i oa, ui iiceoeaii aiioeiiaaiii no?oeoo?e
iiceo, aea oe?iei i?aaenoaaeaii cacia/aii aeua iae?iiaaeiaoi?e, ia?oeie
?aaaothoue ia i?eeiaiiy aoaiieo oa oiii/ii aaaioe, ui aeeeeeathoue
aaaeaiiy ?o iiaoi?iiai aaeeaaiiy (Ai?iaueaaa O.I., 1976, 1995;
Oaiaiaoaeee I.I. n niaao., 1992; Caa?oao Y.Y., 1993; Iyoieoeeay E.I.,
1994; Carlson, R.H., Lydic, 1976; Hunter B.E. et al., 1978; Dyr W. et
al., 1993), oiaoi aeayei i?aia?aoe, yei ao?o/athoueny o iaoaaieici
iiiiaiiiia oa iae?iiaioeaeia, naii aeeeeeathoue aoi?eiii i?eno?anoi,
oi?ioaaiiy iiaeo aai caieueoaiiy iaaooeo iioeaaoeie.

Iieacaii, ui oi?ioaaiiy iioyao aei aoaiieo caeaaeeoue i aiae eiai
aeaniiai iaoaaieicio a i?aaiicii ethaeeie i oaa?ei. Aoaiie – i?e?iaeiee
iaoaaieio i eiai i?enei?aiee iaiii iiaea noaoe iaeiei c oaeoi?ia
iiaeneeaiiy eiai aeciaaiiiai aaeeaaiiy (Ino?ianeee TH.I., 1981; Aiioeia
E.I., 1981; Iyoieoeeay E.I., 1984). Oei oyaeaiiy eyaee a iniiao iiooeo
iiaeo oiii/ieo niieo/aiue, ni?yiiaaieo nai?th aei?th ia iaoaaieici
aoaiieo. O /enei oaeeo niieo/aiue i?eaa?oa? oaaao iaaeeo?iiae, ui aoa
neioaciaaiee a Oe?a?ii. Oea aaaaoieiiiiiaioiee i?aia?ao –
aetheiciaeioeeioi?iiao, yeee ia? aeanoeainoi ca’ycoaaiiy
aoeaoaeueaeaaiaeo – iaeaieueo oiene/iiai iaoaaieioo aoaiieo.
Iiaeoeaoth/e aoeaoaeueaeaaiae, iaaeeo?iiae i?eaii/o? iaeei c iaoaiiciia
iioeaaoei? aei iueiai (Aoeue I.O. n niaao., 1990, 1998; Neieoeeee A.I.,
1990, 1996). Ii?yae c oeei, oeae i?aia?ao ni?ey? neioaco aiiaaiieo
iiiiaiiiia (Neieoeeee A.I., 1990), ?aaoeth? aiino oa aeoeaiinoue
na?ioiiiio (Ia?eiciaa E.I., 1997; Aiaeei A.O. n niaao., 1997).

?acii c oeei, iiaeeaeaii aeineiaeaeaiiy aieeaia iaaeeo?iiaeo ia
iae?ioiciieiai/ii iaoaiicie ?icaeoeo iioyao aei aeeiaieth a eioa?aoo?i
aiaenooii. O oie aea /an ?o ciaiiy oyaey?oueny iaiaoiaeiei ia oieueee o
oai?aoe/iiio aniaeoi i?iaeaie iaaooeo iioeaaoeie, eiie?aoii, iioyao aei
aeeiaieth, aea i aeey ae?iaeaiiy iioeiaeueieo aa?iaioia ?o eoiithaaiiy
oa i?eaii/aiiy.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. Aeineiaeaeaiiy
i?iaaaeaii a eaai?aoi?i? iae?ioiciieiai? oa iioiieiai? Oe?a?inueeiai
iaoeiai-aeineiaeiiai iinoeoooo eeiii/ii? oa aenia?eiaioaeueii?
iaa?ieiai? oa ineoiao?i? a ?aieao eiiieaenieo IAe?: oeo? 10.37.98
“?ic?iaeoe oaoiieiaith ioeiiee ?eceeo aaeeeoeaii? iiaaaeiiee i
noai?aiiy i?ia?ai ia?aeiii? oa aoi?eiii? i?ioieaeoeee” (? 0198V007888),
Iaoeiiiaeueii? i?ia?aie “Iaaeeei-aiieiai/ii oa nioeiaeueii-ineoieiai/ii
aniaeoe ia?eiiaiie” (oeo? 01.20.02/005E-95), ?icaeie “Aea/eoe
iae?iaiieiai/ii iniaeeainoi no/anieo oi?i ia?eiiaiie oa aeeiaieicio i
?ic?iaeyoe a aenia?eiaioi iiaeoiaee aei ?o eoiithaaiiy”.

Iaoa i caaea/i aeineiaeaeaiiy. Iaoith aeineiaeaeaiiy ? c’ynoaaiiy
iae?ioiciieiai/ieo iaoaiiciia aieeao iaaeeo?iiaeo ia aiioeieio
iiaaaeiieo oa aeaeo?e/io aeoeaiinoue iiceo o iioaeoieo oaa?ei oa c
iiaeaethaaiiyi ooo/ii iaaooiai iioyao aei aeeiaieth.

O aiaeiiaiaeiinoi c iaoith aeineiaeaeaiiy ae?iooaaeeny ianooiii caaea/i:

1) aea/aiiy aieeaia iaaeeo?iiaeo ia ?aaeoeith naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono oa iiaaaeiieo iioaeoieo oaa?ei;

2) aea/aiiy aieeaia iaaeeo?iiaeo ia ?aaeoeith naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono oa iiaaaeiieo uo?ia c 14-oe aeaiiith
aeeiaieicaoei?th;

3) aea/aiiy aieeaia iaaeeo?iiaeo ia ?aaeoeith naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono o uo?ia c 6-oe iiny/iith aeeiaieicaoei?th;

4) aea/aiiy aieeaia iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo
iioaeoieo oaa?ei;

5) aea/aiiy aieeaia iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo uo?ia c
14-oe aeaiiith aeeiaieicaoei?th;

6) aea/aiiy aieeaia iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo uo?ia c
6-oe iiny/iith aeeiaieicaoei?th;

7) aea/aiiy aieeaia iaaeeo?iiaeo ia na?oeaai-noaeeiio nenoaio o
iioaeoieo i aeeiaieiciaaieo uo?ia;

8) i?iaaaeaiiy ii?iaiyeueiiai aiaeico oa ocaaaeueiaiiy ?acoeueoaoia
aieeao iaaeeo?iiaeo ia aeyaeaii iieacieee o iioaeoieo oa aeeiaieiciaaieo
uo?ia.

Iaoeiaa iiaecia i oai?aoe/ia cia/aiiy ?iaioe. Aia?oa iieacaii, ui iiaee
i?aia?ao iaaeeo?iiae (aetheiciaeioeeioi?iiao) ia? noeioethth/ee aieea ia
ooieoeiiiaeueiee noai iiceiai? nenoaie iiceoeaiiai iiaee?iieaiiy (ui
aeyaey?oueny o iiaeaeuaiii /anoioe naiinoeioethaaiiy eaoa?aeueiiai
aiiioaeaiono), ia?aaoaeiao? aeaeo?iaaiac iiceo o aie oaeaeeeo ?eoiia oa
i?eaii/o? ?o ia?ieneciaeueii i?iyae.

C’yniaaii, ui aeeiaieicaoeiy uo?ia i?ioyaii 14-oe aeiia i?eaiaeeoue aei
oi?ioaaiiy aeeiaieueii? aeiiiiaioe o iiaaaeiioei. Iicaaaeaiiy /a?aiai?
aeice aoaiieo aeoeao? o oeeo uo?ia i?ioean naiinoeioethaaiiy iiceo,
aeeeeea? aeaneio?iiicaoeith aiiiioaioeiaeia o iaiei?oaeni oa noaeiiio
aeoeaiinoue o eiiai/ieo aiioeiiaaiieo no?oeoo?ao iiceo c iiioeiaoei?th
?? a aiiioaeaioni oa n.accumbens septi. Aia?oa anoaiiaeaii, ui o oeeo
uo?ia iaaeeo?iiae i?eaii/o? aeeiaieueio iioeaaoeith /a?ac ?iceo
aecaeueoaoeith /anoioe ?aaeoei? naiinoeioethaaiiy, neio?iiicaoeith
aiiiioaioeiaeia iiceo (oaoa- i aeueoa-), ?iciiaeie i cieeiaiiy noaeiiii?
aeoeaiinoi, ?? caiiuaiiy ia?ieneciaeueiith a n.accumbens septi. Iiae
eiai aieeaii aeieeathoue iiai no?oeoo?ii-ooieoeiiiaeueii “?iaioie/?
ainaiaei”, ui caaacia/othoue ?icaeoie aeaeo?ia?aoi/iiai oa
iiaaaeiieiaiai nio c aiaeiiaith uo?ia aiae aeeiaieth.

O uo?ia c o?iii/iith 6-oe iiny/iith aeeiaieicaoei?th iaaeeo?iiae
ia?aaoaeiao? i?ioean naiinoeioethaaiiy c aeoeaiei oeieani?yiiaaiei
iiooeii, iaeeaeaiiyi oiiaii-?aoeaeoi?iiai iaaeeo naiinoeioethaaiiy i
aeaneio?iiicaoei?th aiiiioaioeiaeia o iaiei?oaeni, ei?aeyoei?th ii
aaoa-?eoio a aiioeiiaaiieo no?oeoo?ao, i?eaii/o? noaeiiio aeoeaiinoue.

Aia?oa iieacaii, ui iaaeeo?iiae iaaea? ii?iaeicoth/i? aei? ia
na?oeaai-noaeeiio nenoaio aeeiaieiciaaieo uo?ia c iiiiaeaiiyi
niiaaiaeiioaiue ?? neiiaoe/ii? oa ia?aneiiaoe/ii? ?aaoeyoeie.

Aeieacaii, ui iaaeeo?iiae i?eaii/o? iioya aei aeeiaieth /a?ac
aeoeaoth/ee aieea ia iiceiaee noano?ao iiceoeaiiai aiioeieiiai
iiaee?iieaiiy oa neiaeiai i?ioeane, aioeaiieaioiaaiio i ianiaoeeoi/io
aeoeaoth/i nenoaie iiceo.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoaoia. ?acoeueoaoe ?iaioe iathoue
aaciina?aaeith eeiii/io ni?yiiaaiinoue, ?ice?eaathoue iaoaiicie
oa?aiaaoe/ieo aoaeoia aieeaia iaaeeo?iiaeo, ui ni?ey? eeiiioeenoai o
aeai?i oaeoeee eieoaaiiy oai?eo ia aeeiaieici.

Iniaenoee aianie caeiaoaa/a. ?iaioa c eioa?aoo?ieie aeaea?aeaie,
aenia?eiaioe c aeei?enoaiiyi noa?aioaene/ii? oaoiiee, iaoiaeia
naiinoeioethaaiiy iiceo, ?a?no?aoei? aeaeo?e/ii? aeoeaiinoi iiceo oa
aeaeo?ioiciieiai/ieo iieacieeia, a?oa?iaeueiiai oeneo caeieniaii
naiinoieii. Aiaeic ?o ?acoeueoaoia, o oiio /enei iaoaiaoe/iee,
i?iaaaeaii naiinoieii. Iioa?i?aoaoeiy ?acoeueoaoia i?iaaaeaia ca o/anoth
ea?iaieea. Iaienaiiy aeena?oaoei? naiinoieia.

Ai?iaaoeiy ?acoeueoaoia aeena?oaoei?. ?acoeueoaoe oa iniiaii iieiaeaiiy
?iaioe aeeeaaeaii ia Iiaeia?iaeiiio neiiicioii ii ea?aeeieiai? (Eiii,
1997), I-io Iaoeiiiaeueiiio eiia?ani iaa?iiaoieiaia i ineoiao?ia (1997),
THaieaeiie eiioa?aioei? “200 ?ieia Naao?iaie Aea/i” (1997), I-io
Iaoeiiiaeueiiio eiia?ani oiciieiaia Oe?a?ie (Aeiiaoeuee, 1998),
eiioa?aioei? Oe?a?inueeiai IAeI eeiii/ii? oa aenia?eiaioaeueii?
iaa?ieiai? oa ineoiao?i? “Iiai iiaeoiaee aei eoiithaaiiy aaeeeoeaieo
noaiia” (Oa?eia, 1999).

Ioaeieaoei?. Iniiaii iieiaeaiiy ?iaioe aiaeia?aaeaii a 6-oe
ioaeieaoeiyo, c ieo 4 noaooi o oaoiaeo aeo?iaeao.

No?oeoo?a oa ianya aeena?oaoei?. Aeena?oaoeieia ?iaioa neeaaea?oueny c
anooio i ?icaeieia: iaeyae eioa?aoo?e, ianya oa iaoiaee aeineiaeaeaiue,
?acoeueoaoe aeanieo aeineiaeaeaiue, caeeth/aiiy oa aeniiaeia, nienea
aeei?enoaieo aeaea?ae (215 iaeiaioaaiue). ?iaioa aeeeaaeaia ia 134-o
noi?iieao iaoeiiieno, iinoeoue 11 oaaeeoeue oa 15 ?enoieia.

IAOA?IAEE OA IAOIAeE AeINEIAeAEAIIss

Aeineiaeaeaiiy aeeiiaii ia 62-o iaeiiieieo uo?ao-naioeyo aaaith 250-270
a, aieii 8-9 iinyoeia o o?iii/iiio aenia?eiaioi.

Aeei?enoaii iaoiaee:

1. Noa?aioaene/ia oaoiiea aenia?eiaioo. Oaa?eiai c aeei?enoaiiyi
noa?aioaene/iiai aia?aoo oa aoeano iiceo uo?a (Ao?ao ss., Iao?aiue I.,
Caoa? I., 1962) iiae oiiiaioaeiaei ia?eicii, ui aaiaeeee
aioo?ioiuei/a?aaii, iiieaiooaaee iio?iiiai aeaeo?iaee o neeyiie
icieyoei? (aeiaiao? aeaeo?iaeo 80-100 iie?iiia) o eaoa?aeueiee
aiiioaeaion, ?aoeeoey?io oi?iaoeith na?aaeiueiai iiceo, i?ici?o
ia?aoeieo (n.accumbens septi), aiiieaii. O eiaii-oii’yio ei?o aeaeo?iaee
iiieaiooaaee aiiaeo?aeueii oaeiai naiiai aeiaiao?o o aeaeyaei nie?aei
(3-4 aeoea), nie?aeue aoea a aeiaiao?i 0,5 ii. Iiaeeoa?aioiee aeaeo?iae
?icoaoiaoaaany a einooei eiaii? iacooe (7-8 aeoeia).

2. Iaoiae naiinoeioethaaiiy iiceo ii Ieaeno oa Iieia?o (Olds J., Milner
P., 1954). Oeae iaoiae aeei?enoaiee ye iiaeaeue iiceoeaii? aiioeieii?
iiaaaeiiee. Nooue iaoiaeo iieyaa? o ae?iaeaiii iino?oiaioaeueiiai iaaeeo
naiinoeioethaaiiy aiioeiiaaiieo cii iiceo, cie?aia eaoa?aeueiiai
aiiioaeaiono. O?eaaeinoue iaeiiai iineeaiiy – 0,5 naeoiae, /anoioa – 50
Aoe, neea – 20-100 ieA. *anoioo ?aaeoei? naiinoeioethaaiiy ?a?no?oaaee c
aeei?enoaiiyi aaoiiaoe/iiai ei/eeueieea ca eiaeii 5 oaeeei i?ioyaii
aiaeeie.

3. Aeaeo?e/io aeoeaiinoue iiceo ?a?no?oaaee iiii- aai aiiiey?ii c
aeei?enoaiiyi 8-ie eaiaeueiiai aeaeo?iaioeaoaeia?aoo EEG-8 oi?ie
“Iaaeiei?” (Aaia?iy) c aaaaeaiiyi iioi?iaoei? a AII o ?aaeueiiio ?aaeeii
/ano.

4. Nenoiei/iee a?oa?iaeueiee oene ?a?no?oaaee ia oainoiaie a?oa?i? uo?ia
aeaeo?iieaoeciia?aoi/iei iaoiaeii c inoeeeia?aoi/iith iiaeeeaoei?th
(Ieiaeen A.A. n niaao., 1982).

5. ?a?no?aoeiy aeaeo?iea?aeeia?aoi/ieo iieacieeia i?iaiaeeeanue ia
aeaeo?iea?aeeia?aoi AE 2O-02 o o?ueio noaiaea?oieo aiaeaaaeaiiyo.
A?aoiaoaaee aiie?ooaei? oa /anoioi? ia?aiao?e, nenoie?/iee oa
?ioaa?aeueiee iieacieee.

6. Aoieiai/iee iaoiae aeeth/aa niinoa?aaeaiiy ca oaa?eiaie ye o
eiio?iei, oae i ia onio aoaiao aenia?eiaioo a oiiaao icieueiaaiiai
ia?aaoaaiiy oa a a?oiiaie iiaaaeiioei. 1 ie 20% aoaiieo uo?e io?eioaaee
i?eioniai, a iioii o aieueiiio aeai?i. Iaaeeo?iiae a aeici 1,4 a/ea
aaiaeeee ia?i?aeueii. Aeey noaoenoe/ii? ia?iaee aeaieo aoa aeei?enoaiee
iaia?aiao?e/iee e?eoa?ie Aieeieniia i O-e?eoa?ie Nouethaeaioa (Aoaea?
A.A., Aaieei A.A., 1973). Aa?ioieaoeith iiieaioaoei? aeaeo?iaeia
caeienithaaee aeaeo?ieioe/iei niiniaii.

INIIAII ?ACOEUeOAOE AeINEIAeAEAIIss

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aiioeieio iiaaaeiieo, i?i ui naiae/eoue ai?iaiaeia iiaeaeuaiiy /anoioe
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niieo/aiii c oeieani?yiiaaiith aiioeieii-iioeaiaaiith iiaaaeiieith. Aeey
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naiinoeioethaaiiy o uo?a ? 33 (?en.1). Oei ciiie niinoa?iaaeenue
ai?iaeiaae 5-7-ie aeiia iiney aiaeiiie i?aia?aoo. A a?oiiaie iiaaaeiioei
oei uo?e aoee niieieii, niinoa?iaaeinue caieueoaiiy /ano iiaaaeiieiaiai
nio. O 14% uo?ia aiaecia/aeanue oaiaeaioeiy aei cieaeaiiy /anoioe
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oeieani?yiiaaiiai oa?aeoa?o.

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?enoiie 1. Aieea ia?i?aeueiiai aaaaeaiiy iaaeeo?iiaeo ia /anoioo
?aaeoei? naiinoeioethaaiiy eaoa?aeueiiai aiiioaeaiono uo?a ? 33.

A – a ii?ii; A – iiney 7-ai aaaaeaiiy; n – /anoioa iineeaiue
naiinoeioethaaiiy; t-/an (oa.).

Iioiai oa?aeoa?o ciiie oei/aniai? i?aaiicaoei? naiinoeioethaaiiy aoee
aeyaeaii o uo?ia c iiia?aaeiueith 14-oe aeaiiith aeeiaieicaoei?th. O?aaa
neacaoe, ui naia aeeiaieicaoeiy o oei?? a?oie uo?ia o noaii iicaaaeaiiy
/a?aiai? aeice ?icei iiaeaeuoaaea /anoioo naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono c iiaeneeaiiyi iiooeiai? iiaaaeiiee, ui
ni?yiiaaia ia io?eiaiiy iiceoeaii? aiioei? i?e naiinoeioethaaiii. Aeey
iethno?aoei? i?eaaaeaia i?inoi?iai-oei/aniaa oa?aeoa?enoeea i?ioeano
naiinoeioethaaiiy o uo?a ? 9 (?en.2).

Ii/aoeiaa aaaaeaiiy oeei uo?ai iaaeeo?iiaeo aeeeeeaei ?iceo
aecaeueoaoeith /anoioe ?aaeoei? naiinoeioethaaiiy aai ?? niaae. Ianooiii
aaaaeaiiy iaaeeo?iiaeo oeie a?oii oaa?ei aeeeeeaee iinooiiaa aai ?icea
cieaeaiiy /anoioe naiinoeioethaaiiy, i?e oeueiio ia aoei ?? ?iceeaeo
iaaeiei na?aaeiuei?, caieueooaaeeny iaoce aiaeii/eieo iiney na?ie
iineeaiue naiinoeioethaaiue. O 75% uo?ia aiaecia/aeanue iiaia aiaeiiaa
aiae i?eeiaiiy aeeiaieiciaaii? ?aei iaaioue a oiiaao aieiaeii?
iioeaaoei?.

A o?aoie iinoaiiaoei aenia?eiaioo 3-o iiny/ii uo?e uiaeaiii io?eioaaee
i?ioyaii 6-oe iinyoeia aoaiie i oaeei /eiii ia iiiaio aeineiaeaeaiiy
oei/aniai? i?aaiicaoei? /anoioe naiinoeioethaaiiy eaoa?aeueiiai
aiiioaeaiono aoee 9-oe iiny/iiai aieo. ?o iiaaaeiiea aiae?iciyeany
aenieith ?ooiaith aeoeaiinoth, niiioaiith aa?aneaiinoth, ia?iaeei
noi?iaiaeaeoaaeanue noaeiiieie ?aaeoeiyie o aeaeyaei i?aeueieo
aiia?eiiacia, caaaeueiei o?aioiiiyi.

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?enoiie 2. Aieea 14-oe aeaiii? aeeiaieicaoei? ia aeeiaiieo /anoioe
naiinoeioethaaiiy eaoa?aeueiiai aiiioaeaiono uo?a ? 9.

A – a ii?ii; A – iiney 14-oe aeaiii? aeeiaieicaoei? (a noaii
aanoeiaioei?); A – iiney 1-ai aaaaeaiiy; A – iiney 7-ai aaaaeaiiy. n –
/anoioa iineeaiue naiinoeioethaaiiy; t – /an (oa.).

Ia aiaeiiio aiae uo?ia c 14-oe aeaiiith aeeiaieicaoei?th, oey a?oia
uo?ia aaaeei iaaoaaea iino?oiaioaeueiiai iaaeeo naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono. O 88% uo?ia iiae /an naiinoeioethaaiiy
aiaecia/aany ae?aciee ?ooiaee ianiieie, yeee ia aoa ni?yiiaaiee ia
iaaeaeue, iiae /an naiiiiae?aciaiiy oaa?eie ?aaee oa a?ecee iaaeaeue, ca
aeoeaiee ia?iiae naiinoeioethaaiiy aeaaaee aaeeeo /anoioo, iaeiae
noia?ia /anoioa ca 1 aiaeeio aoea iaaenieith. Oei aeaii naiae/aoue i?i
oa, ui o uo?ia c 6-oe iiny/iith aeeiaieicaoei?th no?aaeaeathoue
iianoe/ii i?ioeane, iaaoaaiiy iaaeeo, a aiioeieii ?aaeoei? iathoue
aacoiiaiee oa?aeoa?, aac ei?aeoei? iaiei?oeeaeueieo no?oeoo? iiceo. Aeey
iethno?aoei? i?eaaaeaia i?inoi?iai-oei/aniaa oa?aeoa?enoeea i?ioeano
naiinoeioethaaiiy o uo?a ? 29 (?en.3).

Aaea ia?oi aaaaeaiiy oeie a?oii iaaeeo?iiaeo ia?aaiaeeei i?ioean
naiinoeioethaaiiy ia yeinii iiaee ?iaaiue o 75% uo?ia: aaeeeee ?iceeae
/anoioe ?aaeoei? iaaeiei na?aaeiuei? ca 5 oaeeei ii?aeioaaany c
iiaeaeuaiiyi eieueeinoi iineeaiue naiinoeioethaaiue ca 60 oaeeei, a
ii?iaiyiii c ii/aoeiaith.

?ooiaa aeoeaiinoue iaea oeieani?yiiaaiee oa?aeoa?, uo?e naii ciaoiaeeee
iaaeaeue, aac i?eioniaiai “iaaaaeoaaiiy”. Ianooiii aaaaeaiiy
iaaeeo?iiaeo aeeeeeaee oi ae ciiie, ui i ia?oa.

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ffff030000000000

?enoiie 3. Oa?aeoa?enoeea /anoioe ?aaeoei? naiinoeioethaaiiy
eaoa?aeueiiai aiiioaeaiono o uo?ia c 6-oe iiny/iith aeeiaieicaoei?th
iiney aaaaeaiiy 1,4 a/ea iaaeeo?iiaeo uo?a ? 29.

A – iiney 6-oe iiny/ii? aeeiaieicaoei? (a aanoeiaioei?); A – iiney 1-ai
aaaaeaiiy iaaeeo?iiaeo ia oiii aanoeiaioei?; A – iiney 7-ai aaaaeaiiy
iaaeeo?iiaeo ia oiii 6-oe iiny/ii? aeeiaieicaoei?. n – /anoioa iineeaiue
naiinoeioethaaiiy; t – /an (oa.).

*anoioa i oa?aeoa? ?aaeoei? naiinoeioethaaiiy naiae/eee i?i oa, ui oeae
i?aia?ao iie?auo? iai’youe, aeoeao? nenoaio iiceoeaiiai iiaee?iieaiiy c
oeieani?yiiaaiith iiooeiaith iiaaaeiieith. O 25% uo?ia aaaaeaiiy
iaaeeo?iiaeo cieaeoaaei /anoioo naiinoeioethaaiiy. Aeey oeeo uo?ia aoea
oa?aeoa?iith aeniea aiioeieiinoue, iiooeiaa aeoeaiinoue ye aei-, oae i
iiney i?eeiaiiy yoaiieo, a iioii i iiney i?eeiaiiy iaaeeo?iiaeo c
noaeiiieie i?iyaaie a iiaaaeiioei.

Oaei ?icieoei a aoaeoao iaaeeo?iiaeo iiaeia iiynieoe oaeoaie, yei
iniothoue a eioa?aoo?i, i?i ii/aoeiao aaoa?iaaiiinoue aeeiaieueii?
iioeaaoei?, ui coiiaeaia ?iciiiaiioiith aeoeaiinoth
aeeiaieueaeaaiae?iaaiace, ?icieie nooiaiyie o/anoi aoeaoaeueaeaaiaeo a
ciiithaaiii aeoeaiinoi eaoaeace, iaoaaieicio aeioaiiio (Eaioieeiaa I.I.,
Naoaiianeay A.E., 1992; Heap L., Monica R., Pyatot et al., 1995).

Aieea iaaeeo?iiaeo ia aeaeo?e/io aeoeaiinoue iiceo iioaeoieo uo?ia oa
i?e aeeiaieicaoei?. Ia?oa aaaaeaiiy iaaeeo?iiaeo o iioaeoieo uo?ia
aeeeeeaei oaciai ciiie: ai?iaeiaae 30-oe oaeeei aiaecia/aeanue
aeoeaaoeiy aeaeo?iaaiaco, iioii noiaeinoue ii/aoeiai?, a a ae?oaie oaci
/anoioiee niaeo? aiiiioaioeiaeia aoa oa?aeoa?iei aeey ?icaeoeo
aeaeo?ia?aoi/ii? ea?oeie nio. Iiney 5-6-oe aeaiiiai aaaaeaiiy
iaaeeo?iiaeo o aiaenoaaeaiiio ia?iiaei (aei aaaaeaiiy /a?aiai? aeice) o
63% uo?ia ciiithaaiiy AAA aoee iiaeiaieie aei oeo, ui iienaii i?e
?aaeoei? i/ieoaaiiy: ia?ieneciaeueii i?iyae oaoa- i aeueoa-, oeaiee
aaoa-aa?aoai o aiiioaeaioni, aiiieaiii, n.accumbens septi. Nueiia
aaaaeaiiy iaaeeo?iiaeo ia?aaoaeiaoaaei aeaeo?iaaiac o aie oaeaeeeo
?eoiia, i?eaii/oaaei ia?ieneciaeueii i?iyae aiiiioaioeiaeia, a /a?ac
40-50 oaeeei ?icaeaaeanue oa ia?aaaaeaea iiaieueia aeoeaiinoue,
ia?aaaaeii a eiiai/ieo no?oeoo?ao iiceo. sse iieacaa ei?aeyoeieiee
aiaeic, iaeiaioi ca’ycee ii ?eoiai oeeo no?oeoo? aoee c iiaith ei?ith.
Oei niiaaiaeiioaiiy ia ocaiaeaeothoueny c aeaieie i?i ooieoeiiiaeueiee
noai iiceoeaieo aiioeiiaaiieo no?oeoo?, yei iiae aieeaii iaaeeo?iiaeo
i?iyaeyee aenieo aeoeaiinoue.

Iioiai oa?aeoa?o ciiie aeaeo?e/ii? aeoeaiinoi iiceo aoee aecia/aii i?e
7-ie aeaiiiio aaaaeaiii iaaeeo?iiaeo uo?ai c 14-oe aeaiiith
aeeiaieicaoei?th. O?aaa neacaoe, ui aaaaeaiiy aoaiieo i?ioyaii 14-oe
aeiia ciiithaaei aeaeo?e/io aeoeaiinoue: o iiaie ei?i ia?aaaaeaea
aeaneio?iiicaoeiy aiiiioaioeiaeia oa ?o ia?ieneciaeueii i?iyae o
eiiai/ieo no?oeoo?ao iiceo.

Ii/aoeiaa aaaaeaiiy oeei uo?ai iaaeeo?iiaeo ia?aaoaeiaoaaei /anoioiee
niaeo? o aie neio?iiicaoei? aiiiioaioeiaeia, i?eaii/oaaei noaeiiii
?ic?yaee, i?ioa a aiiioaeaioni oa n.accumbens septi ?a?no?oaaeanue
ia?ieneciaeueia aeoeaiinoue. Aaea aei 6-7-ai aeiia aaaaeaiiy
iaaeeo?iiaeo aiaecia/aeanue oaiaeaioeiy aei aiaeiiaeaiiy aeaeo?e/ii?
aeoeaiinoi iiceo, ui aoea aeanoeaa eiaeiie c cacia/aieo no?oeoo? – iiaie
ei?i, aiiieaiio, aiiioaeaiono, n.accumbens septi oa ?aoeeoey?iie
oi?iaoei? na?aaeiueiai iiceo. Aeeiaiiea ia?aaoaeiae /anoioiiai niaeo?o
AAA aoea aeyaeaia aieueo iiaeeaeaii ca aeiiiiiaith ei?aeyoeieiiai
niaeo?o, aoee iieacaii ai?iaiaeii ca’ycee iiae ie?aieie aiiiioaioeiaeaie
o ?icieo no?oeoo?ao iiceo. Oae, i?e aeeiaieueiie aanoeiaioei? (noaii
iicaaaeaiiy aoaiieo) ca’ycee ?icieo iaoa?iia AAA iaee aeiaaeueiee
oa?aeoa?, aiaeia?aaeaee niaith iaaeioeieueiinoue ?iciiiaiioieo
eiioiieaoeie ye ii /anoioai, oae i ii no?oeoo?ai. O oie aea /an iiney
aaaaeaiiy iaaeeo?iiaeo ei?aeyoeieii ca’ycee ?icieo /anoio niaeo?o AAA ii
no?oeoo?ai aiaeia?aaeaee aaeaioeaiinoue aiaeiiaiaei iiceo: ciaiooaaeenue
ei?aeyoeieii ca’ycee, aiaeaoaaany ?iciiaeie iioaa?aoei? iioyao aei
aeeiaieth oa oi?ioaaiiy iiaeo “?ia?oie/eo ainaiaeia” a caeieniaiii
aaeaeaaoiiai ?iaiy ianiaiiy oa niieith, ?icaeoeo iiaaaeiieiaiai oa
aeaeo?ia?aoi/iiai nio.

O uo?ia o?aouei? a?oie c 6-oe iiny/iith aeeiaieicaoei?th a aeaeo?e/iie
aeoeaiinoi iiai? ei?e oa aiiieaiio ia?aaaaeaea iiaieueia aeoeaiinoue, a
?aoeeoey?iie oi?iaoei? na?aaeiueiai iiceo, aiiioaeaioni oa n.accumbens
septi – ia?ieneciaeueia. Iiney i?eeiaiiy aoaiieo o 60% iineethaaeenue
aai aeieeaee noaeiiii i?iyae (aaoa-aa?aoaia, aeaooiaa aeoeaiinoue,
aieaaeaey?iee ?eoi), oiaoi iaea iinoea caieueoaia ii nooiaith
aeeiaieueia aanoeiaioeiy, ui aoea noeioeueiaaia aoaiieii o iaaeinoaoiie
aeici, uia eoiithaaoe ii/aoeiao. Ii o iaeii?? oaa?eie ia aoei aiaeiiae
aiae io?eiaiiy aeiaeaoeiai? aeice aoaiieo.

Aaaaeaiiy iaaeeo?iiaeo noeioethaaee o 75% aeaeo?iaaiac o aie oaeaeeeo
?eoiia o iiaie ei?i, a eiiai/ieo no?oeoo?ao iiceo ia?aaaaeaea
ia?ieneciaeueia aeoeaiinoue. Iniaeeai oey ia?aaoaeiaa aeaeo?iaaiaco
i?inoaaeoaaeanue i?e ei?aeyoeieiiio aiaeici: aoee aeyaeaii oinii
ei?aeyoeieii ca’ycee /anoioiiai niaeo?o iiai? ei?e oa aiiieaiio,
aiiioaeaiono oa n.accumbens septi ci ca’yceii onio aeineiaeaeaieo
no?oeoo? c iiaith ei?ith; o?eaa?ii no?oeoo?e oi?ioaaiiy aeeiaieueii?
iioeaaoei? (aiiioaeaion i n.accumbens septi) ei?aethaaee ii oaoa- i
aaoa-?eoiai, niaeo? /anoio iiai? ei?e ei?aethaaa ii anii ?eoiai o anio
no?oeoo?ao, ui aeaco? ia iniaeeao ?ieue iaiei?oaeno a iaoieiai/iie
iioaa?aoei? iioyao aei aeeiaieth. O 25% uo?ia, aeey yeeo aoei
oa?aeoa?iei cieaeaiiy /anoioe naiinoeioethaaiiy, aiaecia/aeanue aeniea
noaeiiia aeoeaiinoue, ia?aaaaeii o iiaie ei?i oa aiiieaiii. Aaaaeaiiy
iaaeeo?iiaeo ?? eoiithaaee, a aeaeo?e/iie aeoeaiinoi ?iciiiaiioieo
no?oeoo? ia?aaaaeaee /anoioe, ui aoee oa?aeoa?ieie aeey noaio niieieiiai
ianiaiiy (aeueoa-, oaoa-, a iiaie ei?i e aaoa-?eoiia).

Aieea iaaeeo?iiaeo ia na?oeaai-noaeeiio nenoaio. I?e aiaeici aoaeoia
7-ie aeaiiiai aaaaeaiiy iaaeeo?iiaeo eiio?ieueiei uo?ai aoei aeyaeaii
ianooiia: iiaeaeuaiiy aiieiooaee coaoey O, caieueoaiiy aieueoaaeo QRS,
ciaioaiiy aiieiooaee coaoey ?, cieaeoaaeanue /anoioa na?oeaaeo
nei?i/aiue oa aaee/eia iioaa?aeueiiai iieacieea (oaae.1).

Ciiie nenoiei/iiai oeneo aoee ia nooo?ai. ?acii c oeei, i?e aeyaeaiii o
eiio?iei iciae ii?ooaiiy na?oeaaiai ?eoio (iiiioiiieo oeoieiaeo
aeno?anenoie) oa i?iaiaeeiinoi aecia/aany ea?aeeio?iiiee aoaeo
i?aia?aoo: iiiiaethaaany i?aaeeueiee (neioniaee) na?oeaaee ?eoi. Oea ie
iia’yco?ii c aiaeiiaeaiiyi iaoaaiei/ieo i?ioeania iiiea?aeo, ui
i?eaiaeeoue aei ii?iaeicaoei? aaoiiaoecio eeioei i?iaiaeii? nenoaie
na?oey, a iayaiinoue iiceoeaii? ei?aeyoei? iiae iioaa?aeueiei iieacieeii
oa iieacieeaie a?oa?iaeueiiai oeneo naiae/eee i?i ia?aneiiaoe/iee
iai?yiie aieeaia iaaeeo?iiaeo ia na?oeaai ooieoei?.

O uo?ia c 6-oe iiny/iith aeeiaieicaoei?th aaaaeaiiy iaaeeo?iiaeo oaeiae
aeyaeyee iiceoeaii aieeae ia iieacieee ooieoeiiiaeueiiai noaio
na?oeaai-noaeeiii? nenoaie, yea aoea ii?ooaia aeeiaieicaoei?th. Oae, o
oeeo uo?ia o ii/aoeiaiio oiii a ?acoeueoaoi o?eaaei? aeeiaieicaoei?
?a?no?oaaeinue ciiuaiiy naaiaioo SO c iayaiinoth aeeaieiai coaoey S oa
aenieiai iiceoeaiiai coaoey O (oaae.2).

Oaaeeoey 1

Ciiie iniiaieo AEA-iieacieeia o iioaeoieo uo?ia i?e aaaaeaiii
iaaeeo?iiaeo (Im, n=6).

AEA – iieacieeeEiio?ieue*a?ac 7 aei?a aaaaeaiiy iaaeeo?iiaeoO
iA0,0580,0260,0830,026*? iA0,0310,0230,0250,011QRS
iA0,3470,1680,5210,258*NI %54,838,5756,724,09EI
%54,1719,2330,4212,40***NN oae/i337,7416,43323,2617,55*I?eiioee:

1. O ii?iaiyiii c eiio?ieai: * – p

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