?inoeooo oeoe?iaeo ao?ye?a

OE?A?INUeEI? AEAAeAI?? AA?A?IEO IAOE

Oa?anaiei Ieaa Ieaenaiae?iae/

OAeE 631.527:631.524.86:635.21

Aeei?enoaiiy a naeaeoe??

ia o?oiooi?ino?ee?noue aeo?aeiiai iaoa??aeo

ea?oiie?, noai?aiiai ia iniia? o?eiaaiaoe/ii

a?aeaeaeaieo aeae?a

niaoe?aeuei?noue 06.01.05 — naeaeoe?y ? ian?iieoeoai

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

EE?A — 1998

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia o a?aeae?e? naeaeoe?? ?inoeoooo ea?oiiey?noaa OAAI a

1993-1996 ??.

Iaoeiaee ea??aiee: aeieoi? n?euenueeiainiiaea?nueeeo iaoe,

noa?oee iaoeiaee
ni?a?ia?oiee,

caneoaeaiee i?aoe?aiee
n?euenueeiai

ainiiaea?noaa Oe?a?ie

Inei/oe Aiae??e Aioiiiae/

?inoeooo ea?oiiey?noaa OAAI,

caa?aeoaa/ a?aeae?eo naeaeoe??

Io?oe?ei? iiiiaioe: aeieoi? n?euenueeiainiiaea?nueeeo
iaoe,

i?ioani? eaoaae?e
naeaeoe??

Eiaeooei Iaee? Naiai?aia

Noinueeee aea?aeaaiee
aa?a?iee oi?aa?neoao

eaiaeeaeao
n?euenueeiainiiaea?nueeeo iaoe,

iaoeiaee ni?a?ia?oiee
eaai?aoi???

a?ioaoiieia?? ea?oiie?

Iiaeaeoaiye Oaoyia
Aiaoie?aia

?inoeooo c?iooaaiiai
caiea?ianoaa OAAI

I?ia?aeia onoaiiaa : A?eioea?e?anueeee aea?aeaaiee aa?a?iee
oi?aa?neoao

I?i?noa?noaa aa?ii?iieneiaiai
eiiieaeno Oe?a?ie

i. A?ea Oea?eaa

Caoeno a?aeaoaeaoueny «11» eenoiiaaea 1998 ?ieo i 10 aiaeei? ia

can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee Ae26.360.01 i?e ?inoeooo?

oeoe?iaeo ao?ye?a OAAI

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? ?inoeoooo oeoe?iaeo
ao?ye?a OAAI.

A?aeaoee ia aaoi?aoa?ao a 2-o i?ei??ieeao, caa??ai? ia/aoeith, i?ineii
iaaeneeaoe ca aae?anith: Oe?a?ia, 252650,,ANI i. Ee?a 650, aoe.
Ee?i?/ia, 25.

?inoeooo oeoe?iaeo ao?ye?a OAAI ia ?i’y a/aiiai nae?aoa?y
Niaoe?ae?ciaaii? a/aii? ?aaee.

Aaoi?aoa?ao ?ic?neaiee » ___» ______________ 1998 ?ieo

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee ______________. Eieii??oeue I.I.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. Naiith iioe?aiith ? oeiaei/eiiith oai?iaith
ea?oiie? a on?o e?a?iao, aea ae?iuothoue oeth eoeueoo?o, ? o?oiooi?ic.

Aeiiii?/ii aoaeoeaiei, aeieia?/ii ? nai?oa?ii-a?a??i?/ii aacia/iei
caniaii ai?ioueae c o?oiooi?icii ? aeei?enoaiiy no?eeeo ni?o?a, iniaeeai
c iie?aaiiei eiio?ieai iciaee. I?ioa oey aeanoea?noue o ni?o?a
aioo??oiueiaeaeiaiai (a iaaeao aeaeo Solanum tuberosum) iioiaeaeaiiy
ae?aaeaia a iacia/i?e i???.

Noai?aiiy iiaeo ni?o?a ea?oiie? c iieueiaith no?ee?noth i?ioe
o?oiooi?ico oa eiiieaenii ?ioeo oe?iieo iciae cae?enithaaeinue ia iniia?
aeei?enoaiiy aeae?a S.demissum, S.semidemissum, S.andigenum oa ?ioeo.

A ?inoeooo? ea?oiiey?noaa OAAI noai?aii iiaee 4-6 — aeaeiaee aeo?aeiee
iaoa??ae, cie?aia ca o/anoth a?aeaeaeaiiai aeaeo S.bulbocastanum c
aenieei ??aiai o?oiooi?ino?eeino?. Caeo/aiiy oeueiai iaoa??aeo a
naeaeoe?eio ?iaioo ? aeooaeueiei iai?yieii aeine?aeaeaiue ii noai?aiith
iiaeo o?oiooi?ino?eeeo ni?o?a c eiiieaenii oe?iieo iciae.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. ?iaioa
aeeiioaaeanue ca ieaiii iaoeiai-aeine?aeieo ?ia?o ?inoeoooo
ea?oiiey?noaa OAAI ? iaea ca’ycie c eiai oaiaoeeith :

— Noai?eoe iia? aenieii?iaeoeoeai? ni?oe ea?oiie? c o?iaeaei?noth
450-550 oe/aa, no?ee? i?ioe oai?ia, i?eaeaoi? aeey ?iaeono??aeueieo
oaoiieia?e ae?iaieoeoaa, ?a?no? ?UA01001319? (1993-1995 ??.);

— Noai?eoe aeo?aeiee iaoa??ae oa aenieia?iaeaei? (ia 5-10% aeua
noaiaea?o?a) ni?oe ea?oiie?, no?ee? aei iniiaieo oai?ia, c aenieeie ?
ye?nieie iieacieeaie, i?eaeaoi? aeey iaoai?ciaaiiai ae?iaieoeoaa, ?a?no?
?01.97U005459 (1996-1997 ??.)

Iaoa ? caaaeaiiy aeine?aeaeaiue. Iaoith aeaieo aeine?aeaeaiue ?
iaa?oiooaaiiy iaoiae?a aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia
iniia? Solanum bulbocastanum oa ?ioeo aeae?a a naeaeoe?? ia
o?oiooi?ino?ee?noue c eiiieaenii ?ioeo ainiiaea?nueei-oe?iieo iciae.

Aeey aeinyaiaiiy aeacaii? iaoe aoee iinoaaeai? caaaeaiiy:

— aea/eoe aeo?aeiee iaoa??ae ca eiiieaenii ainiiaea?nueei-oe?iieo iciae
oa caeo/eoe eiai a naeaeoe?eio ?iaioo;

— aeaoe oa?aeoa?enoeeo iauaaee?a neeaaeieo i?aeaeaeiaeo a?a?eae?a ca
no?ee?noth i?ioe o?oiooi?ico eenoe?a ? aoeuea oa aeiaanoe iiaeeea?noue
?o ii?aeiaiiy a i?ioean? naeaeoe??;

— aeyaeoe caeaaei?noue i?ae ?iaeaenii o?aaeaiiy ? oaeoi?aie iieueiai?
no?eeino? i?ioe o?oiooi?ico ? ia oe?e iniia? aeiaanoe iiaeeea?noue
i?aeaeuaiiy ??aiy no?eeino? i?ioe oai?iae;

— i?iaanoe ii??aiyeueio oa?aeoa?enoeeo ??cieo iaoiae?a i?e aeei?enoaii?
neeaaeieo i?aeaeaeiaeo a?a?eae?a a naeaeoe?? ia no?ee?noue i?ioe
o?oiooi?ico, ooca??ico, o?iaeaei?noue, e?ioiaeeno?noue oa ?io? iciaee;

— noai?eoe oe?iiee naeaeoe?eiee iaoa??ae c aenieith no?ee?noth i?ioe
o?oiooi?ico eenoe?a ? aoeuea oa eiiieaenii ?ioeo ainiiaea?nueei-oe?iieo

iciae, i?eeiyoe o/anoue a naeaeoe?? ea?oiie? ia o?oiooi?ino?ee?noue.

Iaoeiaa iiaecia. Aia?oa aena?oeththoueny ?acoeueoaoe aeine?aeaeaiue ii
aeei?enoaiith aeo?aeiiai iaoa??aeo, noai?aiiai ia iniia? o?eiaaiaoe/ii
a?aeaeaeaiiai aeaeo S.bulbocastanum oa ?ioeo, a naeaeoe?eio i?aeoeeo.

Iaa?oioiaaii iaoiaee aeei?enoaiiy aeacaiiai iaoa??aeo a naeaeoe?? ia
o?oiooi?ino?ee?noue a eiiieaen? c ?ioeie oe?iieie iciaeaie, anoaiiaeaia
aeniea eiai aoaeoeai?noue aeey i?aeoe/ii? naeaeoe??.

I?aeoe/ia oe?ii?noue ?iaioe iieyaa? a iaa?oiooaaii? iaoiae?a
aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia iniia? i?aeaeaeiai?
a?a?eaeecaoe??, caeo/aii? eiai a naeaeoe?eio i?aeoeeo, noai?aii?
ia?niaeoeaiiai naeaeoe?eiiai iaoa??aeo oa ni?o?a ea?oiie?, no?eeeo i?ioe
o?oiooi?ico a eiiieaen? c ?ioeie oe?iieie iciaeaie.

Ia caoeno aeiinyoueny iniiai? iieiaeaiiy:

— i?i aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia iniia?
o?eiaaiaoe/ii a?aeaeaeaieo aeae?a a naeaeoe?? ea?oiie?;

— aeniiaie i?i oniaaeeoaaiiy no?eeino? i?ioe o?oiooi?ico, o?iaeaeiino?,
e?ioiaeenoino? oa ?ioeo iciae neeaaeieie i?aeaeaeiaeie a?a?eaeaie;

-iiaeeaeaia iaoeiaa oyaeaiiy i?i iieueiao no?ee?noue i?ioe o?oiooi-

?ico neeaaeieo i?aeaeaeiaeo a?a?eae?a ia iniia? ?? oaeoi??a;

— aeniiaie i?i caeaaei?noue i?ae iieueiaith no?ee?noth i?ioe o?oiooi?ico
eenoe?a ?c no?ee?noth aoeuea, o?iaeaei?noth, e?ioiaeeno?noth ? ?ioeie
iciaeaie, oa iiaeeea?noue ii?aeiaiiy ?o aenieeo iieaciee?a a i?ioean?
naeaeoe??;

Aeaeea?aoe?y iniaenoiai aianeo. Aeine?aeaeaiiy ii oai? aeena?oaoe??
aaoi?ii aeeiiai? iniaenoi. O/anoue aeena?oaioa iieyaa? a io?eiaii?
?acoeueoao?a aeine?aeaeaiue, ?ic?iaoe? i?ia?aie, i?iaaaeaii? aeine?ae?a,
ocaaaeueiaii? aenia?eiaioaeueieo aeaieo.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iniiai? ?acoeueoaoe aeine?aeaeaiue
aeiiia?aeaeenue ia can?aeaii? iaoiaee/ii? eii?n?? oa a?aeae?eo naeaeoe??
?inoeoooo ea?oiiey?noaa (ui??/ii 1994-1997 ??.); i?aeia?iaei?e ia?aae?
“Naeaeoe?y ea?oiie? ia no?ee?noue i?ioe o?oiooi?ico a eiiieaen? c ?ioeie
iciaeaie (Euea?a — Ia?ioeii, 1994); ia eiioa?aioe?? iieiaeeo a/aieo oa
niaoe?ae?no?a “Iaoeia? iniiae aaaeaiiy ea?oiiey?noaa Oe?a?ie a ?eieiaeo
oiiaao” (Iai?oa?aa, 1996), A/ai?e ?aae? ?inoeoooo ea?oiiey?noaa OAAI,
1998 ?.

Ioae?eaoe?y ?acoeueoao?a aeine?aeaeaiue. Ca iaoa??aeaie aeena?oaoe??
iioae?eiaaii 5 iaoeiaeo i?aoeue ? io?eiaii 2 Aaoi?nueeeo na?aeioeoaa ia
ni?oe ea?oiie?.

?aae?caoe?y ?acoeueoao?a aeine?aeaeaiue. ?acoeueoaoe aeine?aeaeaiue ii
aeei?enoaiith aeo?aeiiai iaoa??aeo, noai?aiiai ia aaaaoiaeaeia?e iniia?
aeei?enoiaothoueny o a?aeae?e? naeaeoe?? ?inoeoooo ea?oiiey?noaa oa eiai
ia?aae?.

Ianya oa no?oeoo?a ?iaioe. Aeena?oaoe?y o aeaeyae? ?oeiieno aeeeaaeaia
ia 114 noi??ieao iaoeiiieniiai oaenoo, ai?uo? 31 oaaeeoeth, 11
iaethie?a,

-3-

4 aeiaeaoe?a. Neeaaea?oueny c anooio, 8 ?icae?e?a, aeniiae?a,
i?iiiceoe?e ae?iaieoeoao ? i?aeoe/i?e naeaeoe??, aeiaeaoe?a. Nienie
aeei?enoaieo aeaea?ae ia?aoiao? 215 iaoeiaeo i?aoeue, a oiio /ene? 33
eaoeinueeei o?eooii.

INIIAIA *ANOEIA

A?IEIA?ss PHYTOPHTHORA INFESTANS, OEIAeI*EII?NOUe,

AEO?AeIEE IAOA??AE OA NAEAEOe?ss IA O?OIOOI?INO?EE?NOUe

(IAEssAe E?OA?AOO?E)

Ia i?aenoaa? iaeyaeo e?oa?aoo?e uiaei nenoaie caoiae?a ai?ioueae i?ioe
Ph.infestans aeieeaa?, ui iaaecae/aeii aaeeea ?ieue iaeaaeeoue
noai?aiith oa canoinoaaiith no?eeeo ni?o?a aeey ?o noai?aiiy iaiao?aeii
aeae?eeoe aeaea?aea no?eeino? oa caeo/aoe a naeaeoe?eio ?iaioo iiaee
aeo?aeiee iaoa??ae.

OIIAE, IAOA??AE OA IAOIAeE I?IAAAeAIIss AeINE?AeAEAIUe

?iaioa aeeiioaaeanue a eaai?ioi??? naeaeoe?? ?inoeoooo ea?oiiey?noaa
OAAI i?ioyaii 1993-1996 ?ie?a. Caiaeueiee ianea aeine?aeieo iie?a
n?aici?ie ciaoiaeeoueny a cii? Iie?nny Oe?a?ie. A?oioe
aea?iiai-i?aecieeno?, noi?uai? oa eaaeinoaeeieno?.

A?ae?ioa?i?/i? oiiae a ?iee i?iaaaeaiiy aeine?aeaeaiue a?ae??ciyeenue
a?ae na?aaei?o aaaaoi??/ieo. Ca ia??iae aaaaoaoe?? ea?oiie? a 1993 ?.
aeiaei 450,2 ii iiaae?a, 1994 ?. — 250,6 ii; 1995 ?. — 302,3 ii i?e
na?aaei?e aaaaoi??/i?e — 274 ii. Na?aaeiueiaeiaiaa oaiia?aoo?a ii
aeaeaaeao a 1993 ?. a oe?eiio aoea ieae/a, a 1995 aeua, ii??aiyii c
na?aaeiueith aaaaoi??/iith. A 1994 ?. a?eueo aeaieee aoa eeiaiue oa
ii/aoie na?iiy.

Aeo?aeiei iaoa??aeii a aeine?aeaeaiiyo aoee c?acee, noai?ai? a
eaai?aoi??? aeo?aeiiai iaoa??aeo ?inoeoooo ea?oiiey?noaa ia iniia?
o?eiaaiaoe/ii a?aeaeaeaiiai aeaeo S.bulbocastanum oa S.acaule,
S.demissum, S.andigenum, S.phureja ? ni?o?a ea?oiie? Aea?ea, A?ooa,
Iie?nueea ?iaeaaa, Ia?ea, Euea?a’yiea oa Iai?oa?anueea a?ea.

Ian?iiy io?eioaaee a?ae no?auoaaiiy oa naiicaieeaiiy neeaaeieo
i?aeaeaeiaeo a?a?eae?a. Iin?a ian?iiy ? ae?iuoaaiiy ?icnaaee i?iaiaeeee
a oaieeoe?. N?yioe? ae?iuoaaee iieueiaith eoeueoo?ith a?ey 100 aaiioei?a
eiaeiiai. Aoeueaia? iieie?iiy ae?iuoaaee ia 3 aoeueaiaeo ae?eyieao c
ieiuath aeeaeaiiy 70o28 ni, a ii??aiyii? c aaouee?anueeeie oi?iaie ?
ni?oaie noaiaea?oaie.

Ia i?ioyc? aaaaoaoe?? i?iaiaeeee oaiieia?/i? niinoa?aaeaiiy oa iae?ee
o?aaeaiiy oai?iaaie. Aecia/aiiy iieueiai? no?eeino? i?ioe o?oiooi?ico
iauaaee?a oa aaouee?anueeeo oi?i i?iaiaeeee, ye ia i?e?iaeiueiio
?ioaeoe?eiiio oii?, oae ? a eaai?aoi?ieo oiiaao oeyoii ooo/iiai
ca?aaeaiiy eenoe?a.

Ioe?ieo aoeuea ca no?ee?noth i?ioe o?oiooi?ico i?iaiaeeee oaeiae a
eaai?aoi?ieo oiiaao oeyoii ?o ?iieoeyoe?? caoaeieeii oai?iae.

-4-

CAEO*AIIss AEO?AeIIAI IAOA??AEO, NOAI?AIIAI IA

INIIA? I?AEAEAeIAI? A?A?EAeECAOe?? A NAEAEOe?EIO ?IAIOO

I?aeaeaeia? a?a?eaee oa?aeoa?ecothoueny i?aeaeuaiei ? aenieei ai?noii
e?ioiaeth, iathoue i?ae eouai a?eueoo e?euee?noue aoeuea, i?ae ni?oe,
i?iyaeythoue aenieo no?ee?noue i?ioe o?oiooi?ico, ye ia i?e?iaeiueiio
?ioaeoe?eiiio oii?, oae ? i?e ooo/iiio ca?aaeaii? (oaae.1). I?ioa aiie
iinooiathoueny ni?oai — noaiaea?oai ca o?iaeaei?noth oa na?aaeiueith
ianith aoeueae, aeaye? c ieo — ? ca niaeiaeie yeinoyie, iathoue a?eueo
aeiaa? noieiie, ii??aithth/e ?c ni?oaie — noaiaea?oaie oa aeiaaee
ia??iae aaaaoaoe?? (oaae.1).

I?e i?aeai?? aaouee?anueeeo ia? aeey a?a?eaeecaoe?? ie ea?oaaeenue
i?eioeeiii caa?aaeaiiy o?oiooi?ino?eeino? oa ?ioeo iciae iauaaeeaie
neeaaeieo i?aeaeaeiaeo a?a?eae?a ? ii?aeiaiiy ?o c oe?iieie iciaeaie
naeaeoe?eieo ni?o?a.

Aeacai? i?aeaeaeia? a?a?eaee naiinoi?ni?. Aiie iathoue oa?oeeueiee
ieeie, ooai?ththoue yaiaee a?ae naiicaieeaiiy, a oaeiae caaeia?eueii
no?auothoueny i?ae niaith oa eoeueoo?ieie ni?oaie ea?oiie?.

O?OIOOI?INO?EE?NOUe IAUAAeE?A

NEEAAeIEO I?AEAEAeIAEO A?A?EAe?A

A aeine?aeaeoaaieo eiia?iaoe?yo o?oiooi?ino?ee?noue iauaaee?a neeaaeieo
i?aeaeaeiaeo a?a?eae?a caeaaeeoue a?ae oaiioeiiaiai i?iyaeaiiy iciaee o
aaouee?anueeeo oi?i.

I?e no?auoaaii? neeaaeieo i?aeaeaeiaeo a?a?eae?a c neaaino?eeeie
aaouee?anueeeie oi?iaie na?aaei?e ?iaeaen o?aaeaiiy iauaaee?a aoa aeuei,
i?ae i?e ?o no?auoaaii? ?c a?eueo no?eeeie oi?iaie aai i?e naiicaieeaii?
aeo?aeieo oi?i (oaae.2).

A on?o eiia?iaoe?yo no?auoaaiue ? i?e naiicaieeaii? neeaaeieo
i?aeaeaeiaeo a?a?eae?a aeuaieyeanue cia/ia e?euee?noue aenieino?eeeo
i?ioe o?oiooi?ico iauaaee?a, a oiio /ene?, ye? ca iciaeith ia?aaeuothoue
aaouee?anuee? oi?ie (oaae.2). Caaaeyee oeueiio iiaeia a?ae?a?aoe
o?aina?an?? c aenieith iieueiaith no?ee?noth i?ioe o?oiooi?ico.

Iieueiaa no?ee?noue i?ioe o?oiooi?ico neeaaea?oueny ?c ?? oaeoi??a:
no?ee?noue aei i?iieeiaiiy a?eaa, iioe?aiiy eiai a oeaieiao eenoea oa
no?ee?noue aei ?iciiiaeaiiy.

?ieoaaoe?eiee ia??iae, no?ee?noue aei iioe?aiiy ? nii?iiioaiiy, ye ?
?iaeaen o?aaeaiiy o?oiooi?icii, caeaaeaoue a?ae oaiioeiiaiai i?iyaeaiiy
oeeo iciae o aaouee?anueeeo oi?i. I?ae iieacieeaie oaeoi??a no?eeino?
i?ioe o?oiooi?ico iauaaee?a ? aaouee?anueeeo oi?i i?aeaeaeiaeo a?a?eae?a
aeyaeaia na?aaeiy ? aeniea iiceoeaia ei?aeyoe?eia caeaaei?noue.

I?ae ?iaeaenii o?aaeaiiy o?oiooi?icii eenoe?a iauaaee?a i?aeaeaeiaeo
a?a?eae?a ? ?ieoaaoe?eiei ia??iaeii aeyaeaia neaaea oa na?aaeiy
iaaaoeaia, a oaeiae na?aaeiy iiceoeaia caeaaei?noue (ca ?iee
aeine?aeaeaiue a?ae -0,173+0,148 aei -0,430+0,126 oa a?ae 0,356+0,172
aei 0,600+0,136). Iaaaoeaia ei?aeyoe?eia Oaaeeoey 1 Oa?aeoa?enoeea
neeaaeieo i?aeaeaeiaeo a?a?eae?a (1994-1996 ??.)

Iiia?e i?aeaeaeiaeo a?a?eae?a oa ni?oe noaiaea?oe O?iaeae aoeuea, a/eou
E?euee?noue aoeuea, oo/eou Na?aae- iy iana aoeueae, a Ai?no e?ioia-eth,
% Niaeia? yeino?, aae (1-5) No?ee?noue i?ioe o?oiooi?ico eenoe?a
?iaeaen aoeuea i?e ia i?e- o?aaeai — ooo/iiio ?iaeiueiio
iy eeno- ca?aaeaii? oii?, aae e?a aae(1-9) (1-9)
Aeiaaeeia noi-eii?a, ni Ia??iae aaaaoaoe??, aei?a

88.1439c6(ac.,d,an.,b.,ph.)* 576 17 33,9 20,7 3,7 9,0 2,7 3,0 28 145

85.568c9 (d., b.) 460 10 46,0 18,2 3,6 9,0 5,0 7,5 24 140-145

83.47c51 (ac.,d.,b.,an.,ph.) 486 14 34,7 14,1 3,5 7,5 6,1 8,6 22 130-140

85.368c17(an.,d.,b.) 553 13 42,5 20,8 3,6 9,0 6,0 7,0 21 130-140

86.579c14 (an.,d.,b.,ph.) 557 12 46,4 23,3 4,0 9,0 5,9 6,5 20 140-145

86.331c138 (ac.,an.,d.,b.,ph.) 490 15 32,7 16,8 3,6 8,0 6,0 4,0 17
130-140

No.Na?oaiie ee?anueeee 868 12 72,3 20,8 4,5 8,0 26,9 5,0 10 110-112

No.Eoaianueea 654 7,6 85,4 16,3 4,0 8,0 12,5 5,0 10 120-124

?, % 2,9-3,6

I??05, a/eou 39-52

* Cei?i/aiith a?aeiia?aeathoue aeaee: an.-S.acaule, d.-S.demissum,
an.-S.andigenum, b.-S.bulbocastanum,

ph.-S.phureja

Oaaeeoey 2 Oa?aeoa?enoeea iauaaee?a i?aeaeaeiaeo a?a?eae?a ea?oiie? ca
iieueiaith no?ee?noth

i?ioe o?oiooi?ico (1995-1996 ??.)

Iiia?e iiioey-oe?e Iioiaeaeaiiy ?iaeaen o?aaeaiiy eenoe?a
aaouee?anueeeo oi?i iauaae- e?a,
na- ca?aaeiy ?aaei? A?aenioie
aaiioei?a c ?iaeaenii o?aaeaiiy eenoe?a aei 5,0 5,1-10,0 10,1-20,0
Eiao?oe??io ei?aeyoe?? i?ae ?iaeaenii o?aaeaiiy eenoe?a aaouee?anueeeo
oi?i ? iauaaee?a, r + Sr 1995 1996

94.58 86.579n14 o Aonoi 5,9 30,6 18,2 17,1 16 23 17 0,487+0,165
0,614+0,129

94.59 -”- o Na?oaiie ee?anueeee 5,9 26,9
16,4 16,3 15 19 25 0,779+0,193 0,805+0,094

94.248 ? 86.579n14 5,9 5,9 5,9 5,6 51 14 25 0,505+0,147 0,608+0,188

94.136 86.331n138o88.1439n6 6,0 2,7 4,3 4,8 58 19 18 0,609+0,093
0,716+0,077

94.47 -”- o EA 78.5053 6,0 20,5 13,2 15,3 24 15 26 0,711+0,125
0,721+0,089

94.241 ? 86.331n138 6,0 6,0 6,0 6,2 49 18 27 0,814+0,142 0,794+0,132

No. Eoaianueea 10,0

caeaaei?noue a ?yae? eiia?iaoe?e i?ae oeeie iciaeaie aeaco?, ui
ciaioaiiy ?ieoaaoe?eiiai ia??iaeo i?ecaiaeeoue aei ca?eueoaiiy ?iaeaena
o?aaeaiiy, a ioaea ? aei cieaeaiiy iieueiai? no?eeino? i?ioe
o?oiooi?ico iauaaee?a. Iaaiaee, iiceoeaia ei?aeyoe?eia caeaaei?noue i?ae
oeeie iciaeaie aeaco? ia oa, ui ?ieoaaoe?eiee ia??iae a ie?aieo
eiia?iaoe?yo ia aieeioa ia iieueiao no?ee?noue i?ioe o?oiooi?ico. A
aeaiiio aeiaaeeo aiia caeaaeaea a?ae ?ioeo oaeoi??a.

I?ae ?iaeaenii ? ae?aiao?ii o?aaeaiiy, a oaeiae i?ae ?iaeaenii ?
nii?iiioaiiyi aeyaeaia aeniea iiceoeaia ei?aeyoe?eia caeaaei?noue. Oea
aeaco?, ui ?iaeaen o?aaeaiiy c?inoa? a?ae ca?eueoaiiy ae?aiao?a ?
nii?iiioaiiy ? iaaiaee, a?ae ciaioaiiy iieaciee?a oeeo iciae
i?aeaeuo?oueny iieueiaa no?ee?noue i?ioe o?oiooi?ico.

I?ae ?ieoaaoe?eiei ia??iaeii ? ae?aiao?ii o?aaeaiiy iauaaee?a aeyaeaia
na?aaeiy ? aeniea iaaaoeaia oa na?aaeiy iiceoeaia ei?aeyoe?eia
caeaaei?noue (ca ?iee aeine?aeaeaiue a?ae -0,472+0,116 aei -0,705+0,104
oa a?ae 0,424+0,153 aei 0,544+0,139). Iaa?iaia caeaaei?noue i?ae oeeie
iciaeaie aeaco?, ui ca?eueoaiiy ?ieoaaoe?eiiai ia??iaeo a a?eueoino?
eiia?iaoe?e i?ecaiaeeoue aei ciaioaiiy ae?aiao?a o?aaeaiiy, a oaeiae ?
aei i?aeaeuaiiy iieueiai? no?eeino? i?ioe o?oiooi?ico.

A iauaaeeao a?ae naiicaieeaiiy i?aeaeaeiaeo a?a?eae?a i?ae aeacaieie
iciaeaie niinoa??aaeanue aiaeia?/ia caeaaei?noue.

I?ae ?ieoaaoe?eiei ia??iaeii ? nii?iiioaiiyi a iauaaeeao aeyaeaia
neaaea, na?aaeiy ? aeniea iaaaoeaia oa neaaea ? na?aaeiy iiceoeaia
ei?aeyoe?eia caeaaei?noue (ca ?iee aeine?aeaeaiue a?ae -0,224+0,247 aei
-0,659+0,111 oa a?ae 0,014+0,100 aei 0,344+0,138).

Iaa?iaia caeaaei?noue i?ae aeacaieie iciaeaie aeaco? ia oa, ui
iiaeiaaeaiee ?ieoaaoe?eiee ia??iae i?ecaiaeeoue aei iaio ?ioaineaiiai
nii?iiioaiiy, aiane?aeie /iai i?aeaeuo?oueny iieueiaa no?ee?noue i?ioe
o?oiooi?ico iauaaee?a. A aeaio eiia?iaoe?yo ia aeyaeaii caeaaeiino? i?ae
?ieoaaoe?eiei ia??iaeii ? nii?iiioaiiyi. Oea na?ae/eoue i?i iacaeaaeia
oniaaeeoaaiiy aeacaieo iciae. Oiio i?e iaaiiio i?aeai?? aaouee?anueeeo
ia? iiaeia ii?aeiaoe aenie? iieacieee oeeo oaeoi??a, ui iiceoeaii aieeia
ia ??aaiue iieueiai? no?eeino? i?ioe o?oiooi?ico.

I?ae ae?aiaoi?ii o?aaeaiiy ? nii?iiioaiiyi o iauaaee?a a?ae
aae?inoaaiiy ? naiicaieeaiiy i?aeaeaeiaeo a?a?eae?a aeyaeaia aeniea
iiceoeaia ei?aeyoe?eia caeaaei?noue (ca ?iee aeine?aeaeaiue a?ae
0,618+0,157 aei 0,971+0,060).

Oaeei /eiii, aenie? iieacieee oaeoi??a iieueiai? no?eeino? i?ioe
o?oiooi?ico i?e iaaiiio i?aeai?? aaouee?anueeeo ia? iiaeooue
ii?aeioaaoenue, ui i?ecaaaea aei i?aeaeuaiiy iieueiai? no?eeino? i?ioe
o?oiooi?ico iauaaee?a.

I?e aeei?enoaii? a naeaeoe?? neeaaeieo i?aeaeaeiaeo a?a?eae?a iiaeia
io?eioaaoe aenieino?ee? i?ioe o?oiooi?ico aoeuea iauaaeee ye iaoiaeii
aae?inoaaiiy, oae ? ?o naiicaieeaiiyi.

Iaie, a ?acoeueoao? aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aii ?
aeae?eaii naeaeoe?eiee iaoa??ae, a yeiio ii?aeiothoueny iieueiaa
no?ee?noue i?ioe o?oiooi?ico eenoe?a ? aenie? iieacieee oaeoi??a
no?eeino? c aenieith no?ee?noth aoeuea.

INIIAI? AA?IIII?*I? ICIAEE IAUAAeE?A

NEEAAeIEO I?AEAEAeIAEO A?A?EAe?A

Aiane?aeie aae?inoaaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a niinoa??aaany
i?ii?aeiee oa?aeoa? oniaaeeoaaiiy o?iaeaeiino?, aaoa?icen aai iacia/ia
aeai?an?y (oaae. 3). ??aaiue i?iyaeaiiy iciaee caeaaeaa a?ae
aaouee?anueeeo oi?i, ui caeo/aeenue a no?auoaaiiy.

Ii??aithth/e aae?inoaaiiy oa naiicaieeaiiy i?aeaeaeiaeo a?a?eae?a
aeyaeaii, ui ye iaeiei oae ? ae?oaei iaoiaeii iiaeia io?eioaaoe
aenieio?iaeaei? iauaaeee (oaae. 3).

Na?aaeiy e?euee?noue aoeuea c eoua iauaaee?a ??aiyeanue na?aaei?e iaio
aaouee?anueeeo oi?i, aoea aeua aai ieae/ith ?o. Aaeeeee ?iciao eieeaaiiy
aeacaii? iciaee aea? iiaeeea?noue a?ae?a?aoe aeey i?aeoe/ii? naeaeoe??
oi?ie c iaea?eueoith e?euee?noth aoeuea c eoua.

Na?aaeiy iana aoeueae iauaaee?a, iaea?aeaieo a?ae aae?inoaaiiy,
aei??aithaaea na?aaei?e iaio aaouee?anueeeo oi?i, aoea ieae/ith ?o aai
aeaui aeuith. Niinoa??aaeinue ?icuaieaiiy iauaaee?a ca na?aaeiueith
ianith aoeueae a?ae 10 aei 150 a?ai. A ae?oaiio iaai?? iiioeyoe?e, aea
ii??aithaaeenue iauaaeee a?ae aae?inoaaiiy ? naiicaieeaiiy neeaaeieo
i?aeaeaeiaeo a?a?eae?a, niinoa??aaeinue aiaeia?/ia ?icuaieaiiy ca
na?aaeiueith ianith aoeueae.

Oaeei /eiii, ye aae?inoaaiiyi neeaaeieo i?aeaeaeiaeo a?a?eae?a, oae ? ?o
naiicaieeaiiyi iiaeia iaea?aeaoe ? a?ae?a?aoe aaiioeie c aaeeeith
na?aaeiueith ianith aoeueae.

I?e aae?inoaaii? neeaaeieo i?aeaeaeiaeo a?a?eae?a niinoa??aa?oueny
i?ii?aeiee ? aeii?iaioiee oa?aeoa? oniaaeeoaaiiy e?ioiaeenoino?. A
?acoeueoao? aae?inoaaiiy oa naiicaieeaiiy neeaaeieo i?aeaeaeiaeo
a?a?eae?a io?eiaii iauaaeee c aenieei ai?noii o aoeueaao e?ioiaeth
(oaae. 4).

Aeoiaey/e c iie?aaiiiai eiio?ieth oa oa?aeoa?o oniaaeeoaaiiy
e?ioiaeenoino?, naeaeoe?th ia aenie? iieacieee oe??? iciaee i?e
aeei?enoaii? i?aeaeaeiaeo a?a?eae?a iio??aii i?iaiaeeoe ii oeio
iane/oth/eo no?auoaaiue.

Oae, i?e no?auoaaii? neeaaeiiai i?aeaeaeiaiai a?a?eaeo 85.568n9 c
iecueeie?ioiaeenoei ni?oii I?eiea, na?aaeiy e?ioiaeeno?noue iauaaee?a ii
?ieai noaiiaeea 16,5% ? 16,6%, a i?e no?auoaaii? c aenieie?ioiaeenoei
ni?oii A?ei?onueea 3 — a?aeiia?aeii 19,5% ? 18,9%. I?e oeueiio
aeueiey?oueny ? a?eueoa e?euee?noue aaiioei?a c e?ioiaeeno?noth aeua
20%.

Oaeei /eiii, aeo?aeiee iaoa??ae , noai?aiee ia iniia? a?aeaeaeaieo
aeae?a, iiaeia aeei?enoiaoaaoe oaeiae a naeaeoe?? ia e?ioiaeeno?noue.

Oaaeeoey 3 Oa?aeoa?enoeea iauaaee?a ca o?iaeaei?noth, iaea?aeaieo a?ae
naiicaieeaiiy

oa aae?inoaaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a

Iiia?e iiioey-oe?e Iioiaeaeaiiy Na?aaeiy o?iaeaei?noue, a/eou
aaouee?anueeeo iauaaee?a, oi?i I+m d t
A?aenioie aaiioei?a c o?iaeaei?noth, a/eou 501-750 751-1000 >1000

1 9 9 5

94.248 ? 86.579c14 550 464+35 0 — 26 20 0

94.58 86.579c14 x Aonoi 609 386+33 -78 1,6 18 0 0

94.59 -”- o Na?oaiie ee?anueeee 655 557+38 +93 1,8 17 26 0

94.241 ? 86.331n138 441 718+46 0 — 15 29 19

94.136 86.331n138 o 88.1439n6 394 439+31 -279 5,0 22 10 0

94.47 -”- o EA 78.5053 633 380+33 -338 6,0 23 0 0

Co. Eoaianueea 602+30

1 9 9 6

94.248 ? 86.579c14 565 623+35 0 — 34 31 16

94.58 86.579c14 x Aonoi 657 617+37 -6 0,1 27 25 15

94.59 -”- o Na?oaiie ee?anueeee 771 780+41 +157 2,9 23 29 28

94.241 ? 86.331n138 540 797+44 0 — 20 35 30

94.136 86.331n138 o 88.1439n6 573 785+43 -12 0,8 27 20 35

94.47 -”- o EA 78.5053 650 679+43 -118 1,9 20 40 15

Co. Eoaianueea 732+35

Oaaeeoey 4 Oa?aeoa?enoeea iauaaee?a iaea?aeaieo a?ae aae?inoaaiiy oa
naiicaieeaiiy

neeaaeieo i?aeaeaeiaeo a?a?eae?a, ca ai?noii e?ioiaeth

Iiia?e iiioey-oe?e Iioiaeaeaiiy Na?aaeiy e?ioiaeeno?noue, %
aaouee?anueeeo iauaaee?a, oi?i I+m d t
A?aenioie aaiioei?a c e?ioiaeeno?noth 15,1-18,0 18,1-20,0 >20

1 9 9 5

94.248 ? 86.579c14 23,9 22,9+0,32 0 — 6 21 69

94.58 86.579c14 x Aonoi 20,9 19,1+0,22 -3,8 9,8 28 37 26

94.59 -”- o Na?oaiie ee?anueeee 23,9 22,7+0,27 -0,2 0,5 8 24 67

94.241 ? 86.331n138 16,9 17,4+0,25 0 — 36 23 19

94.136 86.331n138 o 88.1439n6 18,7 17,6+0,24 +0,2 0,6 33 29 22

94.47 -”- o EA 78.5053 14,9 15,4+0,35 -2,0 4,6 41 18 14

Co. Eoaianueea 16,0+0,22

1 9 9 6

94.248 ? 86.579c14 22,8 21,9+0,33 0 — 12 32 52

94.58 86.579c14 x Aonoi 18,1 19,0+0,32 -2,0 4,3 30 39 24

94.59 -”- o Na?oaiie ee?anueeee 20,9 20,5+0,29 -0,5 1,1 12 35 45

94.241 ? 86.331n138 16,7 16,8+0,26 0 — 48 9 8

94.136 86.331n138 o 88.1439n6 18,8 18,1+0,28 +1,3 3,4 31 19 18

94.47 -”- o EA 78.5053 15,8 15,5+0,27 -1,3 3,5 34 7 5

Co. Eoaianueea 14,2+0,25

NOAI?AIIss O?OIOOI?INO?EEIAI NAEAEOe?EIIAI

IAOA??AEO A EIIIEAEN? C ?IOEIE ICIAEAIE

A iauaaeeao a?ae aae?inoaaiiy ? naiicaieeaiiiy neeaaeieo i?aeaeaeiaeo
a?a?eae?a aeyaeaia neaaea iaaaoeaia ? neaaea iiceoeaia ei?aeyoe?eia
caeaaei?noue i?ae ?iaeaenii o?aaeaiiy eenoe?a ? o?iaeaei?noth (ca ?iee
aeine?aeaeaiue a?ae -0,116+0,146 aei -0,289+0,228 oa a?ae 0,130+0,129
aei 0,342+0,129). A?aenooi?noue caeaaeiino? i?ae aeacaieie iciaeaie
na?ae/eoue i?i iacaeaaeia ?o oniaaeeoaaiiy. Oiio, i?e iaaiiio i?aeai??
aaouee?anueeeo ia? ? iaiao?aeiiio ianyc? ?ia?o iiaeia ii?aeiaoe aenie?
iieacieee iieueiai? no?eeino? i?ioe o?oiooi?ico ? o?iaeaeiino?.

A?aenooi?noue ei?aeyoe?eii? caeaaeiino? i?ae ?iaeaenii o?aaeaiiy
o?oiooi-?icii eenoe?a ? e?ioiaeeno?noth iauaaee?a (ca ?iee
aeine?aeaeaiue a?ae 0,018+ 0,101 aei -0,104+0,126 oa a?ae 0,136+0,150
aei 0,317+0,212) na?ae/eoue i?i iacaeaaeia oniaaeeoaaiiy aeacaieo iciae
? iiaeeea?noue ?o ii?aeiaiiy a i?ioean? naeaeoe??.

I?aeae?ath/e aeey a?a?eaeecaoe?? aaouee?anuee? oi?ie c aenieei
oaiioeiiaei i?iyaeaiiyi aeniei? iieueiai? no?eeino? i?ioe o?oiooi?ico ?
aenieei ai?noii e?ioiaeth, iiaeia iaea?aeaoe naeaeoe?eiee iaoa??ae, yeee
ae ii?aeioaaa a nia? aenie? iieacieee oeeo iciae.

A?aenooi?noue ei?aeyoe?eii? caeaaeiino? i?ae e?ioiaeeno?noth ?
o?iaeaei?noth a ?yae? eiia?iaoe?e (aeaco?, ui oe? iciaee
oniaaeeiaothoueny iacaeaaeii ? i?e oe?eani?yiiaai?e ?iaio? ?o aenie?
iieacieee iiaeia ii?aeiaoe.

I?ae no?ee?noth iauaaee?a i?ioe ooca??ico ? ?iaeaenii o?aaeaiiy
o?oiooi?icii eenoe?a, aeyaeaia neaaa iaaaoeaia ? neaaa iiceoeaia
ei?aeyoe?eia caeaaei?noue (ca ?iee aeine?aeaeaiue a?ae 0,093+0,145 aei
-0,315+0,165 oa a?ae 0,136+0,194 aei 0,232+0,203).
A?aenooi?noue caeaaeiino? i?ae aeacaieie iciaeaie na?ae/eoue i?i
iacaeaaeia oniaaeeoaaiiy oeeo iciae ? iiaeeea?noue ?o ii?aeiaiiy a
i?ioean? naeaeoe??.

Iiae?aia caeaaei?noue aeyaeaia i?ae no?ee?noth aoeuea i?ioe ooca??ico ?
no?ee?noth i?ioe o?oiooi?ico aoeuea.

Iaie noai?aii iaoa??ae, yeee ii?aeio? aenieo no?ee?noue i?ioe
o?oiooi?ico eenoe?a ? aoeuea c i?aeaeuaiith no?ee?noth i?ioe ooca??ico.

NOAI?AIIss Oe?IIIAI NAEAEOe?EIIAI IAOA??AEO

Aiane?aeie aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia iniia?
a?aeaeaeaieo aeae?a, iaie aeae?eaii eiia?iaoe?? no?auoaaiiy ?
naeaeoe?eiee iaoa??ae, yeee oa?aeoa?eco?oueny eiiieaenii oe?iieo iciae
(oaae. 5).

Iaie, a ni?aaaoi?noa?, noai?aii 4 iiaeo ni?oe ea?oiie?: ?aii?e Iia?iue,
na?aaeiueinoeae? ssa?? oa Neia’yiea, na?aaeiuei?aii?e Eoiaaa. Ia?o? aeaa
i?ioiaeyoue aea?aeaaia ni?oiaei?iaoaaiiy, inoaii? aeaa — caianaii aei
?a?no?o ni?o?a ?ineei Oe?a?ie a?aeiia?aeii c 1999 ? 1998 ?ie?a.

Oaaeeoey 5 A?a?eaee ea?oiie? c aenieith no?ee?noth i?ioe o?oiooi?ico
(1995-1996 ??.)

Iiia?e a?a?eae?a Iioiaeaeaiiy A?oia noeaeino? Oaeoi?e iieueiai?
no?eeino? ?ieoaaoe?eiee ae?aiao? nii?iiioai- ia??iae, aei?a
o?aaeaiiy, ii iy, aae (1-3) ?iaeaen o?aaeaiiy eenoe?a No?ee?noue i?ioe
o?oiooi?ico aoeuea, aae (1-9)

No. Na?oaiie ee?anueeee na?aaeiuei?aiiy 3,0 32,0 3,0 40,0 5,0

94.58-21 86.579n14 o Aonoi -”- 6,0 5,0 0 1,0 8,0

94.35-53 85.568n9 o EA 78.5053 -”- 5,5 10,0 0,5 2,5 9,0

No. Eoaianueea na?aaeiueinoeaea 5,0 15,0 1,0 10,0 5,0

94.58-11 86.579n14 o Aonoi -”- 5,5 10,0 0 2,5 9,0

94.248-18 ? 86.579n14 -”- 7,0 0 0 0 9,0

94.248-27 -”- -”- 6,0 5,0 0,5 1,0 9,0

No. Ca?aai na?aaeiueii?ciy 5,0 20,0 1,5 20,0 8,0

94.160-21 85.568n9 o A?ei?onueea 3 -”- 5,5 5,0 0,5 1,2 9,0

94.160-44 -”- -”- 7,0 0 0 0 9,0

94.248-9 ? 86.579n14 -”- 7,0 0 0 0 9,0

94.248-42 -”- -”- 7,0 0 0 0 9,0

AEIIII?*IA AOAEOEAI?NOUe AE?IUOAAIIss

IIAEO NI?O?A EA?OIIE?

Ainiiaea?nueeo aoaeoeai?noue anoaiiaethaaee oeyoii ii??aiyiiy
?acoeueoao?a ae?iaieoeoaa i?iaeoeoe?? iiaeo ni?o?a c ni?oaie —
noaiaea?oaie. Oae, i?e aeei?enoaii? iiaeo ni?o?a Eoiaaa, Ceia’yiea ?
ssa?? io?eiaii i?eaaaeo o?iaeath a ii??aiyii? ?c ni?oaie noaiaea?oaie
Iaanueea ? Aao/einueea a?ae 37 oe/aa aei 167 oe/aa.

Iaea?eueo iiaii aeiiii?/iee aoaeo iiaeia aecia/eoe aa?o?nieie
iieacieeaie. Oae, ia?aaeuaiiy /enoiai i?eaooeo a?ae ae?iuoaaiiy iiaiai
na?aaeiuei?aiiueiai ni?oo Eoiaaa iaae noaiaea?oii Iaanueea noaiiaeoue
967 a?eaaiue. A a?oi? na?aaeiueinoeaeeo ia?aaeuaiiy /enoiai i?eaooeo
noaiiaeoue a?ae 4322 a?eai? o ni?oo ssa?? aei 9055 a?eaaiue ni?oo
Neia’yiea iaae noaiaea?oii Aao/einueea.

AENIIAEE

1. Iaa?oioiaaii iaoiaee aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia
iniia? o?eiaaiaoe/ii a?aeaeaeaieo aeae?a (S.bulbocastanum, S.demissum,
S.acaule, S.andigenum, S.phureja) a naeaeoe?? ea?oiie? ia
o?oiooi?ino?ee?noue oa ?io? iciaee.

2. Neeaaei? i?aeaeaeia? a?a?eaee ii??aiyii c ni?oaie oa?aeoa?ecothoueny:
aaaaoiaoeueaia?noth 10-17 oooe/eou, i?aeaeuaiei ai?noii e?ioiaeth aei
23,3%, no?ee?noth i?ioe o?oiooi?ico eenoe?a aei 9,0 aae?a, aoeuea aei
8,6 aaea.

3. Aeei?enoaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a a naeaeoe?? ia iieueiao
no?ee?noue i?ioe o?oiooi?ico (eenoe?a ? aoeuea) oa ?io? iciaee
iaiao?aeii i?iaiaeeoe oeyoii ?o oe?eani?yiiaaiiai aae?inoaaiiy,
naiicaieeaiiy oa no?auoaaiiy i?ae niaith.

4. Anoaiiaeaiee aenieee iiceoeaiee eiao?oe??io ei?aeyoe?? i?ae
iieueiaith no?ee?noth i?ioe o?oiooi?ico eenoe?a neeaaeieo i?aeaeaeiaeo
a?a?eae?a oa ?o iauaaee?a (0,487…0,814), ui na?ae/eoue i?i
caaeainiaai?noue aaiiio aeo?aeieo a no?auoaaii? oi?i ca aoaeoeaieie
aaiaie eiio?ieth iciaee. Aeuaieaiiy iauaaee?a c no?ee?noth aeuith, i?ae
aaouee?anueeeo oi?i noaiiaeoue 59,8%.

5. Aeiaaaeaia aeniea caeaaei?noue i?iyao neeaaeiaeo o?oiooi?ino?eeino?
neeaaeieo aaaaoiaeaeiaeo a?a?eae?a oa ?o iauaaee?a: ca ?ieoaaoe?eiei
ia??iaeii (r=0,582…0,835), ca no?ee?noth i?ioe iioe?aiiy a?eaa
(r=0,526…0,786), ca no?ee?noth i?ioe nii?iiioaiiy (r=0,579…0,776).
Aeieacaii, ui a i?ioean? aae?inoaaiiy iaoa??aeo, yeee aea/aee, iaea?eueo
ao?a/a?oueny no?ee?noue i?ioe i?iieeiaiiy a?eaa ? caeeoathoueny aenieeie
no?ee?noue i?ioe iioe?aiiy oa ?iciiiaeaiiy eiai.

6. Aeyaeaia aeniea ei?aeyoe?eia caeaaei?noue no?eeino? i?ioe o?oiooi?ico
aoeuea aaouee?anueeeo oi?i oa iauaaee?a (0,568…0,778). Aeuaieaiiy
aaiioei?a c no?ee?noth aeuith i?ae o caeo/aieo a no?auoaaiiy oi?i
neeaaea?

38-80%, ui na?ae/eoue i?i aenieo /anoioo o?aina?an??.

7. Aeyaeaia neaaea ? na?aaeiy iaaaoeaia oa iiceoeaia ei?aeyoe?eia
caeaaei?noue i?ae ?iaeaenii o?aaeaiiy o?oiooi?icii eenoe?a ? no?ee?noth
aoeuea iauaaee?a i?aeaeaeiaeo a?a?eae?a. A?aenooi?noue ei?aeyoe?eii?
caeaaei?no? i?ae aeacaieie iciaeaie na?ae/eoue i?i iacaeaaeia ?o
oniaaeeoaaiiy ? iiaeeea?noue ii?aeiaiiy aenieeo iieaciee?a oeeo iciae
i?e iaaiiio i?aeai?? aaouee?anueeeo ia?. Ia ii?aeiaiiy a iauaaee?a
o?oiooi?ino?eeino? eenoe?a ? aoeuea aieeaathoue aaouee?anuee? oi?ie, ui
caeo/athoueny a a?a?eaeecaoe?th. A eiia?iaoe?? 85.568n9 o I?eiea ii
?ieao r=0,492…0,531, a a 86.579n14 o Aonoi r=0,355…0,376. Aeae?eai?
iauaaeee ??cieo a?oi noeaeino?, o yeeo ii?aeio?oueny aeniea
o?oiooi?ino?ee?noue eenoe?a ? aoeuea.

8. Oe?iea aaiaoe/ia iniiaa neeaaeieo i?aeaeaeiaeo a?a?eae?a aea?
iiaeeea?noue iaea?aeoaaoe aaoa?iceniee aoaeo ca o?iaeaei?noth oa
e?euee?noth aoeuea c eoua.

Aeae?eaii eiia?iaoe?? no?auoaaiue oa naiicaieeaiiy neeaaeieo
i?aeaeaeiaeo a?a?eae?a, ye? ca o?iaeaei?noth ia?aaeuothoue na?aaeith
iaio aaouee?anueeeo oi?i: 93 ae o 88.1439n6, 85.568n9 o
A?ei?onueea 3, 86.331n138 oa ?io?.

A on?o eiia?iaoe?yo aeuaiey?oueny cia/ia e?euee?noue o?aina?aneaieo ca
aenieith o?iaeaei?noth oa ?? aeaiaioaie iauaaee?a.

9. A iauaaee?a, iaea?aeaieo a?ae aae?inoaaiiy oa naiicaieeaiiy neeaaeieo
i?aeaeaeiaeo a?a?eae?a, aeyaeaia neaaea iaaaoeaia -0,116…-0,289 ?
iiceoeaia 0,130…0,342 ei?aeyoe?eia caeaaei?no? i?ae ?iaeaenii
o?aaeaiiy o?oiooi?icii eenoe?a ? o?iaeaei?noth. Oea na?ae/eoue i?i
iacaeaaeia ?o oniaaeeoaaiiy ? iiaeeea?noue ii?aeiaiiy a iaeiiio aaiioei?
aenieeo iieaciee?a oeeo iciae.

Aeei?enoaiiy a naeaeoe?? aeo?aeiiai iaoa??aeo, noai?aiiai ia iniia?
o?eiaaiaoe/ii a?aeaeaeaieo aeae?a, aea? ciiao iaea?aeoaaoe iauaaeee c
aenieei ai?noii o aoeueaao e?ioiaeth. I?iyaeaiiy iciaee o iauaaee?a
caeaaeeoue a?ae oaiioeiiaiai ?? i?iyaeaiiy o aaouee?anueeeo oi?i.

A naeaeoe?? ia e?ioiaeeno?noue iaiao?aeii i?iaiaeeoe iane/oth/? no?a-

uoaaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a oa iaea?aeoaaoe iauaaeee a?ae
?o naiicaieeaiiy.

10. Anoaiiaeaia neaaea iiceoeaia 0,136…0,317 ? iaaaoeaia -0,018…
-0,104 ei?aeyoe?eia caeaaei?noue i?ae ?iaeaenii o?aaeaiiy o?oiooi?icii
eenoe?a ? e?ioiaeeno?noth, neaaea iaaaoeaia -0,098…-0,209 ? neaaea
iiceoeaia 0,029…0,323 i?ae o?iaeaei?noth ? e?ioiaeeno?noth iauaaee?a
neeaaeieo i?aeaeaeiaeo a?a?eae?a.

A?aenooi?noue ei?aeyoe?eii? caeaaeiino? i?ae aeacaieie iciaeaie a ?yae?
eiia?iaoe?e aeaco? ia iacaeaaeia oniaaeeoaaiiy ? i?i iiaeeea?noue
ii?aeiaiiy ?o a i?ioean? naeaeoe??.

11. Aeei?enoaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a a naeaeoe?? ea?oiie?
aea? ciiao noai?eoe naeaeoe?eiee iaoa??ae c aenieith no?ee?noth i?ioe
nooi? ooca??icii? aieeecie.

Anoaiiaeaia iiaeeea?noue ii?aeiaiiy no?eeino? i?ioe ooca??ico ?c
iieueiaith no?ee?noth i?ioe o?oiooi?ico.

12. Aeei?enoiaoth/e a naeaeoe?? neeaaei? i?aeaeaeia? a?a?eaee, iaie
aeae?eaii eiia?iaoe?? no?auoaaiiy oa noai?aii naeaeoe?eiee iaoa??ae
ea?oiie? c aenieith no?ee?noth i?ioe nooi? ooca??icii? aieeecie.

Anoaiiaeaii i?i iacaeaaeii oniaaeeoaaiiy iieueiai? no?eeino? i?ioe
o?oiooi?ico ? ooca??ico, a ioaea, ? iiaeeea?noue ?o ii?aeiaiiy a
i?ioean? naeaeoe??.

13. Aiane?aeie aeei?enoaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a iaie
noai?aii naeaeoe?eiee iaoa??ae ea?oiie? c eiiieaenii
ainiiaea?nueei-oe?iieo iciae: o?iaeaei?noue — iiiaae 1000 a/eou,
e?ioiaeeno?noue 20%, ?iaeaen o?aaeaiiy eenoe?a o?oiooi?icii — 1,2,
no?ee?noue aoeuea i?ioe o?oiooi?ico — 9,0 aae?a, no?ee?noue i?ioe
ooca??ico — 7,0 aae?a oa ?ioa.

14. Noai?aii a ni?aaaoi?noa? /ioe?e ni?oe ea?oiie?: ?aii?e Iia?iue,
na?aaeiuei?aii?e Eoiaaa, na?aaeiueinoeae? ssa?? oa Neia’yiea,. Eoiaaa oa
Neia’yiea caianai? aei ?a?no?o ni?o?a ?ineei Oe?a?ie, ?io? aeaa
i?ioiaeyoue aea?aeaaia ni?oiaei?iaoaaiiy.

?AEIIAIAeAOe??

Ae?iaieoeoao.

Ai?iaaaeeoe iia?, a?aeiinii no?ee? i?ioe o?oiooi?ico oa c eiiieaenii
?ioeo ainiiaea?nueei-oe?iieo iciae ni?oe ea?oiie?, caianai? aei ?a?no?o
ni?o?a ?ineei Oe?a?ie: na?aaeiuei?aii?e Eoiaaa — c 1998 ?ieo oa
na?aaeiueinoeaeee Neia’yiea — c 1999 ?ieo.

I?iaanoe ?iciiiaeaiiy iiaeo ni?o?a ea?oiie?, ui i?ioiaeyoue aea?aeaaia
ni?oiaei?iaoaaiiy: ?aii?e Iia?iue oa na?aaeiueinoeaeee ssa??.

I?aeoe/i?e naeaeoe??.

?aeiiaiaeoaaoe aeo?aeiee oa naeaeoe?eiee iaoa??ae ea?oiie?, noai?aiee ia
iniia? o?eiaaiaoe/ii a?aeaeaeaieo aeae?a: 86.579n14, 86.368n17,
86.331n138, 88.1439n6, 85.568n9, 94.35-35 (85.568n9 o EA 78.5053),
94.35-11 (85.568n9 o EA 78.5053), 94.160-43 (85.568n9 o A?ei?onueea 3),
94.58-4 (86.579n14 o Aonoi), 94.248-18 (?1 86.579n14), 94.248-27 (?1
86.579n14), 94.35-9 (85.568n9 o EA 78.5053), 94.160-18 (85.568n9 o
A?ei?onueea 3), 94.160-21 (85.569n9 o A?ei?onueea 3), 94.160-44
(85.568n9 o A?ei?onueea 3), 94.248-24 (?1 86.579n14), 94.248-42 (?1
86.579n14) oa ?io?.

Eiia?iaoe?? no?auoaaiue: 85.568n9 o A?ei?onueea 3, ?1 86.331n138,
?1 86.579n14, 85.568n9 o EA 78.5053, 86.579n14 o Aonoi, 86.579n14 o
Na?oaiie ee?anueeee, 85.368n17 o Na?oaiie ee?anueeee, 85.368n17 o
Aaeeia, Ia?oiia o 88.1439n6, Aaeeia o 86.331n138.

Iaoiaee aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia iniia?
o?eiaaiaoe/ii a?aeaeaeaieo aeae?a.

NIENIE IIOAE?EIAAIEO ?IA?O II OAI? AeENA?OAOe??

1. Oa?anaiei I.I. Aeei?enoaiiy neeaaeieo i?aeaeaeiaeo a?a?eae?a ea?oiie?
a naeaeoe?? ia iieueiao no?ee?noue i?ioe o?oiooi?ico //A?niee aa?a?ii?
iaoee. — 1997. — ?5. — n.71-74.

2. A.N. ?741. Oe?a?ia. Ni?o ea?oiie? — Eoiaaa (Inei/oe A.A….,Oa?anaiei
I.I. oa ?io? (Oe?a?ia)). Caianaii aei ?a?no?o ni?o?a ?ineei Oe?a?ie c
1998 ?. — 1n.

3. A.N. ?763. Oe?a?ia. Ni?o ea?oiie? — Neia’yiea (Inei/oe
A.A….,Oa?anaiei I.I. oa ?io? (Oe?a?ia)). Caianaii aei ?a?no?o ni?o?a
?ineei Oe?a?ie c 1999 ?. — 1n.

4. Oa?anaiei I.I., Inei/oe A.A. I?aeaeuaiiy iieueiai? no?eeino? i?ioe
o?oiooi?ico a iioiinoa? neeaaeieo i?aeaeaeiaeo a?a?eae?a
ea?oiie?//Caoeno ?ineei. — 1998. — ?5.-N.17.

5. Oa?anaiei I.I. Aeei?enoaiiy a naeaeoe?? aeo?aeiiai iaoa??aeo,

noai?aiiai ia iniia? o?eiaaiaoe/ii a?aeaeaeaieo aeae?a
//Ea?oiiey?noai.-E.,1998.-Aei.28.-N.70.

6. Inei/oe A.A., Iaca? N.A., A?eoeaiei E.I., Oeiioaiei O.A., Inei/oe

A.A., Oa?anaiei I.I Iia? ni?oe ea?oiie? Ia??e, ?aeo?n ?
Eoiaaa.//Ea?oiiey?noai.-E.,1998.-Aei.28.-N.109.

7. Oa?anaiei I.I., Inei/oe A.A. Aea/aiiy iiaiai aeo?aeiiai iaoa??aeo
aeey naeaeoe?? ea?oiie? ia o?oiooi?ino?ee?noue//Naeaeoe?y ea?oiie? ia
no?ee?noue i?ioe o?oiooi?ico a eiiieaen? c ?ioeie iciaeaie: Oace
aeii./?i.-o. ea?oiiey?noaa OAAI. — Iai?oa?aa — Ia?ioeii, 1994. — N.52.

8. Oa?anaiei I.I. I?i ii?aeiaiiy iieueiai? no?eeino? i?ioe o?oiooi?ico
eenoe?a ? aoeuea ea?oiie?//Iaoeia? iniiae aaaeaiiy ea?oiiey?noaa Oe?a?ie
a ?eieiaeo oiiaao: Oace.aeii.eiio.iie.a/ai. ? niaoe?ae?no?a, 23-24
o?aaiy 1996 ?./?i.-o. ea?oiiey?noaa OAAI. — Iai?oa?aa. — 1996. — N.32.

9. Oa?anaiei I.I. Oniaaeeoaaiiy o?oiooi?ino?eeino? i?e aeei?enoaiiy
neeaaeieo i?aeaeaeiaeo a?a?eae?a ea?oiie?//I?iaeaie caoenoo ?ineei a?ae
oe?aeeeaeo i?aai?ci?a a no/anieo aeiiii?/ieo oa aeieia?/ieo oiiaao.;
Oac.aeii. — Ee?a. — 1996. — n.62.

A I I O A Oe ? ss

Oa?anaiei I.I. Aeei?enoaiiy a naeaeoe?? ia o?oiooi?ino?ee?noue
aeo?aeiiai iaoa??aeo ea?oiie?, noai?aiiai ia iniia? o?eiaaiaoe/ii
a?aeaeaeaieo aeae?a. — ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
n?eueneiainiiaea?nueeeo iaoe ca niaoe?aeuei?noth 06.01.05 — naeaeoe?y ?
ian?iieoeoai. — ?inoeooo oeoe?iaeo ao?ye?a OAAI, Ee?a, 1998.

Caoeua?oueny aeena?oaoe?y, a ye?e iiaeai? ?acoeueoaoe aeine?aeaeaiue ii
iaa?oiooaaiith iaoiae?a aeei?enoaiiy aeo?aeiiai iaoa??aeo, noai?aiiai ia
iniia? Solanum bulbocastanum oa ?ioeo aeae?a, a naeaeoe?? ia
o?oiooi?ino?ee?noue a eiiieaen? c ?ioeie ainiiaea?nueei-oe?iieie
iciaeaie.

Aeo?aeiee iaoa??ae iiaeia caeo/eoe a naeaeoe?eio ?iaioo ye iaoiaeii
aae?inoaaiiy, oae ? iaea?aeaiiyi iauaaee?a a?ae eiai naiicaieeaiiy, a
oaeiae no?auoaaiiyi i?ae niaith. Aeioe?eueii i?aeae?aoe aaouee?anuee?
ia?e i?e naeaeoe?? ia iieueiao no?ee?noue i?ioe o?oiooi?ico ca ??
neeaaeiaeie.

-Eeth/ia? neiaa: ea?oiiey, i?aeaeaeia? a?a?eaee, o?oiooi?ic, no?ee?noue,
ainiiaea?nuee? oa?aeoa?enoeee.

A I I I O A Oe E ss

Oa?anaiei I.A. Eniieueciaaiea a naeaeoeee ia oeoiooi?ionoie/eainoue
enoiaeiiai iaoa?eaea ea?oioaey, iieo/aiiiai i?e o/anoee oeeiaaiaoe/anee
ioaeaeaiiuo aeaeia. — ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa
naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe 06.01.05 — naeaeoeey e
naiaiiaiaenoai. — Einoeooo naoa?iie naaeeu OAAI, Eeaa, 1998.

Caueuaaony aeenna?oaoeey, a eioi?ie i?aaenoaaeaiu ?acoeueoaou
enneaaeiaaiee ii iainiiaaieth iaoiaeia eniieueciaaiey enoiaeiiai
iaoa?eaea, iieo/aiiiai i?e o/anoee Solanum bulbocastanum e ae?oaeo
aeaeia, a naeaeoeee ia oeoiooi?ionoie/eainoue n eiiieaenii ae?oaeo
oicyenoaaiii-oeaiiuo i?eciaeia.

Enoiaeiue iaoa?eae iiaeii eniieueciaaoue a naeaeoeeiiiie ?aaioa eae
iaoiaeii aae?inniaaieai, oae e iieo/aieai iioiinoaa io aai naiiiiueaiey,
a oaeaea no?aueaaiey eo iaaeaeo niaie. AEaeaoaeueii iiaeae?aoue
?iaeeoaeueneea oi?iu i?e naeaeoeee ia iieaaoth onoie/eainoue i?ioea
oeoiooi?ica ca aa ninoaaiuie.

Eeth/aaua neiaa: ea?oioaeue, iaaeaeaeiaua aea?eaeu, oeoiooi?ic,
onoie/eainoue, oicyenoaaiiua oa?aeoa?enoeee.

A N N O T A T I O N

Tarasenko O.O. Utilization of Potato Parent Material Obtained with
Involvement of Phylogenetically Remote Species in Breeding for
Resistance to Late Blight. — Manuscript.

Dissertation for submission for the degree of Candidate of Agricultural
Science on speciality 06.01.05 — breeding and seed production. — UAAS
Institute of Agriculture, Kyiv, 1998.

A thesis is defended, it contains results of research on grounding
methods of using parent material, obtained with involvement of Solanum
bulbocastanum and other species, in breeding for resistance to late
blight parallel with other economic characters.

The parent material can be used in breeding process as by backcrossing
method so by obtaining progeny from self-pollynation and by crossing. It
is advisable to select parent forms in breeding for field resistance to
the potato late blight in accordance with its components.

Key words: potatoes, interspecific hybrids, late blight resistance,
economic characters.

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