Aei?i?iiao?ianueeee aea?aeaaiee oi?aa?neoao

Eoeueaa/ei TH??e Ethoeeiiae/

OAeE 591.55+592:615

NOAI NO?OEOO?II? I?AAI?CAOe?? AACO?AAAOIEO OAA?EI I?AeNOEEEE NOAIIAEO
E?N?A A OIIAAO I?IIENEIAIAI CAA?OAeIAIIss

— aeieia?y

Aaoi?aoa?ao

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Aei?i?iiao?ianuee – 1999

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia ia eaoaae?? ciieia?? oa aeieia?? Aei?i?iiao?ianueeiai
aea?aeaaiiai oi?aa?neoaoo I?i?noa?noaa ina?oe Oe?a?ie.

Iaoeiaee ea??aiee: Eaiaeeaeao
a?ieia?/ieo iaoe, aeioeaio

Aa?nia A?eoi? Ieaenaiae?iae/,

eaoaae?a ciieia?? oa aeieia??

Aei?i?iiao?ianueeiai aea?aeaaiiai

oi?aa?neoaoo

Io?oe?ei? iiiiaioe:

Aeieoi? a?ieia?/ieo iaoe, i?ioani? Aiinoieia Eaii?ae Aai?a?eiae/,
N?ioa?iiieuenueeee aea?aeoi?aa?neoao, eaoaae?a aeieia?? oa
?aoe?iiaeueiiai i?e?iaei-ei?enooaaiiy, caa?aeoaa/.

Aeieoi? a?ieia?/ieo iaoe, aeioeaio ss?ioaiei Ieeiea Ieeieaeiae/,
Aeiiaoeueeee aea?aeoi?aa?neoao, eaoaae?a ciieia??, caa?aeoaa/

I?ia?aeia onoaiiaa:

Iaoe?iiaeueiee iaaeaaia?/iee oi?aa?neoao ?iai? I.I. Ae?aaiiaiiaa,
eaoaae?a ciieia??, i. Ee?a

Caoeno a?aeaoaeaoueny « 1 »a?oaeiy 1999 ?. i 12:00 aiaeei? ia can?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae 08.051.04, Aei?i?iiao?ianueeee
aea?aeaaiee oi?aa?neoao I?i?noa?noaa ina?oe Oe?a?ie, 320625,
i.Aei?i?iiao?ianuee, i?iaoeie Iaoeiaee, 13, ei?i. 17, aoae. 611.

C aeena?oaoe??th iiaeia iciaeiieoeny o a?ae?ioaoe? Aei?i?iiao?ianueeiai
aea?aeaaiiai oi?aa?neoaoo, 320625, i.Aei?i?iiao?ianuee, i?ia. Iaoeiaee,
13.

Aaoi?aoa?ao ?ic?neaiee 29 aeiaoiy 1999?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee _______________________ Aeoaeia A.I.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. Caoeno iaaeieeoiueiai na?aaeiaeua caaaeaee aoa, ? ?
aoaea iaei??th c aeooaeueieo i?iaeai aeey eiaeii? e?a?ie. Caa?oaeiaiiy
aoiinoa?e oiene/ieie i?iaeoeoaie iaiineoue oeiaeo ia o?eueee
iaaeieeoiueiio na?aaeiaeuo, aea e caei?ia’th ethaeeie. A ca’yceo c
??ciei noaiii aeaa?aaeaoe?? a?inoa?e ae??oaeueia cia/aiiy ia no/aniiio
aoai? iaaoaa? ?aa?iiaeueiee aniaeo i?e?iaeiei?enooaaiiy. Ana a?eueoa
oaaaa aei i?iaeai ioi?iie i?e?iaee iaoiiaeea iaiao?aei?noue i?iaaaeaiiy
a?i?iaeeeaoe?eieo aeine?aeaeaiue c aeei?enoaiiyi ??cieo a?ieia?/ieo
nenoai ye a ?aieao iaoe?iiaeueii?, oae ? i?aeia?iaeii? ae?yeueiino?.
Na?aae ??ciiiai?oieo a?i?iaeeeaoi??a oaeeo ye i?e?ii?aai?cie, ?ineeie,
o?aaaoi? oaa?eie, ii i?aao a?aeia i?noea caeiathoue aaco?aaaoi? oaa?eie.
Iaaeaenieee oaoiiaaiiee oa aio?iiiaaiiee oene, iacaaeainiaai?noue
i?e?iaei?ano?niiai iioaioe?aeo i?ecaiaeeoue aei aeieeiaiiy no?eeeo
iani?eyoeeaeo ci?i a aeinenoaiao, aeaa?aaeaoe?? i?e?iaeiiai na?aaeiaeua.
Ia oii? oeeo ci?i aeieea? iiaeeea?noue aeei?enoaiiy aeieo i?aaenoaaiee?a
aaco?aaaoieo i?aenoeeee aeey a?i?iaeeeaoe??, ioe?iee noaio ai?aaeieo oa
ooo/ieo e?niaeo a?iaaioeaiic?a a oiiaao noaiiaiai I?eaei?i?ia’y i?ae
aieeaii ??cieo oi?i aio?iiiaaiieo /eiiee?a.

Ca’ycie aeena?oaoe?? c ieaiii IAe? eaoaae?e ciieia?? oa aeieia?? AeAeO.
?iaioa aeeiioaaeanue ca?aeii c ae?th/ith iaoeiaith i?ia?aiith
«A?ieia?/i? ?ano?ne, ?o ?aoe?iiaeueia aeei?enoaiiy, a?aeoai?aiiy oa
ioi?iia» (?icae?e «Oaa?eiiee na?o, ?aoe?iiaeueia aeei?enoaiiy oa
ioi?iia»). Ca?aeii c oe??th i?ia?aiith i?iaiaeeeenue aeine?aee a iaaeao
aea?aeathaeaeaoieo ieaiiaeo oai Eiiieaenii? aeniaaeeoe??, eaoaae?e
ciieia?? oa aeieia?? Aei?i?iiao?ianueeiai aea?aeaaiiai oi?aa?neoaoo (??
aea?ae-?a?no?aoe??: 0296V006370 (1994-1996 ??.), 0197V000680 (c 1997 ?.
ii ianooiiee /an). Cacia/aia oaiaoeea eii?aeeio?oueny I?i?noa?noaii
ina?oe Oe?a?ie, Iaoeiaith ?aaeith c i?iaeai a?oioiciaanoaa IAI Oe?a?ie.

Iaoa ?iaioe oa caaaeaiiy aeine?aeaeaiiy.

Iniiaiith iaoith aeine?aeaeaiiy ? aeyaeaiiy iiaeeeaino? aeei?enoaiiy
i?aenoeeeiaeo aaco?aaaoieo aeey a?i?iaeeeaoe?? oa ioe?iee noaio
ai?aaeieo eiiieaen?a i?e noeoean?eieo ci?iao i?e?iaeieo oa ooo/ieo
e?niaeo a?iaaioeaiic?a a oiiaao noaiiaiai I?eaei?i?ia’y i?ae aieeaii
??cieo oi?i aio?ii?/ieo /eiiee?a. Caaaeaiiyie aeineeaeaeaiiy aoee:

SYMBOL 45 \f «Symbol» \s 12 — aeyaeoe aeaeiaee neeaae, oaeniiii?/io
no?oeoo?o oa ?o ci?ie o aaco?aaaoieo i?aenoeeee i?e?iaeieo e?niaeo oa
ooo/ieo e?niianaaeaeaiue, ye? ciaoiaeyoueny i?ae aieeaii i?iieneiaiai
caa?oaeiaiiy;

SYMBOL 45 \f «Symbol» \s 12 — aecia/eoe ci?ie neiaeieia?/ieo iieaciee?a
i?aenoeeeiaeo aaco?aaaoieo i?ae aieeaii i?iieneiaiai caa?oaeiaiiy, oa
iiaeeeaino? ?o aeei?enoaiiy a a?i?iaeeeaoe?eieo aeine?aeaeaiiyo;

SYMBOL 45 \f «Symbol» \s 12 — aeine?aeeoe iniaeeaino? oi?ioaaiiy oaoie
aaco?aaaoieo i?aenoeeee ia ae?eyioe? e?niai? ?aeoeueoeaaoe??;

SYMBOL 45 \f «Symbol» \s 12 — aecia/eoe aieea e?nioeiieia?/ieo oaeoi??a
ia no?oeoo?o i?aenoeeeiai? aioiiioaoie ae?eyiee e?niai? ?aeoeueoeaaoe??;

SYMBOL 45 \f «Symbol» \s 12 — aeine?aeeoe iniaeeaino? iaeiie/aiiy
aaaeeeo iaoae?a aaco?aaaoieie i?aenoeeee a oiiaii /enoeo oa i?aeaeaieo
i?iieneiaiio caa?oaeiaiith i?e?iaeieo oa ooo/ieo e?niianaaeaeaiiyo;

SYMBOL 45 \f «Symbol» \s 12 — aeei?enoaoe iiaeaeuei? a?oie aaco?aaaoieo
aeey i?iaaaeaiiy aeioieneei-eia?/ieo aeine?aeaeaiue;

SYMBOL 45 \f «Symbol» \s 12 — ?icaeyiooe ci?ie a eiiieaenao
aaco?aaaoieo i?aenoeeee o ca’yceo c i?aeoii-eaiiyi i?e?iaeieo caieaaieo
a?iaaioeaiic?a aiane?aeie oaooieo i?ae?iaie;

SYMBOL 45 \f «Symbol» \s 12 — aecia/eoe iiaeeeaino? aeei?enoaiiy
iieaciee?a noi?nii? ci?ie ?ineeiiiai iie?eao oa oaoie aaco?aaaoieo
i?aenoeeee ia oaoi?noe/iiio, ii?oi-eia?/iiio, aeioieneeieia?/iiio ??aiyo
aeey a?i?iaeeeaoe?eieo aeine?aeaeaiue.

Iaoeiaa iiaecia io?eiaieo ?acoeueoao?a.

Aeine?aeaeaiiy i?iaiaeeeenue a noia??iaeono??aeueiiio ?aa?ii? Oe?a?ie,
cie?aia a ?aeii? i?iieneiai? ciie i.Aei?i?iaeca?aeeinueea, ia oa?eoi???
Cao?aeiiai Aeiiaano, yea ciaoiaeeoueny i?ae aieeaii a??ie/iaeiaoaii?
i?iieneiaino?. Aiie aacothoueny ye ia oaoi?noe/ieo, oae ? ia
aeioieneeieia?/ieo ci?iao, ye? a?aeaoaathoueny o oaoi? aaco?aaaoieo
i?aenoeeee. Aia?oa aeey oeueiai ?aa?iio ?icaeyiooi noi?ni? ci?ie
?ineeiiiai iie?eao oa oaa?eiiiai ianaeaiiy i?aenoeeee aiane?aeie
oaoiiaaiiiai aieeao. Aeai?, ye? ?nioaaee ?ai?oa, aena?oethaaee eeoa
oaoi?noe/i? aai oei?enoe/i? ci?ie ?ineeiiiai oa oaa?eiiiai na?oo. Iaie
iaea?aeai? iia?, cia/ii iiai?o? aeai?, ye? iiaeooue aooe aeei?enoai? a
iii?oi?eiaiaeo aeine?aeaeaiiyo c iaoith ?iaeeeaoe?? iaaaoeaiiai aieeao
caa?oaeiaiiy ia aeinenoaie.

I?aeoe/ia cia/aiiy io?eiaieo ?acoeueoao?a.

?acoeueoaoe aeine?aeaeaiue a i?iieneia?e cii? i. Aei?i?iaeca?aeeinueea
iiaeooue aooe aeei?enoai? a «I?iaeo? i?ia?aie aeoiaeo i.
Aei?i?iaeca?aeeinueea c aeieia?/ii? e?ece», ?ic?iaeaii? I?ii?iiii
Oe?a?ie, Oe?a?inueeei aea?aeaaiei iaoeiai-aeine?aeiei ?inoeoooii acioii?
i?iieneiaino?. Aeine?aeaeaiiy, i?iaaaeai? ia ae?eyioe? e?niai?
?aeoeueoeaaoe??, iiaeooue aooe aeei?enoai? aeey aeai?o eiino?oeoe??
ooo/ieo a?oio?a oa no?eeeo aea?aaieo ii??ae ia oaooieo a?aeaaeao
Cao?aeiiai Aeiiaano. Iaoa??aee aeena?oaoe?? aeei?enoiaothoueny a
niaoeeo?nao aeey nooaeaio?a aeueo o/aiaeo caeeaae?a, eaeoe?yo c ioi?iie
iaaeieeoiueiai i?e?iaeiiai na?aaeiaeua o/iyi oe?e oa oe?ieei aa?noaai
ianaeaiiy.

Iniaenoa o/anoue aaoi?a a io?eiaieo ?acoeueoaoao, ye? i?aaenoaaeai? a
aeena?oaoe??. Aeena?oaoe?y ? iniaenoith iaoeiaith i?aoeath, yeo aeeiiaii
aaoi?ii i?ioyaii 5 ?ie?a o neeaae? Eiiieaenii? aeniaaeeoe?? AeAeO c
aea/aiiy e?n?a noaiiai? ciie Oe?a?ie. Aaoi? a?aa iniaenoo o/anoue o
cae?aii?, aecia/aii? oa ia?iaoe? aeine?aeaeaiue iieueiaiai iaoa??aeo, a
i?iaaaeaii? eaai?aoi?ieo aeine?aeaeaiue, i?aeaioiaoe? iaoeiaeo
ioae?eaoe?e. Aaoi?ii i?aeaioiaeaii 8 naiino?eieo ioae?eaoe?e, a a
i?aoeyo, ui iioae?eiaai? a ni?aaaoi?noa?, /anoea iniaenoi? o/ano?
noaiiaeoue 25-50 %.

Ai?iaaaeaeaiiy ?acoeueoao?a ?iaioe. Iaoa??aee ?iaioe aeena?oaioa
Eoeueaa/ei TH.E. ? eiie?aoi? ?aeiiaiaeaoe?? ui aei aeei?enoaiiy
oa?aeoa?enoee aaco?aaaoieo i?aenoeeee noaiiaeo e?n?a aeicaieythoue
i?iaiaeeoe aiae?c aieeao oaoiiaaiieo /eiiee?a ia caaaeueio no?oeoo?o oa
ooieoe?iioaaiiy aaco?aaaoieo, ?o ?ieue a i?aeo?eiaii? aiiainoaco
aeinenoai, iaeiie/aii? oiene/ieo ?a/iaei.

Oe? iaoa??aee oa ?aeiiaiaeaoe?? aeei?enoaii oa ai?iaaaeaeaii Oi?aae?iiyi
aeieia?/ii? aaciaee c iaoith caa?aaeaiiy a?i??ciiiai?oiino? oa ioi?iie
aeinenoai ?aa?iio ia aenia?eiaioaeueii-ae?iaie/?e ae?eyioe? e?niai?
?aeoeueoeaaoe?? AeOE Iaaeia?aaeaoa?eey. Iaoa??aee c oaie aeena?oaoe?eii?
?iaioe aeei?enoaii i?e neeaaeaii? ca?o?a oa oaiaoe/ieo ieai?a a IAe?
a?ieia?? AeAeO. Aiie oa?eoee aei o/aiaeo i?ia?ai oa aeei?enoiaothoueny a
o/aiaiio i?ioean? a eo?nao caaaeueieo oa niaoe?aeueieo aeenoeeie?i ia
eaoaae?? ciieia?? oa aeieia?? Aei?i?iiao?ianueeiai aea?aeaaiiai
oi?aa?neoaoo.

Ai?iaaoe?y ?iaioe. Iaoa??aee aeena?oaoe?? i?ieoee ai?iaaoe?th ia
Iaoe?iiaeuei?e eiioa?aioe?? ii caa?aaeaiith a?i??ciiiai?oiino? a Oe?a?i?
(Eai?a, 1997), ia ia?aae? c aeieia?? oa ?iaeaia???
(Aei?i?iaeca?aeeinuee, 1996), ia ia?aae? «Oaa?eiiee na?o A?ei-?onueeiai
Iie?nny oa eiai ?aoe?iiaeueia aeei?enoaiiy» (Aiiaeue, 1998),
i?aeia?iaei?e iaoeiai-i?aeoe/i?e eiioa?aioe?? «Sustainable divelopment:
system analysis in ecology» (Naaanoiiieue, 1996), i?aeia?iaeieo
eiioa?aioe?yo «O?aioe?y oa Oe?a?ia, iaoeiai-i?aeoe/iee aeina?ae o
eiioaeno? ae?aeiao iaoe?iiaeueieo eoeueoo?» (Aei?i?iiao?ianuee, 1995,
1997, 1998), ia 1 i?aeia?iaei?e eiioa?aioe?? «No?eea caa?oaeiaiiy
iaa-eieeoiueiai na?aaeiaeua oa aeieia?/ia aaciaea» (Aei?i?iiao?ianuee,
1995).

Ioae?eaoe??. Ca oaiith aeena?oaoe?? iioae?eiaaii 19 iaoeiaeo ?ia?o,
na?aae yeeo 8 naiino?eieo.

No?oeoo?a ? ianya ?iaioe. Iaoa??aee aeena?oaoe?? aeeeaaeai? ia 238
noi??ieao, aeania oaeno caeia? 142 noi??iee. Aeena?oaoe?y neeaaea?oueny
?c anooio, 9 ?icae?e?a, aeniiae?a oa ?aeiiaiaeaoe?e, nienea aeei?enoaieo
aeaea?ae. Nienie e?oa?aoo?e i?noeoue 267 iaeiaioaaiiy, na?aae yeeo 28
?iicaiieie iiaaie.

CI?NO ?IAIOE

O anooi? iaa?oioiao?oueny aeooaeuei?noue oa noai iaoeiai? i?iaeaie, iaoa
oa caaaeaiiy aeine?aeaeaiue.

INIIAI? *EIIEEE, ssE? I?ECAIAessOUe AeI AIO?III-OAOIIAAIIEO CI?I A
AEINENOAIAO

A ?icae?e? ?icaeyaea?oueny aieea ia iaaeieeoi? na?aaeiaeua oaeeo
aio?iiiaaiieo /eiiee?a ye o?aai?caoe?y, n?euenueea ainiiaea?noai,
a??ie/a ae?iaieoeoai, i?iieneia? aeeeaee. Aeiaaaeaii, ui iaea?eueo
iaaacia/aieie aeey aeiae?eey ? aeeeaee i?iieneiaiai ae?iaieoeoaa oa
aeiaoaaiiy ei?enieo eiiaeei. Oe? /eiieee iaaaoeaii aieeaathoue ye ia
o?oi- oae ? ia ciioeaiic.

Iaei??th c neeaaeiaeo /anoei ciioeaiico aenooiathoue aaco?aaaoi? oaa?eie
i?aenoeeee. Oaoi?noe/i? ? no?oeoo?i? ci?ie a ?o iiioeyoe?yo, iniaeeaino?
iaeiie/aiiy aaaeeeo iaoae?a i?aaenoaaieeaie ??cieo a?oi
aeei?enoiaothoueny a a?i?iaeeeaoe?eieo aeine?aeaeaiiyo i?e oaoiiaaiiiio
caa?oaeiaii? na?aaeiaeua.

Iniiaia /anoeia i?aoeue i?enay/aia ie?aii oaoi?noe/iei, no?oeoo?iei
ci?iai a iiioeyoe?yo oaa?ei, aeioieneeieia?/iei aeine?aeaeaiiyi ie?aieo
?o a?oi.

O?CEEI-AAIA?AO?*IA OA?AEOA?ENOEEA ?AEIIO AeINE?AeAEAIUe

?aeii aeine?aeaeaiue iaeaaeeoue aei i?aeciie ni?aaaei?o noai?a noaiiai?
ciie Oe?a?ie. A ?icae?e? iaaiaeyoueny aeai? i?i caaaeuei?
o?ceei-aaia?ao?/i? oiiae ?aa?iio, ee?iao, ?aeue?o oa aaiii?oieia?th
a?oioo, a?ae?ieia?/i? oiiae, ?ineeii?noue oa oaa?eiiiee na?o, ye?
aena?oeai? a i?aoeyo a?aeiieo a/aieo (Eaa?aiei,1954; Aunioeeiai,1962;
Naiaithou,1948; Eaeeieiie,1972; O?aaeaaaa, Aoeaoiaa,1969;
Aaeueaa?aea,1971 oa ?ioeo)

IAOA??AEE, IAOIAeIEIA?ss OA IAOIAeEEE AeINE?AeAEAIUe

Iaoa??aee aeine?aeaeaiue iaea?aeai? o neeaae? Eiiieaenii? aeniaaeeoe??
Aei?i?iiao?ianueeiai aea?aeoi?aa?neoaoo ii aea/aiith e?n?a noaiiai? ciie
Oe?a?ie a ia??iae c 1994 ii 1998 ??.

Ca?? iaoa??aeo i?iaiaeeany ii caaaeuei?e iaoiaeeoe? a?oioiai-ciieia?/ieo
aeine?aeaeaiue (Aeey?ia, 1975) noi?nii c aeei?enoaiiyi ianoie Aa?aa?a.
Ca ia??iae aeine?aeaeaiue c?a?aii 1200 i?ia. Ai?no aaaeeeo iaoae?a a
aaco?aaaoieo i?aenoeeee aeeiiaii iaoiaeii aoiiii-aani?aoe?eii?
niaeo?ioioiiao??? ia AAS-30 (Oaaacia, Oeaeaa, 1983). I?aeaioiaeaii 460
i?ia ? i?iaaaeaii iiiaae 3220 aeaiaioi-aecia/aiue. E??i oeueiai
i?iaiae?ciaaii ai?no aaaeeeo iaoae?a a 50 i?iaao a?oioo oa i?aenoeeee,
oa 45 i?iaao oei?enoe/iiai iaoa??aeo. Iaoaiaoe/iee ia?ia?oie
?acoeueoao?a aeeiiaii ca noaiaea?oieie iaoiaeeeaie (Eaeei, 1990) ia
eiiiuethoa?? IBM PC/AT 486 c aeei?enoaiiyi iaeaoa Exel-97.

A ?icae?e? iaaaaeaii oa?aeoa?enoeeo ae?eyiie, ia yeeo i?iaiaeeeenue
aeine?aeaeaiiy. Ca eiio?ieuei? i?eeiyo? ae?eyiee aeieia?/iiai i?io?eth
ia oa?eoi??? I?enaia?nueeiai i?aeia?iaeiiai a?inoa?iiai noaoe?iia?o.
Ae?eyiee, ye? ciaoiaeyoueny i?ae aieeaii i?iieneiaiai caa?oaeiaiiy,
?icoaoiaai? iiaeeco i.Aei?i?iaeca?aeeinueea, a ae?eyiee e?niai?
?aeoeueoeaaoe?? ia oa?eoi???, yea i?eeyaa? aei oaoo Cao?aeiiai Aeiiaano.

CI?IE NO?OEOO?E IANAEAIIss AACO?AAAOIEO I?AeNOEEEE I?E?IAeIEO E?NIAEO OA
OOO*IEO IANAAeAEAIUe I?Ae AIEEAII I?IIENEIAIAI CAA?OAeIAIIss

Iniaeeaino? ci?ie oaoie aaco?aaaoieo i?aenoeeee ooo/ieo a?eiaeaoe??aeo
ianaaeaeaiue i?ae aieeaii i?iieneiaiai caa?oaeiaiiy. I?iaaaeaii
ii??aiyiiy oaoie aaco?aaaoieo oaa?ei i?aenoeeee c eiio?ieueieo ooo/ieo
ianaaeaeaiue oa ianaaeaeaiue, ui ciaoiaeyoueny i?ae aieeaii oeaiaioiiai
ae?iaieoeoaa, a oaeiae Aei?i?iaeca?aeeinueeiai ae?iaie/iai ia’?aeiaiiy
«Acio» (AeAI «Acio»). A cii? ae?? i?iieneiaiai caa?oaeiaiiy e?euee?noue
aeae?a oaa?ei ciaioo?oueny a 1,2-1,3 ?ace o ii??aiyii? c eiio?ieueieie
ianaaeaeaiiyie, aea ii?yae c oeei ca?eueoo?oueny /enaeuei?noue oaa?ei,
e?euee?noue aeae?a noia?aeiieiaio?a. A cii? ae?? AeAI «Acio» oe?eeii
ci?ith?oueny aeaeiaee neeaae /a?a’ye?a. Cai?noue Lumbricus terrestris,
ca?a?no?iaaiiai o eiio?ieueieo ianaaeaeaiiyo, c’yaeythoueny Dendrobaena
octaedra, Eisenia foetida, Octolasium complanatum. A?aeaoaathoueny ci?ie
neeaaeo Staphylinidae, Mollusca. O cii? caa?oaeiaiiy c’yaey?oueny
Cochlicopa lubrica, yea aoea a?aenooiy o eiio?ieueieo ianaaeaeaiiyo. O
cii? ae?? oeaiaioiiai ae?iaieoeoaa a 2,4 ?ace ciaioo?oueny e?euee?noue
?iaeei eeano eiiao. A?aenooi? i?aaenoaaieee Dermestidae, Tenebrionidae,
Gryllidae. A?aeaoaa?oueny ciaioaiiy cia/aiue ?iaeaeno Oaiiiia,
?ic?aoiaaiiai ii /enaeueiino? c 2,22-2,67 aei 1,36-1,71, oa ii a?iian? —
c 2,87-3,14 aei 2,17-2,73 o ii??aiyii? c eiio?ieueieie ae?eyieaie. Aeey
ooieoe?iiaeueieo a?oi a cii? i?iieneiaiai caa?oaeiaiiy a?aeaoaa?oueny
ciaioaiiy /enaeueiino? i?aaenoaaiee?a o?oioaa?a oa ca?eueoaiiy
nai?ioaa?a i?e

aeineoue aenie?e /enaeueiino? ciioaa?a (?en.1). Iaiao?aeii a?aecia/eoe,
ui oaoia aaco?aaaoieo oaa?ei ooo/ieo ianaaeaeaiue a cii? ae?? AeAI
«Acio» iaioa i?aeeyaa? ci?iai ii??aiyii c oaoiith ooo/ieo ianaaeaeaiue
oeaiaioiiai ae?iaieoeoaa, ui iiaeeeai iia’ycaii c iaeiiai aieo c
ia?eoi?/i?noth ?iaioe AeAI «Acio», a c ae?oaiai c niaeo?ii i?iieneiaeo
aeeeae?a i?aei?e?inoa.

Iniaeeaino? ci?i oaoie aaco?aaaoieo i?aenoeeee caieaaieo e?niaeo
a?iaaioeaiic?a i?ae aieeaii i?iieneiaiai caa?oaeiaiiy. sse ? a ooo/ieo
ianaaeaeaiiyo, ca?a?no?iaaii ciaioaiiy e?eueeino? aeae?a aaco?aaaoieo a
1,5 ?ace i?e ca?eueoaii? o 2 ?ace e?eueeino? aeae?a noia?aeiieiaio?a,
aei yeeo iaeaaeaoue Vallonia costata (7,19 %), Staphylinus caesareus
(8,63 %), Pterostichus melanarius (6,83%). A?aeaoaathoueny ? ye?ni?
ci?ie na?aae aaco?aaaoieo. Cieeathoue Succinea oblonga, Succinea putris,
Lumbricus rubellus, Eisenia tetraedra. Ia ca?a?no?iaaii i?aaenoaaiee?a
Trombiculidae. Ci?ie a?aeaoaathoueny na?aae ooieoe?iiaeueieo a?oi
aaco?aaaoieo. sseui /enaeuei?noue ciioaa?a ia eiio?ieueieo ae?eyieao
noaiiaeoue 35,0-35,8 aec/i2, oi i?ae aieeaii i?iieneiaiai caa?oaeiaiiy
/enaeuei?noue o?oioaa?a ciaioo?oueny o 1,8 ?ace, a nai?ioaa?a o 2,1
?ace. A?aeaoaa?oueny cieaeaiiy ?iaeaeno aeaeiai? ??ciiiai?oiino?,
?ic?aoiaaiiai ca /enaeuei?noth oa a?iianith: 2,83-3,09 i?ioe 1,72-1,94
(ca /enaeuei?noth), oa 2,19-2,39 i?ioe 1,54-1,98 (ii a?iian?).

Ae?AAIINOEEA NOAIO E?NIAEO IANAAeAEAIUe IA Ae?EssIEAO ?AEOEUeOEAAOe?? A
CAO?AeIIIO AeIIAAN? CA AeIIIIIAITH EIIIEAEN?A AACO?AAAOIEO I?AeNOEEEE

Aiae?c ni?eueiino? oaoie aaco?aaaoieo i?aenoeeee ??cieo
aea?aaii-/aaa?ieeiaeo ii??ae oa aa??aio?a ianeiieo a?oio?a ae?eyiee
e?niai? ?aeoeueoeaaoe??. Aeey ianaeaiiy aaco?aaaoieo i?aenoeeee i’yoe
aa??aio?a a?oio?a a ianaaeaeaii? aeaoe?? a?ei?, ea?aaaie, a’yca
iecueeiai, ianeeiee aocueeieenoi?, ae?th/eie i?i?iaeueia ye?nia oa
e?euee?nia ni?euei?noue ca?a?no?iaaia aeey ia?oiai aa??aioo oa ianooiieo
ca iei aa??aio?a a?oio?a. Iaeneiaeueii aeecuee? oaoie aaco?aaaoieo 3-4
oa 3-5 aa??aio?a a?oio?a. Ia caaaeath/e ia iecueeo ye?nio ni?euei?noue
oaoie aaco?aaaoieo a caeaaeiino? a?ae neeaaeo aea?aaii-/aaa?ieeiaeo
ianaaeaeaiue caa?oa? ia naaa oaaao, ui iaeneiaeueii aeecuee? eiiieaene
aaco?aaaoieo i?aenoeeee aeaoe?? a?ei?, a’yca iecueeiai, ianeeiee
aocueeieenoi?, a eiao?oe??ioe ni?eueiino? ii AEaea?o o aaco?aaaoieo c
/aaa?ieeiaeo ianaaeaeaiue ieae/? ii??aiyii c aea?aaieie.

E?euee?nio ni?euei?noue aaco?aaaoieo i?aenoeeee aea?aaii-/aaa?ieeiaeo
ianaaeaeaiue ii ciaioaiith iiaeia iinoaaeoe o oae?e iine?aeiaiino?: a’yc
iecueeee — aeaoe?y a?ea; aeaoe?y a?ea — ae?th/eia; ianeeiea aocueeieenoa
— aeaoe?y a?ea; ianeeiea aocueeieenoa –

ea?aaaia; ianeeiea aocueeieenoa — a’yc iecueeee; ianeeiea aocueeieenoa —
ae?th/eia; ea?aaaia — ae?th/eia; ea?aaaia — aeaoe?y a?ea; a’yc iecueeee
— ae?th/eia.

?iciiae?e oaoie aaco?aaaoieo i?aenoeeee ia ae?eyioe? e?niai?
?aeoeueoeaaoe?? a caeaaeiino? a?ae neeaaeo aea?aaii-/aaa?ieeiaeo
ianaaeaeaiue. Iniaeeaino? oi?ioaaiiy ooo/ieo aaeaoioii?a ia i?noe?
oaooieo a?aeaae?a cia/ii aieeaathoue ia ?iciiae?e a ieo i?aaenoaaiee?a
aaco?aaaoieo i?aenoeeee. Aaeeea cia/aiiy ia? oei aea?aainoith (no?oeoo?a
oa neeaae). O aea?aaii-/aaa?ieeiaeo ianaaeaeaiiyo ia ca?a?no?iaaii
??ceeo ci?i e?eueeino? oaenii?a aaco?aaaoieo, ?o /enei aei??aith? 14-16.
Ne?ae a?aecia/eoe ca?eueoaiiy /enaeueiino? aaco?aaaoieo a ianaaeaeaiiyo
ianeeiee

aocueeieenoi?, a’yca iecueeiai o ii??aiyii? c aeaoe??th a?eith —
Mollusca 6,4 aec ia 10e/ae?a i?ioe 4,6 aec ia 10 e/ae?a, Tenebrionidae
5,6 aec ia 10 e/ae?a i?ioe 1,9 aec ia 10e/ae?a, Silphinidae 13,1 aec ia
10 e/ae?a i?ioe 9,3 aec ia 10 e/ae?a.

?iciiae?e oaoie aaco?aaaoieo i?aenoeeee ia ae?eyioe? e?niai?
?aeoeueoeaaoe?? caeaaeii a?ae aa??aioo ooo/ieo a?oio?a o
aea?aaii-/aaa?ieeiaeo ianaaeaeaiiyo. ?icaeyaeath/e ?iciiae?e
aaco?aaaoieo o i’yoe aa??aioao a?oio?a, iaiao?aeii i?aee?aneeoe ?o
??ciee ye?niee oa e?euee?niee neeaae aeey ie?aii? ii?iaee. ssae?i oaoie
aaco?aaaoieo ia an?o aa??aioao a?oio?a neeaaeathoue i?aaenoaaieee eiiao.
Noia?ia /enaeuei?noue aaco?aaaoieo aeey ae?oaiai aa??aioo a?oio?a
neeaaea? 98 aec ia 10 e/ae?a, /aoaa?oiai aa??aioo a?oio?a — 102 aec ia
10 e/ae?a, o?aoueiai — 196,5 aec ia 10 e/ae?a, i’yoiai — 131 aec ia 10
e/ae?a.

Iniaeeaino? ci?ie neiaeieia?/ieo oa?aeoa?enoee aaco?aaaoieo i?aenoeeee
??cieo aa??aio?a ianaaeaeaiue ae?eyiee e?niai? ?aeoeueoeaaoe?? ia
oa?eoi??? Cao?aeiiai Aeiiaano. Na?aae ii??ae aea?aaii-/aaa?ieeiaeo
ianaaeaeaiue iaea?eueoa /enei aeae?a oaa?ei (27) a?aecia/aii o
ianaaeaeaii? aeaoe?? a?ei?, a iaea?eueoa noia?ia u?euei?noue — 123, 4
aec ia 10 e/ae?a o ianaaeaeaii? a’yca iecueeiai. I?i?iaeueia /enei
aeae?a (14) i?e i?i?iaeuei?e noia?i?e u?eueiino? (75,9 aec ia 10 e/ae?a)
ca?a?no?iaaii a ianaaeaeaii? ea?aaaie. Iaeneiaeueia e?euee?noue aeae?a
(23-27) i?eoiaeeoueny ia aea?aai? ianaaeaeaiiy, a /aaa?ieeiaeo
ianaaeaeaiiyo ?o e?euee?noue a 1,8 ?ac iaioa. ?iaeaen ??ciiiai?ooy
Oaiiiia ?ic?aoiaaiee aeey an?o aa??aio?a ianaaeaeaiue ii aeeiai?/i?e
u?eueiino? (2,16-2,9) aeuee ca ?iaeaen ??ciiiai?ooy, ?ic?aoiaaiee ii
a?iian? (1,99-2,1). Aeiii?o?/iee aiae?c noaio oaoie i?aenoeeee ia
ae?eyioe? ?aeoeueoeaaoe?? iieaco?, ui a oeaiioe/iiio a?aeiioaii?
ia?aaaaeathoue e?nia? aeaee — 41,5 %, noaiia? aeaee neeaaeathoue 38,5 %,
a eo/i? — 20 %. Aaeeea aeiey e?niaeo aeae?a ii??aiyii c noaiiaeie
na?ae/eoue i?i ni?eyoeea?noue e?ni?ineeiieo oiia ia ae?eyioe?. Iniiaiith
oaiaeaioe??th ?icaeoeo oa?oioaaiiy ? noaiiaeaiiy e?niaiai oeio
e?oaiia?ao ?a/iaei.

CI?IE NOAIO NO?OEOO?E AACO?AAAOIEO I?AeNOEEEE A CAIEAAIEO A?IAAIOeAIICAO
??*EE NAIA?E IA OA?EOI??? CAO?AeIIAI AeIIAANO, ssE? CIAOIAessOUeNss I?Ae
OAOIIAAIIEI IAAAIOAAEAIIssI

I?inaaei/i? yaeua i?e caeiaoaaii? eai’yiiai aoa?eey ? iiaeaeueoa
i?aeoiieaiiy caieaae ?.Naia?e i?ecaiaeeoue aei ciaioaiiy o 2 ?ace
e?eueeino? aeae?a aaco?aaaoieo i?aenoeeee. Neeaae aaco?aaaoieo
i?aenoeeee o i?aeoiieai?e caieaa? ?.Naia?e ia ??ai? oaenii?a iieacaii o
oaaeeoe? 1.

Aeey i?aeoiieaieo oa eiio?ieueieo ae?eyiie o?eueee 9 aeae?a aoee
ni?eueieie. ?iaeaen noiaeino? oa?oioaaiue aei??aith? 0,169, ui aeaco? ia
cia/io ci?io no?oeoo?e oaoie. Ca ?aooiie a?aenooiino? ia ae?eyieao
i?aeoiieaii? iieie aaeeeeo oo?oi?a, /a?a’ye?a a?aeaoaa?oueny ca?eueoaiiy
na?aaeiueiai cia/aiiy a?iiane oaa?ei ia eiio?ieueieo ae?eyieao (0,33
a/i2 i?ioe 0,096 a/i2.). I?aeoiieaiiy caieaaieo a?iaaioeaiic?a ia
oa?eoi??? Cao?aeiiai Aeiiaano i?ecaiaeeoue aei ciaioaiiy ?iaeaena
Oaiiiia, ?ic?aoiaaiiai ii aeeiai?/i?e u?eueiino? — 2,20 ii??aiyii c
3,09. Oeae ?iaeaen, ?ic?aoiaaiee ii a?iian?, a cii? i?aeoiieaiiy
noaiiaeoue 1,34, o eiio?ie? — 2,19. Ia ae?eyieao i?aeoiieaiiy e?nia?
aeaee neeaaeathoue (57 %) a?ae caaaeueii? /enaeueiino?, i?e aeineoue
aenieiio a?aenioeo eo/ieo aeae?a — (38 %), ui a?aeiia?aea? i?ioeano
ieoeia?iiy ? iiaeaeueoiio ia?aoiaeo eo/iiai a?iaaioeaiico o aieioiee.
Aeineoue aenieee ia ae?eyioe? i?aeoiieaiiy a?aenioie a?a?io?e?a — 19 %,
oa oeueo?aa?a?io?e?a — 5 %, na?aae yeeo iiaeia a?aecia/eoe oae? aeaee ye
Agonum obscurum, Stenolophus proximus oa ?io?, a oaeiae cacia/eoe aeaee
?iaeeeaoi?e canieaiiy — Anisodactilus binotatus, Dischirius sp. Ci?ie
noaio oa?oioaaiue aaco?aaaoieo oaa?ei aeathoue iiaeeea?noue
aeei?enoiaoaaoe ?o aeey ae?aaiinooaaiiy i?e?iaeieo e?niaeo
a?iaaioeaiic?a a oiiaao ?ioaineaiiai aeiaoaaiiy ei?enieo eiiaeei.

Oaaeeoey 1

Neeaae aaco?aaaoieo i?aenoeeee ia ae?eyioe? caieaae ?.Naia?a o cii?
i?aeoiieaiiy ia oa?eoi??? Cao?aeiiai Aeiiaano (ia ??ai? oaenii?a)

Oaenii Aaniethoia /enaeuei?noue,

aec/i2 Aeii?ioaaiiy, % Aeeiai?/ia u?euei?noue

10 e/ae Aeii?ioaaiiy, %

Aranei

Diplopoda

Chilopoda

Isopoda

Insenta

Carabidae

Staphylinidae

Silphidae

Elateridae

Histeridae

Scarabaeidae

Tenebrionidae 8,2

0,7

0,25

1,35

18,0

10,6

2,25

4,2

0,5

0,45

– 28,6

2,06

0,9

4,77

63,71

58,88

12,5

23,33

2,77

2,5

– 6,0

1,57

0,33

11,84

5,53

0,14

4,57

0,42

0,71

0,33

0,14 30,39

7,95

1,67

59,9

46,70

1,18

38,59

3,54

5,99

2,78

1,18

INIAEEAINO? AEOIOEssOe?? AAAEEEO IAOAE?A AACO?AAAOIEIE I?AeNOEEEE
I?E?IAeIEO OA OOO*IEO E?NIAEO A?IAAIOeAIIC?A

Iaeiie/aiiy aaaeeeo iaoae?a aaco?aaaoieie i?aenoeeee i?e?iaeieo e?niaeo
ianaaeaeaiue. C’yniaaii, ui a oiiaao i?iieneiaiai caa?oaeiaiiy
iaea?eueoa e?euee?noue cae?ca i?noeoueny a i?aai?ci? Isopoda oa
Lumbricidae — 5970 ia/ea oa 4210ia/ea a?aeiia?aeii, na?aae eiiao o
i?aaenoaaiee?a Silphidae — 3150 ia/ea oa Carabidae — 2890ia/ea.
Ia?aaiaoeue o a?eueo?e i??? iaeiie/othoue i?aaenoaaieee Lumbricidae —
362,0 ia/ea oa Aranei — 303,0 ia/ea, Cu — Lumbricidae — 201,0 ia/ea oa
Carabidae — 92,0 ia/ea, Zn — Isopoda — 1365,0 ia/ea oa Carabidae — 1280
ia/ea. Ii e?eueeino? iaeiie/aiiy Ni noi?oue ia i’yoiio i?noe? i?ney Fe,
Mn, Zn, Cu. Na?aae iaea?eueo oiene/ieo aeaiaio?a Pb o cii? caa?oaeiaiiy
o a?eueo?e i??? iaeiie/othoue i?aaenoaaieee Arane? — 45,0 ia/ea oa
Stapholinidae — 69,0 ia/ea, a Cd — Arane? — 96,0 ia/ea oa Carabidae —
29,0 ia/ea. Ai?no Pb o i?aaenoaaiee?a Carabidae, Stapholinidae,
Elateridae c ciie caa?oaeiaiiy o 3 ?ace a?eueoa ii??aiyii c
eiio?ieueieie ae?eyieaie. ?iaeaen a?ieia?/iiai iaeiie/aiiy (?AI), yeee
iieaco? iiaeeeaino? aeoioeyoe?? aaaeeeo iaoae?a aaco?aaaoieie,
iai?eeeaae nai?ioaaaie, o eaioethaao aeeaeaiiy (ca noano?ao aeeaeaiiy
nai?ioaa?a i?eeiyoi i?aenoeeeo) o i?e?iaeieo e?niaeo a?iaaioeaiicao aeey
Cd aeuee i?ae aeey Pb (oaaeeoey 2). A naith /a?ao ?iaeaen a?iiaeiie/aiiy
Cd o oaa?ei c caa?oaeaieo ae?eyiie o 3 ?ace aeuee, i?ae eiio?ieueieo.
O?aaa a?aecia/eoe, ui ?AI aeey an?o i?e?iaeaiaio?a i?aaenoaaieeaie
nai?ioaa?a c caa?oaeiaieo ae?eyiie aeuee ii??aiyii c eiio?ieueieie
ae?eyieaie.

Oaaeeoey 2

?iaeaen a?ieia?/iiai iaeiie/aiiy i?aaenoaaieeaie Lumbricidae oa Isopoda
a caieaaieo a?iaaioeaiicao ?.?. Naia?e oa Eiiiieyiee.

I?noea caeiae/? Oaenii ?iaeaen a?ieia?/iiai iaeiie/aiiy

Fe Mn Cu Zn Ni Pb Cd

Ciia ca-a?oaeiaiiy Isopoda 6,01 1,57 16,50 12,63 10,50 7,50 21,00

Eiio?ieue Isopoda 5,07 0,61 32,25 12,39 7,06 6,25 7,75

Ciia ca-a?oaeiaiiy Lumbricidae 4,21 5,58 20,12 2,96 4,28 5,00 6,00

Eiio?ieue Lumbricidae 3,34 1,13 15,75 3,04 3,46 2,75 2,00

Iaeiie/aiiy aaaeeeo iaoae?a aaco?aaaoieie i?aenoeeee c ooo/ieo e?niaeo
ianaaeaeaiue. A oiiaao noaiiaiai I?eaei?i?ia’y i?aeoe/ii a?ey eiaeiiai
i?aei?e?inoaa ?icoaoiaai? ooo/i? ianaaeaeaiiy. O cii? ae?? AeAI «Acio»
iaea?eueoa eiioeaio?aoe?y Fe aeyaeaia aeey ?aeiiiae?aieo — 4894 ia/ea oa
/a?a’ye?a — 3624 ia/ea, a na?aae eiiao aeey ia?oai?ae?a — 3254 ia/ea. O
cii? ae?? oeaiaioiiai ae?iaieoeoaa aeoeaii iaeiie/othoue Fe ?aeiiiae?ai?
— 4894 ia/ea oa iaaoeiiiae?ai? — 2450 ia/ea, a iaeiaioa eiioeaio?aoe?y
cae?ca ii??aiyii c ?ioeie i?e?iaeaiaioaie (0,075 ia/ea) a?aecia/aia aeey
/a?a’ye?a, ui iiaeeeai, iia’ycaii c? neeaaeii aeeeae?a oeaiaioiiai
ae?iaieoeoaa. Mn a cii? ae?? AeAI «Acio» o a?eueo?e i??? iaeiie/othoue
?aeiiiae?ai? — 1316 ia/ea, oa ia?ai?aee — 1326 ia/ea, a o cii? ae??
oeaiaioiiai ae?iaieoeoaa ?aeiiiae?ai? — 917 ia/ea oa /a?a’yee — 587
ia/ea. I?aeue ye o eiio?ieueieo, oae ? o caa?oaeiaieo ooo/ieo
ianaaeaeaiiyo a a?eueo?e i??? iaeiie/o?oueny ?aeiiiae?aieie — 379-408
ia/ea oa iaaoeiiiae?aieie — 140-156 ia/ea. Ia caaaeath/e ia aenieee
??aaiue iaeiie/aiiy Zn ieanoei/anoiaoneie o ianaaeaeaiiyo c ciie ae??
AeAI «Acio» eiai eiioeaio?aoe?y o 3 ?ace ieae/a ii??aiyii c ?aeiiii
oeaiaioiiai ae?iaieoeoaa — 826 ia/ea oa 2210 ia/ea a?aeiia?aeii.
Nai???aeii a?aeaoaa?oueny iaeiie/aiiy Ni o /a?a’ye?a. sseui o cii? ae??
aiie iaeiie/ththoue eiai 2 ia/ea, eiio?ieueieo ianaaeaeaiiyo 28 ia/ea,
oi o cii? ae?? AeAI «Acio» 280 ia/ea. Naeiaoeue o cii? oaoiiaaiiiai
caa?oaeiaiiy o a?eueo?e i??? iaeiie/othoue ?aeiiiae?ai? — 207-455ia/ea
oa noao?e?i?aee — 48-92 ia/ea, a Cd — ?aeiiiae?ai? — 43-171 ia/ea oa
iaaoeiiiae?ai? — 42-60 ia/ea. ?AI Fe aeey nai?ioaa?a (/a?a’yee,
?aeiiiae?ai?) c eiio?ieueieo oa caa?oaeiaieo ianaaeaeaiue ciaoiaeeoueny
ia ??ai? 4,9-6,75, aea aeey /a?a’ye?a c ciie ae?? oeaiaioiiai
ae?iaieoeoaa a?i noaiiaeoue 0,0001. Aeey Mn oa Ni a?i aeua o
ianaaeaeaiiyo caa?oaeiaieo i?iieneiaeie aeeeaeaie. Ia caa?oaeiaieo
ae?eyieao ?aeiiiae?ai? caeaoi? iaeiie/oaaoe a?eueoa Pb (?AI 103,0;
227,5) oa iaioa Cd (?AI 43,0; 21,0), a /a?a’yee iaaiaee a?eueoa Cd (?AI
6,0; 6,55) oa iaioa Pb (?AI 2,0; 4,0).

Aeei?enoaiiy iiaeaeueieo a?oi aaco?aaaoieo aeey i?iaaaeaiiy
aeioieneeieia?/ieo aeine?aeaeaiue. Aeey oaeaeei? ioe?iee aio?iiiaaiiiai
aieeao ia a?iaaioeaiic aeioe?eueii aeei?enoiaoaaoe iiaeaeuei? a?oie
aaco?aaaoieo oaa?ei, ye? caeaoi? iaeiie/oaaoe aaaee? iaoaee ca ei?ioeee
a?ae??cie /ano, ui aeicaiey? nai?/anii i?iaiaeeoe ?iaeeeaoe?th
caa?oaeiaiiy a?iaaioeaiico. A oiiaao noaiiaiai I?eaei?i?ia’y
aeei?enoiaoaaeenue i?aaenoaaieee oo?oi?a, ye? a?aeiia?aeathoue an?i
aeiiaai aei iiaeaeueieo a?oi. Aiie caeaoi? iaeiie/oaaoe aaaee? iaoaee a
aaeee?e e?eueeino?, caa??aath/e i?e oeueiio aenie? aaeaioeai?
iiaeeeaino?. Iaeiie/aiiy aaaeeeo iaoae?a ia ??ai? o?io?/ieo a?oi ia?
nai? iniaeeaino?. sseui o ooo/ieo ianaaeaeaiiyo i?ae aieeaii
i?iieneiaiai caa?oaeiaiiy i?enio?oioaae iaeiie/othoue o 4 ?ace a?eueoa
Fe, o 1,5-2 ?ace a?eueoa Zn, Pb, Cd ii??aiyii c ciioaaaie, a inoaii? o
1,5-2 ?ace a?eueoa Ni, oi o caieaaieo a?iaaioeaiicao oe? ci?ie ia
iano?eueee /?oei aecia/ai?. I?ae aieeaii i?iieneiaiai caa?oaeiaiiy
i?aaenoaaieee ciioaa?a iaeiie/othoue o 1,5 ?ace a?eueoa Pb, i?ae
i?enio?oioaae, a i?enio?oioaae o 1,2-2 ?ace a?eueoa Cd i?ae ciioaae. A
naith /a?ao eiioeaio?aoe?y aaaeeeo iaoae?a a i?aai?ci? oo?oi?a
aecia/a?oueny ?o aeaeiaeie iniaeeainoyie. Oae, Pterostichus melanarius
oa Poecilus cupreus o a?eueo?e i??? aeoioeththoue Mn oa Cu (342,8 ia/ea;
98,4 ia/ea a?aeiia?aeii), a Carabus granulatus — Ni (369,5 ia/ea), aea
an? aeaee oo?oi?a caeaoi? iaeiie/oaaoe aaeeeo e?euee?noue oaeeo
aenieioiene/ieo i?e?iaeaiaio?a ye Pb (29,1-37,5 ia/ea) oa Cd (23,1-82,7
ia/ea). Aeei?enoaiiy ?o aeey ?iaeeeaoe?? caa?oaeiaiiy aeiae?eey
aei?aaaeaii oei, ui aiie iiaeeiathoue o?eueee ?ooeea? oi?ie aeaiaio?a ?
oiio eiioeaio?aoe?y caa?oaeithaa/a o oaa?eiao a?aeaeaa? oaeoe/ia
caa?oaeiaiiy aeinenoai, a ia iioaioe?eia.

Caeaaei?noue oaoi?noe/ieo iieaciee?a a?ae a?io?i?/ieo ia?aiao??a oa
i?ia?aeueiiai iai?io o aaco?aaaoieo i?aenoeeee c a?iaaioeaiic?a,
i?aeaeaiieo i?iieneiaiio caa?oaeiaiith. Ia’?eoaie aeine?aeaeaiue aoee
i?aaenoaaieee iaaoeiiiae?aieo, /a?a’ye?a oa oo?oi?a c eiio?ieueieo oa
caa?oaeiaieo aeeeaeaie o?i?/ii? i?iieneiaino?, oeaiaioiiai ae?iaieoeoaa
ooo/ieo ianaaeaeaiue. Na?aae i?e?iaeaiaio?a ia?ai? Mn (ye a?iaaiiee), Pb
oa Cd (ye aenieioiene/i?). Iaeneiaeueia iaeiie/aiiy Mn (1120 ia/ea)
ca?a?no?iaaii aeey oo?oi?a c ianaaeaeaiue a cii? ae?? o?i?/ii?
i?iieneiaino?, i?e oeueiio ?o aeaeiaa ??ciiiai?ooy nei?i/o?oueny a 1,3
?ace, a a?iiana c?inoa? a 1,5 ?ace. Ai?no Pb oa Cd o oo?oi?a c
ianaaeaeaiue o cii? ae?? oeaiaioiiai ae?iaieoeoaa ia ??ai? 25 ia/ea oa
a?aeiia?aeii 19 ia/ea, ui a 1,3-1,8 ?ac?a aeua o ii??aiyii? c
eiio?ieueieie ianaaeaeaiiyie oa o 2,2 ?ace ieae/a ii??aiyii c ai?noii ?o
o aeoe?a c ciie ae?? o?i?/ii? i?iieneiaino?. Aeeeaee oeaiaioiiai
ae?iaieoeoaa i?ecaiaeyoue aei ca?eueoaiiy o 1,2 ?ace aeaeiaiai neeaaeo
aeoe?a oa o 5 ?ac?a /enaeueiino?, ui aeaco? ia ?oi? aenie? aaeaioaoe?ei?
iiaeeeaino?. Ii?yae c oeei o oo?oi?a c caa?oaeiaieo a?ioii?a
cieaeo?oueny e?euee?noue ni?eueieo e?i?ae?a a 1,3 ?ace i?e ca?eueoaii? a
?o neeaae? ??aiy a?eueieo aee?ieo eeneio a 6,8 ?ac oa oieanoa?eio a 1,2
?ace. O iaaoeiiiae?aieo iaeneiaeueia eiioeaio?aoe?y Mn (420 ia/ea)
a?aecia/aia o cii? ae?? AeAI «Acio». Oea o 1,3 ?ace aeua ii??aiyii c
eiio?ieai. Ai?no Pb oa Cd o oaa?ei c ciie ae?? i?iieneiaeo i?aei?e?inoa
o 2 ?ace aeua ii??aiyii c eiio?ieai. Ii?yae c oeei /enaeuei?noue iaaoe?a
o cii? ae?? oeaiaioiiai ae?iaieoeoaa o 1,6 ?ace, a i?aei?e?inoa o?i?/ii?
i?iieneiaino? o 1,2 ?ace iaioa ii??aiyii c eiio?ieai. A?iiana iaaoe?a o
cii? ae?? i?aei?e?inoa o?i?/ii? i?iieneiaino? ca?eueoo?oueny o 6 ?ac
ii??aiyii c eiio?ieai. O /a?a’ye?a iaeneiaeueia eiioeaio?aoe?y Mn (587
ia/ea) i?eoaiaiia oaa?eiai c ciie ae?? i?aei?e?inoa oeaiaioii?
i?iieneiaino?. Ai?no Pb o /a?a’ye?a c ciie ae?? i?aei?e?inoa o?i?/ii?
i?iieneiaino? o 4, a Cd o 1,5 ?ace aeua ii??aiyii c eiio?ieai. I?e
oeueiio e?euee?noue aeae?a /a?a’ye?a a cii? ae?? oeaiaioiiai
ae?iaieoeoaa a 2, a o?i?/ii? i?iieneiaino? a 3 ?ace a?eueoa ii??aiyii c
eiio?ieai.

ACA?IIIIA’ssCAI? CI?IE O ?INEEIIIIO IIE?EA? OA IANAEAII? AACO?AAAOIEO
I?AeNOEEEE A AEINENOAIAO, I?AeAeAIEO OAOIIAAIIIIO AIEEAO

Ci?ie ?ineeiiiai iie?eao oa oaa?eiiiai ianaeaiiy i?aenoeeee caieaaieo
a?iaaioeaiic?a ?.Naia?a, i?aeaeaieo oaoiiaaii?e ae??, aiane?aeie
aeiaoaaiiy eai’yiiai aoa?eey. Oaooi? i?ae?iaee oa iia’ycai? c ieie
i?inaaei/i? yaeua, ye? i?ecaiaeyoue aei i?aeoiieaiiy caieaaieo
a?iaaioeaiic?a ??/ee Naia?e ia oa?eoi??? Cao?aeiiai Aeiiaano, iaaaoeaii
aieeaathoue ia o?eueee ia aea?aaino?e oa o?aa’yiee iie?ea, aea e ia
i?aaenoaaiee?a aaco?aaaoieo oaa?ei i?aenoeeee. Ci?ia aii?oaoo
aea?aainoith i?ae aieeaii i?aeoiieaiiy i?ecaiaeeoue aei ci?i ?ineeiiiai
iie?eao oa iaeeaaea? a?aeaeoie ia oi?ioaaiiy i?aenoeeee — na?aaeiaeua
iaoeaiiy i?aenoeei/ieo oaa?ei. A?aenooi?noue /?oei ae?aaeaii? e?niai?
i?aenoeeee aieeaa? ia oiiae iaoeaiiy oa neeaae aaco?aaaoieo oaa?ei. Ia
ae?eyieao a cii? oaoiiaaiiiai aieeao a 2,5 ?ace ciaioo?oueny e?euee?noue
aeae?a aaco?aaaoieo ii??aiyii c eiio?ieueieie a?iaaioeaiicaie, oa a 2-3
?ace ?o /enaeuei?noue. A cii? i?aeoiieaiiy cieeathoue i?aaenoaaieee
oeiia? aeey eiio?ieueii? caieaae — Calosoma inquisitor, Carabus
marginalis, Philonthus laevicolis oa ?io?. sse?ni? oa e?euee?ni? ci?ie
aaco?aaaoieo i?aenoeeee a?aeaoaathoueny ia o?eueee ia aeaeiaiio ??ai?,
aea e ia ??ai? oaenii?a — cieeathoue i?aaenoaaieee Trombiculidae,
Lumbricidae. Oaa?eie i?aaenoaaeai? a?eueo ae??ai?oeie oi?iaie, ye?
iathoue ieae/o a?iiano ii??aiyii c i?aaenoaaieeaie eiio?ieueieo
a?iaaioeaiic?a, aea iathoue aeu? ?ai?iaeoeoeai? cae?aiino?, e?aua
aeaeeaathoue o iani?eyoeeaeo oiiaao. Oaoiiaaiia iaaaioaaeaiiy aieeaa? ia
aeiii?o?/i? oa?aeoa?enoeee ?ineei oa oaa?ei. sse ? o o?aa’yienoeo ?ineei
a cii? i?aeoiieaiiy o aaco?aaaoieo oaa?ei aeineoue aenieee a?aenioie
eo/ieo aeae?a — 38 % ii??aiyii c eiio?ieueieie ae?eyieaie — 25 %,
noaiia? aeaee i?aaenoaaeai? ie?aieie aecaiiey?aie. sse na?aae ?ineeiiiai
iie?eao, oae ? na?aae aaco?aaaoieo i?aenoeeee, ciaeaeaii i?aaenoaaiee?a
aaeia?iio?a, ye? iiaeooue aeei?enoiaoaaoenue ye ?iaeeeaoi?e canieaiiy
a?oioo.

Ii?oieia?/i? ci?ie ?ineeiiiai iie?eao oa oaa?eiiiai ianaeaiiy
i?aenoeeee, ye a?aeiia?aeue ia caa?oaeiaiiy iaaeieeoiueiai na?aaeiaeua
i?iieneiaeie aeeeaeaie. Aeeeaee acioiiai ae?iaieoeoaa aieeaathoue ia
?ai?iaeoeoeai? cae?aiino? aea?aaii-/aaa?ieeiaeo ?ineei, aeeeeeathoue o
ieo ii?oieia?/i? ci?ie (Noiey?aieiaa, Eoeooeei, 1988). Caaaeai? aaoi?e
i?iaiaeeee aeine?aeaeaiiy o ooo/ieo e?niaeo ianaaeaeaiiyo o cii? ae??
AeAI «Acio», oiio iaea?aeai? aeai? aoee aeei?enoai? aeey aeyaeaiiy
noi?nieo ci?i ?ineeiiiai iie?eao oa oaa?eiiiai ianaeaiiy i?aenoeeee.
Aaoi?e a?aecia/athoue iaaaoeaiee aieea caa?oaeiaiiy ia ii?oieia?/i?
iniaeeaino? nooea?oue, cieaeaiiy e?eueeino? aooii?a oa ea?o?a,
e?eueeino? oa iane ian?iiy aea?aa. Ii?oieia?/i? ci?ie o aaco?aaaoieo
i?aenoeeee i?ae aieeaii i?iieneiaiai caa?oaeiaiiy iaie
aeine?aeaeoaaeenue ia i?eeeaae? Carabus granulatus, Pterostichus
melanarius, Staphylinus caesareus. Aei??ai i?aeeyaaee aieiaa,
ia?aaeiueinieiea, iaaee?eea, aoneee, e?ioe?aee. O?eueee o Pterostichus
melanarius aeinoia??ii ca?eueoo?oueny aeiaaeeia aonee?a ii??aiyii c
eiio?ieueieie ae?eyieaie. ?ioeo aeinoia??ieo ii?oieia?/ieo ci?i o
i?aaenoaaiee?a aaco?aaaoieo ia ca?a?no?iaaii. Oaeei /eiii ii?oieia?/i?
ci?ie ye o ?ineei oae ? aaco?aaaoieo ? aaeaeaaoiei a?aeaoeii aeeaeo
i?aai?ci?a ia i?iieneiaa caa?oaeiaiiy.

Iniaeeaino? iaeiie/aiiy ?ia??ae??io?a i?iieneiaiai ae?iaieoeoaa
?ineeiiei iie?eaii oa aaco?aaaoieie i?aenoeeee. I?iaaaeaii
aeine?aeaeaiiy ii aeyaeaiith iniaeeainoae iaeiie/aiiy aaaeeeo iaoae?a a
nenoai? a?oio — i?aenoeeea — ?ineeia — aaco?aaaoia oaa?eia. Aeineoue
neeueia oaoiiaaiia iaaaioaaeaiiy a oiiaao i?iieneiaiai caa?oaeiaiiy
cacia? iaeei c iaeaaaeeea?oeo no?oeoo?ieo eiiiiiaio?a a?iaaioeaiico —
e?niaa i?aenoeeea. Ia ae?eyieao a?ey i?iieneiaeo eiiieaen?a
ca?eueoo?oueny ai?no Cu o 2 ?ace, Zn o 1,6 ?ace, Ni o 1,3 ii??aiyii c
eiio?ieai. Ai?no Pb a i?aenoeeoe? c ciie i?iieneiaiai caa?oaeiaiiy
noaiiaeoue 2,4-2,7 ia/ea i?ioe 1,5ia/ea o eiio?ie?, a Cd 1,4-2,2 ia/ea
i?ioe 0,61 ia/ea o eiio?ie?. Iaeiie/aiiy i?e?iaeaiaio?a o a?oio? o cii?
i?iieneiaiai caa?oaeiaiiy aeaui ieae/a ii??aiyii c i?aenoeeeith. A
?acoeueoao? aiae?co iaeiie/aiiy i?e?iaeaiaio?a o i?aenoeeoe? oa a?oio?
iiaeia c?iaeoe i?aenoiie, ui i?aenoeeea aenooia? aeoeaiei a?io?eueo?ii ?
aeeiio? caoeni? ooieoe?? ia oeyoo i?iieeiaiiy aaaeeeo iaoae?a o a?oio.
Nooi?iue aeoioeyoe?? ?ineeiaie i?e?iaeaiaio?a (ia i?eeeaae? i’yoi/ieea)
iiaeia ioe?ieoe ii eiao?oe??ioo a?ieia?/iiai iaeiie/aiiy (EAI). A?i
na?ae/eoue, ui o cii? i?iieneiaiai caa?oaeiaiiy eenoy o?aa’yienoeo
?ineei o a?eueo?e i??? iaeiie/othoue Cu (EAI Cu 8,6), a o iaio?e Cd (EAI
Cd 0,12). Ei??iiy o a?eueo?e i??? iaeiie/othoue Ni (EAI Ni 5,5), a o
iaio?e Cd (EAI Cd 0,1). O i?aaenoaaiee?a aaco?aaaoieo i?aenoeeee
(?aeiiiae?ai?) o cii? i?iieneiaiai caa?oaeiaiiy iaea?eueoee aenieee ?AI
aeey oaeiai aenieioiene/iiai aeaiaioo ye Pb (?AI 227,5), a iaea?eueo
iecueeee aeey Fe (?AI 4,7). Eiiieaeni? aeine?aeaeaiiy i?aaenoaaiee?a
oaa?eiiiai oa ?ineeiiiai na?oo aeathoue iiaeeea?noue iaoe aeieeaaei?o?
?acoeueoaoe i?i nooi?iue caa?oaeiaiiy iaaeieeoiueiai na?aaeiaeua ??cieie
/eiieeaie aio?ii?/iiai iioiaeaeaiiy.

IIAEEEA?NOUe AEEI?ENOAIIss AACO?AAAOIEO I?AeNOEEEE AeEss A?I?IAeEEAOe??
NOAIO ?O IANAEAIIss OA E?NIAEO IANAAeAEAIUe

Iaea?eueo oi/ieie neiaeieia?/ieie oa?aeoa?enoeeaie aeey a?i?iaeeeaoe??
aenooia? aeaeiaa ?icia?ooy (?iaeaen Oaiiiia, N?iiniia oa ?io?),
aaaaonoai ianaeaiiy, no?oeoo?a aeii?ioaaiiy. Iaio iaae?eiei aeey
aeei?enoaiiy ? oaeee iieaciee, ye iayai?noue /e a?aenooi?noue
i?aaenoaaiee?a aaeeeeo nenoaiaoe/ieo a?oi. Ne?ae a?aoiaoaaoe
aaeaioaoe?th aaaaoueio aeae?a aei oiia ?nioaaiiy ia caa?oaeiaieo
ae?eyieao. ?acoeueoaoe ii??aiyiiy ?iaeeeaoe?eieo oa?aeoa?enoee
aaco?aaaoieo i?aenoeeee ooo/ieo ianaaeaeaiue c ciie ae?? AeAI «Acio» oa
oeaiaioiiai ae?iaieoeoaa iieacothoue, ui ni?eueiei aeey ieo ? cieaeaiiy
?iaeaena a?ieia?/ii? ??ciiiai?oiino? a 1,1-1,2 ?ace, a?aenooi?noue o
oaoi? aeeieiiiae, ca?eueoaiiy e?eueeino? aeae?a noao?e?i?ae, ca?eueoaiiy
/enaeueiino? nai?ioaa?a o 3-4 ?ace oa cieaeaiiy /enaeueiino? o?oioaa?a o
3-7 ?ac?a. Na?aae oaeoeueoaoeaieo a?i?iaeeeaoi?ieo oa?aeoa?enoee iiaeia
aeei?enoaoe aeii?iaioio no?oeoo?o aeiuiaeo /a?a’ye?a. Ia ae?eyieao,
caa?oaeiaieo aeeeaeaie AeAI «Acio» c’yaeythoueny Dendrobaena octaedra,
Octolasium complonatum, Lumbricus rubellus, o e?ninioaao a cii? ae??
oeaiaioiiai ae?iaieoeoaa Eisenia nordenscioldi, a ia iaio ae?eyieao
Eisenia foetida. Neeaaei?oa c’ynoaaoe a?i?iaeeeaoe?ei? oa?aeoa?enoeee
aeey aaco?aaaoieo i?aenoeeee caieaaieo e?niaeo a?iaaioeaiic?a. Aeaye?
iieacieee ni?euei?, a aeaye? i?ioeeaaei?. Oae ?iaeaen a?ioe/iiai
??ciiiai?ooy cieaeo?oueny a 1,4 ?ace. Cieaeaiiy e?eueeino? aeae?a oaoie
iiaeia aeei?enoaoe ye aeiaeaoeiaee iieaciee. No?oeoo?a aeii?ioaaiiy
aeae?a aaco?aaaoieo i?aenoeeee eiio?ieueieo oa caa?oaeiaieo caieaaieo
a?iaaioeaiic?a aeineoue niaoeeo?/ia ? iiaea aooe aeei?enoaia a
a?i?iaeeeaoe?eieo oe?eyo. O?eueee ia caa?oaeiaieo ae?eyieao na?aae
aeiuiaeo /a?a’ye?a c’yaey?oueny Eisenia nordenscioldi, na?aae ?aaeee?a
Vallonia costata, na?aae oo?oi?a Pterostichus anthracinus, Pterostichus
nigrita, Harpalus amplicollis, na?aae ieanoei/aoiaoneo Valgus
hemipterus.

AENIIAEE OA ?AEIIAIAeAOe??

I?iieneia? caa?oaeiaiiy a ooo/ieo oa i?e?iaeieo e?niaeo ianaaeaeaiiyo
yaeythoueny i?e/eiith cieaeaiiy a ieo aeaeiai? ??ciiiai?oiino?
aaco?aaaoieo i?aenoeeee, ca?eueoaiiy /enaeueiino? aeae?a oa iia’ycaia c
oeei ca?eueoaiiy e?eueeino? noia?aeiieiaio?a, ui aeaco? ia ii?ooaiiy a
eiiieaenao i?aenoeeeiaeo aaco?aaaoieo. Aeeeaee i?iieneiaeo i?aei?e?inoa
ni?eythoue ciaioaiith ?iaeaen?a ??ciiiai?oiino?, nei?i/aiith e?eueeino?
oaenii?a ca ?aooiie cieeiaiiy i?aaenoaaiee?a ie?aieo ?iaeei. Ia ??ai?
ooieoe?iiaeueieo a?oi a?aecia/aii cieaeaiiy /enaeueiino? i?aaenoaaiee?a
o?oioaa?a oa ca?eueoaiiy nai?ioaa?a, i?e aenie?e /enaeueiino? ciioaa?a.

I?aeoiieaiiy caieaaieo a?iaaioeaiic?a ia oa?eoi??? Cao?aeiiai Aeiiaano
aaaea aei cieaeaiiy ?iaeaena ??ciiiai?oiino?, ?ic?aoiaaiiai ii
/enaeueiino? c 3,09 aei 2,02 oa a?iian? c 2,19 aei 1,34 ii??aiyii c
aoaeiiieie a?iaaioeaiicaie. Iecueea /enaeuei?noue aaco?aaaoieo i?e
iecuee?e aeaeia?e ??ciiiai?oiino? ia ae?eyieao caieaae oa ciie
i?aeoiieaiiy na?ae/aoue, ui oaoia aaco?aaaoieo ciaoiaeeoueny a neeueii
i?eai?/aiiiio noai?. Aeiii?o?/iee aiae?c oaoie aena?oeth? c?inoaiiy
/anoee iacia?a?io?eueieo eiiieaen?a ? oiio iniaeeai i?eaeaoiee aeey
a?i?iaeeeaoe?? ci?i a i?e?iaeieo caieaaieo a?iaaioeaiicao i?ae
oaoiiaaiiei aieeaii.

Aenieee ai?no aaaeeeo iaoae?a a i?aai?ci? aaco?aaaoieo oaa?ei c ooo/ieo
oa i?e?iaeieo e?niaeo ianaaeaeaiue, ye? ciaoiaeyoueny i?ae aieeaii
i?iieneiaeo aeeeae?a, aeaco? ia i?aeaeuaiee ??aaiue caa?oaeiaiiy oeueiai
?aeiio oa aeicaiey? aeei?enoiaoaaoe oeeo oaa?ei ye a?i?iaeeeaoi??a
caa?oaeiaiiy o iii?oi?eiaiaeo aeine?aeaeaiiyo iaaeieeoiueiai
na?aaeiaeua.

Aeiaaaeaii, ui a yeino? iiaeaeueii? a?oie i?e i?iaaaeaii?
aeioieneeieia?/ieo aeine?aeaeaiue a oiiaao noaiiaiai I?eaei?i?ia’y
iiaeia aeei?enoiaoaaoe oo?oi?a (Carabidae). Iath/e aenieo /enaeuei?noue,
aeaeiaa ??ciiiai?ooy, oe?iea ?iciianthae-aeaiiy, aiie iaeiie/othoue
aaeeeo e?euee?noue i?e?iaeaiaio?a, iiaeeiath/e o?eueee ?ooeea? oi?ie, a
oea cia/eoue, ui eiioeaio?aoe?y caa?oaeithaa/a a i?aai?ci? oaa?ei aoaea
a?aeia?aaeaoe oaeoe/io caa?oaeiaii?noue aeinenoai, a ia iioaioe?eio.

??ci? a?oie aaco?aaaoieo oaa?ei, iaeiie/oth/e a i?aai?ci? aaeeeo
e?euee?noue aaaeeeo iaoae?a, ?acii c oei iathoue ??cio nooi?iue
aaeaioaoe?? aei aeaea?ae ?o i?iieneiai? ai?n??. Iaeiie/aiiy
aaco?aaaoieie i?e?iaeaiaio?a, ? a oiio /ene? oaeeo aenieioiene/ieo ye
naeiaoeue oa eaaei?e, aieeaa? ia ?o aeaeiaee neeaae, /enaeuei?noue oa
a?iiano.

Ia ia?oiio aa??aio? a?oio?a ae?eyiee e?niai? ?aeoeueoeaaoe??, aea
aei?noe? e?ni?ineeii? oiiae i?ecaiaeyoue aei aeiaaeaiiy aea?aainoith,
aaco?aaaoi? ia ciaoiaeyoue ni?eyoeeaeo oiia aeey aeeooy ? oaoia
i?aaenoaaeaia aeiaaeeiaeie i?aaenoaaieeaie. Ia ae?oaiio aa??aio?
a?oio?a, ia aeeaey/enue ia oa, ui aenioa aea?aa, c?ieioo?noue ?o e?ii,
oa?aeoa? caieiaeaiiy aeaeae? a?ae iioeiaeueieo aeey e?no, /enaeuei?noue
i?aaenoaaiee?a i?aenoeeeiai? oaoie c?inoa? o 5 ?ac?a ii??aiyii c ia?oei
aa??aioii, aea ia noa? iaeneiaeueiith. Ia o?aoueiio-i’yoiio aa??aioao
a?oio?a o ca’yceo c ca?eueoaiiyi oaeoi??a neeueaaoecaoe??, /enaeuei?noue
oa aeaeiaa ??ciiiai?ooy oaoie aaco?aaaoieo aeinyaa? naiai iaeneiaeueiiai
cia/aiiy – 131 – 196,5 aec. ia 10 e/ae?a.

Ia ?iciiae?e oa no?oeoo?o ianaeaiiy oaa?ei ia ae?eyioe? e?niai?
?aeoeueoeaaoe?? aaeeeee aieea iathoue e?ni?ineeii? oiiae, iniaeeai
aeieia?/ia no?oeoo?a aea?aai-noith. A aea?aaieo ianaaeaeaiiyo, oaeeo ye
ianeeiea aocueeieenoa, a’yc iecueeee, aeaoe?y a?ea, aaco?aaaoi?
i?aenoeeee ciaoiaeyoue aeey naaa a?eueo ni?eyoeea? oiiae ?nioaaiiy
ii??aiyii c /aaa?ieeiaeie ianaaeaeaiiyie. Ia aeeaey/enue ia iecueeo
ye?nio ni?euei?noue i?aenoeeeiai? oaoie a caeaaeiino? a?ae neeaaeo
aea?aaii-/aaa?ieeiaeo ianaaeaeaiue iaeneiaeueii aeecuee? ianaeaiiy
aaco?aaaoieo i?aenoeeee aeaoe?? a?ei?, a’yca iecueeiai, ianeeiee
aocueeieenoi?. No?oeoo?a oaoie aaco?aaaoieo aeaco? ia oa, ui ia
ae?eyioe? e?niai? ?aeoeueoeaaoe?? a?aeaoaa?oueny noaiiaeaiiy
a?iaaioeaiico, a?ieia?/iee e?oaiia?a yeiai ? i?ii?aeiei i?ae e?niaei oa
noaiiaei.

Iaieio/? ci?ie, aiane?aeie ae?iaie/i? ae?yeueiino? ethaeeie, iniaeeai ia
nai?o ?aii?o noaae?yo, ia caaaeaee iiaeooue aooe nai?/anii cao?eniaai?.
A oeeo oiiaao ci?ie ii?oieia?/ieo oa?aeoa?enoee, aeaeiaiai neeaaeo,
iniaeeaino? iaeiie/aiiy i?iien-eiaeo aeeeae?a ?ineeiaie oa aaco?aaaoieie
i?aenoeeee iiaeooue aooe iaae?eiei e?eoa-???i ioe?iee oeo aeano?oeoeaieo
i?ioean?a, ye? a?aeaoaathoueny a aeinenoaiao i?ae aieeaii ??cieo oi?i
caa?oaeiaiiy.

Aeey a?i?iaeeeaoe?? caa?oaeiaiiy iacaiieo a?iaaioeaiic?a iiaeia
?aeiiaiaeoaaoe oae? iniaeeaino? no?oeoo?e aaco?aaaoieo: cieaeaiiy
?iaeaeno a?ieia?/iiai ??ciiiai?ooy, ci?ie no?oeoo?e aeii?ioaaiiy ia
??ai? ?iaeei oa aeae?a, ci?ie ni?aa?aeiioaiiy o?io?/ieo oa?oioaaiue.
?io? iieacieee iiaeooue aeei?enoiaoaaoenue aeiaeaoeiai a ie?aieo
aeiaaeeao.

Iaea?aeai? ?acoeueoaoe aeicaieythoue ?aeiiaiaeoaaoe iii?oi?eiaia?
aeine?aeaeaiiy ??cieo ca noaiii o?ainoi?iaoe?? aeinenoai i?iaiaeeoe ia
??ai? oa?oiiaaiue, oa aeiiiaithaaoe ?o ia aeaeiaiio, iiioeyoe?eiiio oa
?ioeo ??aiyo. Iaaeith e?eoe/iiai noaio ianaeaiiy aaco?aaaoieo i?aenoeeee
ne?ae aaaaeaoe ?iciiae?e iniaei ia ??ai? aeaeo, iine?eueee a?i i?aeoe/ii
ia a?aeiiaey?oueny.

Nienie iioae?eiaaieo i?aoeue

Naciiiua, aiaeiaua e aucaaiiua aio?iiiaaiiuie oaeoi?aie eciaiaiey
no?oeoo?u iiioeyoeee ii/aaiiuo e iacaiiuo aaniicaiii/iuo aeeaioiuo a
iaeioi?uo aeiaaioeaiicao oeaio?aeueiiai noaiiiai I?eaeiai?iauey /Aa?nia
A.A., Ieeeiaiei A.O., AEoeia A.A., Eoeueaa/ei TH.E., Eenaiei O.E. //
Aanoiee Aeiai?iiao?ianeiai oieaa?neoaoa. — Aeiai?iiao?iane: AeAO, 1995,
aui. 2. — N. 177-184.

?ac?aaioea iaoiaea eiiieaeniie yeieiai-aeioeie/aneie ioeaiee ninoiyiey
iiioeyoeee aeeaioiuo a oaoiiaaiiuo yeinenoaiao I?eaeiai?iauey / Ienth?a
A.I., AEoeia A.A., Iieic I.A., Nooaiiaa A.I., Aanni A.ss., ?aaa A.A.,
Eoeueaa/ei TH.E. // Aanoiee AeAO. — Aeiai?iiao?iane: AeAO, 1996, aui. 2.
— N. 177-185.

Eoeueaa/ei TH.E. Aaniicaiii/iua iiaenoeeee eae eiaeeeaoi? ninoiyiey
iieiaiiuo aeiaaioeaiicia Caiaaeiiai Aeiiaanna, iaoiaeyueony iiae
oaoiiaaiiie iaa?oceie // Ieoaiiy noaiiaiai e?niciaanoaa oa e?niai?
?aeoeueoeaaoe?? caiaeue. I?aeaoc. ca. iaoe. i?. — Aei?i?iiao?ianuee:
AeAeO, 1998, ei. 2. — N. 174-179.

Eoeueaa/ei TH.E. Iniaaiiinoe iaeiieaiey oyaeaeuo iaoaeeia aeoaeaeeoeaie
a aeiaaioeaiicao, iiaeaa?aeaiiuo oeie/aneiio caa?yciaieth // A?niee
Aei?i?iiao?ianueeiai oi?aa?neoaoo. A?ieia?y. Aeieia?y. —
Aei?i?iiao?ianeue: AeAeO, 1998, aei. 4. — N. 63-68.

Eoeueaa/ei TH.E. Aaniicaiii/iua aeeaioiua iiaenoeeee eneonnoaaiiuo
aaeiaeaoeeaauo ianaaeaeaiee eae aeieiaeeeaoi?u caa?yciaiey ie?oaeathuae
n?aaeu aua?inaie i?iiuoeaiiiai i?iecaiaenoaa // I?eaei?i?ianueeee
iaoeiaee a?niee. A?ieia?y, n?euenueea ainiiaea?noai oa aaoa?aia??y. —
1998. — ?113 (180). — N. 121-125.

Ciiaaia?aoe/aneee aiaeec ii/aaiiie iiaenoeei/iie oaoiu noaiiiai
I?eaeiai?iauey / AEoeia A.A., Ieeeiaiei A.O., Aa?nia A.A., Nie?iia
TH.A., Eoeueaa/ei TH.E., Eenaiei O.E. // Aii?inu noaiiiai eaniaaaeaiey e
eaniie ?aeoeueoeaaoeee caiaeue. Iaaeaoc. na. iao/i. o?. —
Aeiai?iiao?iane: AeAeO, 1997. — N. 89-92.

N?aaieoaeueiay oa?aeoa?enoeea niaea?aeaiey iaeioi?uo iee?iyeaiaioia a
i?aaiecia aaniicaiii/iuo, caiiiaiaeiuo e i?aniueathueony a oneiaeyo
oaoiiaaiiiai aeeyiey /Ienth?a A.I., Aanni A.ss., AEoeia A.A., Eoeueaa/ei
TH.E., ?oceia A.E., Ia?/aieianeay A.A. // A?niee Aei?i?iiao?ianueeiai
oi?aa?neoaoo. A?ieia?y. Aeieia?y, aei. 3, 1997. — N. 133-143.

Eaaeano?iaay oa?aeoa?enoeea ianaeaiey aaniicaiii/iuo aeeaioiuo iniiaiuo
aeiaaioeaiioe/aneeo eaoai noaiiie ciiu Oe?aeiu (i?eaeieeiii-aaei/iue
eaiaeoaoo) /Aa?nia A.A., Ei?aaeaa A.I., Ieeeiaiei A.O., Eoeueaa/ei
TH.E., Nie?iia TH.A. // Aii?inu noaiiiai eaniaaaeaiey e eaniie
?aeoeueoeaaoeee caiaeue. Iaaeaoc. na. iao/i. o?. — Aeiai?iiao?iane:
AeAO, 1997. — N. 132-142.

Anoanoaaiiay e aio?iiiaaiiay aeeiaieea no?oeoo?u ianaeaiey ii/aaiiuo e
iacaiiuo aaniicaiii/iuo aeeaioiuo a iaeioi?uo aeiaaioeaiicao
oeaio?aeueiiai noaiiiai I?eaeiai?iauey / Aa?nia A.A., AEoeia A.A.,
Eoeueaa/ei TH.E., Eenaiei O.E. //Onoie/eaia ?acaeoea: caa?yciaiea
ie?oaeathuae n?aaeu e yeieiae/aneay aaciianiinoue. Oac. aeiee. 1
Iaaeaeoia?. iao/i.-i?aeo. eiio. — Aeiai?iiao?iane, 1995. — O. 1. — N.
78-79.

Eoeueaa/ei TH.E. Eiiieaenu aaniicaiii/iuo, eae iieacaoaeue ninoiyiey
ae?aaaniuo ianaaeaeaiee a oneiaeyo noaie // Aanoiee Aeiai?iiao?ianeiai
oieaa?neoaoa. Aeieiaey. Yeieiaey. — Aeiai?iiao?iane: AeAO, 1993, aui. 1.
— N. 61.

I?iaeaiu io?aiu yioiiioaoiu eaiaeoaooia Aeiai?iiao?iaueiu, iaoiaeyueony
iiae oa?icie en/aciiaaiey / Aa?nia A.A., Eenaiei O.E., Eoeueaa/ei TH.E.,
AEoeia A.A. //4-a I?aeia?iae. eiio. O?aioe?y oa Oe?a?ia,
iaoeiai-i?aeoe/iee aeina?ae o eiioaeno? ae?aeiao iaoe?iiaeueieo
eoeueoo?. — Aei?i?iiao?ianuee, 1997. — O. 2. — N. 6-7.

Eoeueaa/ei TH.E. I?i cia/aiiy ?ioi?iaoe?? c aioiiieia?/ii?
a?i?iaeeeaoe?? i?iieneiaeo caa?oaeiaiue aeey ioi?iie iaaeieeoiueiai
na?aaeiaeua ? oi?ioaaiiy aeieia?/ii? eoeueoo?e // 4-a I?aeia?iae. eiio.
O?aioe?y oa Oe?a?ia, iaoeiai-i?aeoe/iee aeina?ae o eiioaeno? ae?aeiao
iaoe?iiaeueieo eoeueoo?. — Aei?i?iiao?ianuee, 1997. — O. 2. — N. 43.

Nie?iia TH.A., Eoeueaa/ei TH.E., AEoeia I.A. Aaco?aaaoi? oaa?eie a
aeieia?/iiio iii?oi?eiao iaaeieeoiueiai na?aaeiaeua // 2 I?aeia?. eiio.
O?aioe?y oa Oe?a?ia: iaoeiai-i?aeoe/iee aeina?ae. — Aei?i?iiao?ianuee,
1995. — N. 62-63.

Eoeueaa/ei TH.E., *oi?eia I.A. Aaniicaiii/iua iiaenoeeee eae
aeieiaeeeaoi?u oaoiiaaiiiai caa?yciaiey ie?oaeathuae n?aaeu // Yeieiaey
e iieiaeaaeue. Iaoa?eaeu 1 Iaaeaeoia?. iao/i.-i?ae. eiio. — Aiiaeue,
1998. — O. 1. — N. 105.

Eoeueaa/ei TH.E., Aa?nia A.I., E?naiei O.?. Aieea aaaeeeo iaoae?a ia
no?oeoo?o aeayeeo eiiieaen?a aaco?aaaoieo c i?aenoeeie ooo/ieo e?niaeo
ianaaeaeaiue // Caa?aaeaiiy a?i??ciiiai?oiino? a Oe?a?i?. Iaoe?iiaeueia
eiio. — Eai?a, 1997. — N.40.

Eoeueaa/ei TH.E. Aeeyiea i?iiuoeaiiiai i?iecaiaenoaa ia iaeioi?ua a?oiiu
aaniicaiii/iuo iiaenoeeee eneonnoaaiiuo aaeiaeaoeeaauo ianaaeaeaiee // 5
I?aeia?. eiio. «O?aioe?y oa Oe?a?ia, iaoeiai-i?aeoe/iee aeina?ae o
eiioaeno? ae?aeiao iaoe?iiaeueieo eoeueoo?». — Aei?i?iiao?ianuee, 1998.
— O. 2. — N. 8-9.

Eoeueaa/ei TH.E. AEoaeaeeoeu eae aeieiaeeeaoi?u caa?yciaiey ii/au
oyaeaeuie iaoaeeaie a oneiaeyo noaiiiai I?eaeiai?iauey // 1-y
Iaaeaeoia?. iao/i.-i?ae. eiio. — Aeiai?iiao?iane, 1995. — O. 2. — N.
76-77.

Kulbachko Y.L. Influence of cement productiion on some biochemical
indices of ground Beetles (Carabidae) // Sustainable development: system
analysis in ecology. 2 nd Practical Conference Sevastopol. Ukraine,
1996. — P. 142-143.

Aanni A.ss., Eoeueaa/ei TH.E., Ienth?a A.I. Eciaiaiea iaeioi?uo
aeioeie/aneeo iieacaoaeae iicaiii/iuo e aaniicaiii/iuo aeeaioiuo a
oneiaeyo i?iiuoeaiiiai caa?yciaiey n?aaeu iaeoaiey // Anaoe?a?inueea
iaoeiai-iaoiaee/ia eiioa?aioe?y «Aeieia?y oa ?iaeaia??y. Noai,
iane?aeee, oeyoe ooai?aiiy aeieia?/ii /enoeo oaoiieia?e». —
Aei?i?iaeca?aeeinuee, 1996. — N. 41-42.

Eoeueaa/ei TH.E. Ninoiyiea no?oeoo?iie i?aaiecaoeee aaniicaiii/iuo
aeeaioiuo iiaenoeeee noaiiuo eania a oneiaeyo i?iiuoeaiiiai caa?yciaiey:
?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo iaoe
ii niaoeeaeueiinoe 03.00.16. — «Yeieiaey». — Aeiai?iiao?ianeee
ainoaea?noaaiiue oieaa?neoao, Aeiai?iiao?iane, 1999.

Aeenna?oaoeey iinayuaia aiciiaeiinoe eniieueciaaiey aeiea
i?aaenoaaeoaeae aaniicaiii/iuo iiaenoeeee aeey aeieiaeeeaoeeiiiuo
enneaaeiaaiee, ioeaiee ninoiyiey anoanoaaiiuo e eneonnoaaiiuo eaniuo
aeiaaioeaiicia noaiiiai I?eaeiai?iauey a oneiaeyo aeeyiey ?acee/iuo
aio?iiiaaiiuo oaeoi?ia.

Iieacaii, /oi i?iiuoeaiiia caa?yciaiea i?eaiaeeo e ea/anoaaiiui e
eiee/anoaaiiui eciaiaieyi a eiiieaenao aaniicaiii/iuo iiaenoeeee
anoanoaaiiuo e eneonnoaaiiuo eaniuo aeiaaioeaiicia. Oiaiueoaaony eiaeaen
aeaeiaiai ?aciiia?acey, a ia o?iaia ooieoeeiiaeueiuo a?oii oiaiueoaaony
/eneaiiinoue i?aaenoaaeoaeae oeoioaaia, oaaee/eaaaony /eneaiiinoue
nai?ioaaia i?e aeiaieueii aunieie /eneaiiinoe ciioaaia.

?anniio?aiu iniaaiiinoe oi?ie?iaaiey oaoiu aaniicaiii/iuo iiaenoeeee
eneonnoaaiiuo yaeaoioiiia ia ianoa oaooiuo ioaaeia. A oi?ie?iaaiee e
?ani?aaeaeaiee ii/aaiii-ciieiae/aneeo eiiieaenia aieueoia cia/aiea eiaao
oei ae?aainoiy, iniaaiii aai no?oeoo?a e ninoaa. Ia o/anoea eaniie
?aeoeueoeaaoeee /eneaiiinoue e aeaeiaia ?aciiia?acea iiaenoeei/iuo
aaniicaiii/iuo aic?anoaao io eonoa?ieeiauo e ae?aaaniui ianaaeaeaieyi.
ssae?i oaoiu aaniicaiii/iuo ia iyoe aa?eaioao eneonnoaaiiuo ii/aia?oioia
ninoaaeytho i?aaenoaaeoaee ianaeiiuo, ca n/ao eioi?uo eciaiyaony
aeeiaie/aneay ieioiinoue e o?iaaiue aeiieie?iaaiey yieaaeiuo
a?oiie?iaie. Eiaeaen aeaeiaiai ?aciiia?acey aeey /eneaiiinoe iniaae
ii/oe aaeaia auoa /ai aeey eo aeiiannu. A oeaiioe/aneii ioiioaiee ia
o/anoea eaniie ?aeoeueoeaaoeee i?aiaeaaeatho eaniua aeaeu, /eneaiiinoue
noaiiuo aeaeia a 1,2 ?aca ieaea, a /eneaiiinoue eoaiauo a 2 ?aca ieaea
ii n?aaiaieth n eaniuie e noaiiuie. Aieueoay aeiey eaniuo aeaeia
naeaeaoaeuenoaoao i aeaaii?eyoiuo eani?anoeoaeueiuo oneiaee ia o/anoea.
Iieacaoaee eciaiaiey ninoiyiey a?oiie?iaie aaniicaiii/iuo iiaenoeeee a
oneiaeyo iiaeoiieaiey i?e i?inaaei/iuo i?ioeannao a iieiao ?ae iiaeii
eniieueciaaoue aeey aeeaaiinoeee anoanoaaiiuo eaniuo aeiaaioeaiicia a
oneiaeyo eioaineaiie aeiau/e eaiaiiiai oaey.

Enneaaeiaaiu iniaaiiinoe iaeiieaiey oyaeaeuo iaoaeeia ?acee/iuie
a?oiiaie aaniicaiii/iuo iiaenoeeee a anoanoaaiiuo e eneonnoaaiiuo eaniuo
ianaaeaeaieyo a oneiaeyo i?iiuoeaiiiai caa?yciaiey. Ioia/aii nieaeaiea
eiaeaena aeieiae/aneiai iaeiieaiey iee?iyeaiaioia i?aaenoaaeoaeyie
nai?ioaaia a ciia i?iiuoeaiiuo aua?inia. Aianoa n oai i?aaenoaaeoaee
?aeiia?aciuo (iie?eoeu) niiniaiuo iaeaieeaaoue a aieueoae ia?a naeiaoe e
a iaiueoae eaaeiee, a /a?ae iaiai?io a aieueoae ia?a eaaeiee e a
iaiueoae naeiaoe. Iainiiaaia aiciiaeiinoue a ciia i?iiuoeaiiiai
caa?yciaiey a oneiaeyo noaiiiai I?eaeiai?iauey a ea/anoaa iiaeaeueiie
a?oiiu aeey i?iaaaeaiey yeioieneeieiae/aneeo enneaaeiaaiee
eniieueciaaoue aeoaeaeeoe.

Iieacaia caaeneiinoue oaoienoe/aneeo iieacaoaeae io aeioeie/aneeo
ia?aiao?ia e ieia?aeueiiai iaiaia o ?acee/iuo i?aaenoaaeoaeae
aaniicaiii/iuo iiaenoeeee. A /anoiinoe, o aeoaeaeeoe ioia/aii nieaeaiea
aeaeiaiai ?aciiia?acey, oaaee/aiea aeiiannu i?e nieaeaiee eiee/anoaa
iaueo eeieaeia e oaaee/aiey a eo ninoaaa naiaiaeiuo aee?iuo eeneio e
oieanoa?eia.

Aia?aua ?anniio?aiu acaeiinaycaiiua eciaiaiey ii?oieiae/aneeo
oa?aeoa?enoee, iniaaiiinoe iaeiieaiey iee?iyeaiaioia ?anoeoaeueiui
iie?iaii e ianaeaieai aaniicaiii/iuo iiaenoeeee iiae aeeyieai
i?iiuoeaiiiai caa?yciaiey a oneiaeyo noaiiiai I?eaeiai?iauey. Ioia/aiu
neiyeieiae/aneea oa?aeoa?enoeee aaniicaiii/iuo iiaenoeeee, eioi?ua
iiaeii eniieueciaaoue a aeieiaeeeaoeeiiiuo oeaeyo.

Eeth/aaua neiaa: oaoiiaaiiia caa?yciaiea, oyaeaeua iaoaeeu,
oaeniiiie/aneay no?oeoo?a, aeieiaeeeaoeey.

Eoeueaa/ei TH.E. Noai no?oeoo?ii? i?aai?caoe?? aaco?aaaoieo oaa?ei
i?aenoeeee noaiiaeo e?n?a a oiiaao i?iieneiaiai caa?oaeiaiiy: ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
ca niaoe?aeuei?noth 03.00.16 — «Aeieia?y». — Aei?i?iiao?ianueeee
aea?aeaaiee oi?aa?neoao, Aei?i?iiao?ianuee, 1999.

Aeena?oaoe?y i?enay/aia ieoaiiyi aeei?enoaiiy ??cieo a?oi ? aeae?a
aaco?aaaoieo i?aenoeeee aeey a?i?iaeeeaoe?? oa ioe?iee noaio e?niaeo
a?iaaioeaiic?a noaiiaiai I?eaei?i?ia’y a oiiaao oeneo ??cieo oi?i
aio?iiiaaiieo /eiiee?a. C’yniaaii ci?ie neiaeieia?/ieo iieaciee?a,
aeaeiaiai neeaaeo oa oaeniiii?/ii? no?oeoo?e i?aenoeeeiaeo aaco?aaaoieo
i?ae aieeaii i?iieneiaiai caa?oaeiaiiy. ?icaeyiooi iniaeeaino?
oi?ioaaiiy oaoie aaco?aaaoieo i?aenoeeee ia ae?eyieao e?niai?
?aeoeueoeaaoe??. Aecia/aii iniaeeaino? iaeiie/aiiy aaaeeeo iaoae?a
??cieie a?oiaie aaco?aaaoieo i?aenoeeee, noi?ni? ci?ie ?ineeiiiai
iie?eao oa oaoie aaco?aaaoieo i?aenoeeee ia oaoi?noe/iiio,
ii?oieia?/iiio, aeioieneeieia?/iiio ??aiyo n iiaeaeueoei aeei?enoaiiyi
?o aeey a?i?iaeeeaoe?eieo aeine?aeaeaiue. ?acoeueoaoe ?iaioe ciaeoee
ai?iaaaeaeaiiy a i?e?iaeiioi?iiieo aeieia?/ieo caoiaeao, iaa/aeueiiio
i?ioean?.

Eeth/ia? neiaa: oaoiiaaiia caa?oaeiaiiy, aaaee? iaoaee, oaeniiii?/ia
no?oeoo?a, a?i?iaeeeaoe?y.

Kulbachko Yu.L. The state of structural organization of invertebrates in
a litter of steppe forests under conditions of industrial pollution. —
Manuscript.

The dissertation to obtain the degree of Candidate of Science (Biology)
in speciality 03.00.16 — Ecology. — Dniepropetrovsk State University,
Dniepropetrovsk, 1999.

The thesis is devoted to the problems of different invertebrate groups
and species of soil-cover utilization for bioindication and evalution of
steppe Pridneprovye forest biogeocoenosis state in the conditions of
antropogenous factors effect. Speceis composition alterations of
synecological indicies and taxonomy structure of soil-cover
invertebrates under the influence of industrial pollution have been
determined. Deculiarities of soil-cover invertebrates fauna formation on
the lots of wood recultivation have been considered. Peculiarities of
heavy metals accumulation by different groups of soil-cover
invertebrates of plant cover and soil-cover invertebrates fauna
interconnected changes on the fauna, morphology, ecology level with
further their utilization for bioindication investigations have been
determined. Our investigations resulted in environment protection
measures and training course.

Key-words: technogenous pollution, heavy metals, taxonomy structure,
bioindication.

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