?INOEOOO CAIEA?IANOAA

OE?A?INUeEA AEAAeAI?ss AA?A?IEO IAOE

?OAAI ?IIA IEAEN??AIA

OAeE 633.15:631.52

NAEAEOe?ss A?A?EAe?A EOEO?OAeCE IA NO?EE?NOUe AeI OAI?IA A OIIAAO
E?NINOAIO OE?A?IE

06.01.05 – naeaeoe?y ? ian?iieoeoai

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

Ee?a — 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia a Oiainuee?e n?euenueeiainiiaea?nuee?e aeaaeai??.

Iaoeiaee ea??aiee: aeieoi? n?euenueeiainiiaea?nueeeo iaoe, i?ioani?

*o/i?e ?aai Iao?iae/, caa?aeoaa/ eaoaae?e

aaiaoeee, naeaeoe?? oa ian?iieoeoaa,

Oiainueea n?euenueeiainiiaea?nueea aeaaeai?y.

Io?oe?ei? iiiiaioe: aeieoi? n?euenueeiainiiaea?nueeeo iaoe, i?ioani?

ssoianueeee ?aai A?eoi?iae/, aieiaiee iaoeiaee

ni?a?ia?oiee a?aeae?eo naeaeoe?? e?oi’yieo eoeueoo?

?inoeoooo caiea?ianoaa OAAI,

eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe

IA?OIIAIEI Aiaoie?e Einoyioeiiae/,

aeioeaio eaoaae?e naeaeoe?? oa o?c?ieia?? ?ineei,

Iaoe?iiaeueiee aa?a?iee oi?aa?neoao.

I?ia?aeia onoaiiaa: A?iieoeueeee aea?aeaaiee n?euenueeiainiiaea?nueeee
?inoeooo.

Caoeno a?aeaoaeaoueny “20”aeiaoiy 1999 ?. ia can?aeaii?

Niaoe?ae?ciaaii? a/aii? ?aaee Ae 27.361.01 i?e ?inoeooo? caiea?ianoaa
OAAI

ca aae?anith:

255205, nio. *aaaie Ee?ai-Nayoioeinueeiai ?eiio Ee?anueei? iaeano?,

?inoeooo caiea?ianoaa OAAI, ia ?i’y a/aiiai nae?aoa?y Niaoe?aaee.

C aeena?oaoe??th iiaeia iciaeiieoeny o a?ae?ioaoe? ?inoeoooo.

A?aeceae ia aaoi?aoa?ao o aeaio i?ei??ieeao, caa??aieo ia/aoeith,

i?ineii iaaeneeaoe ia ?i’y a/aiiai nae?aoa?y Niaoe?aaee.

Aaoi?aoa?ao ?ic?neaii “17 ” aa?aniy 1999 ?.

A/aiee nae?aoa?

Niaoe?ae?ciaaii? a/aii? ?aaee,

eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe
E?aa/aiei E.I.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. A no/anieo aeiiii?/ieo oiiaao ca?eueoaiiy
ae?iaieoeoaa ca?ia ? ei?i?a a cia/i?e i??? caeaaeeoue a?ae oni?o?a
naeaeoe??. Iaeiei ?c aenieiaoaeoeaieo oeyo?a ae??oaiiy i?iaeaie
ca?eueoaiiy ae?iaieoeoaa ca?ia eoeo?oaece oa iie?ioaiiy eiai yeino? ?
noai?aiiy iiaeo aenieia?iaeaeieo, no?eeeo aei iaea?eueo oeiaei/eiieo
oai?ia a?a?eae?a eoeo?oaece. Na?oiaee aeina?ae iieaco?, ui naeaeoe?y ia
i?iaeoeoeai?noue ? ye?noue i?iaeoeoe?? aac iaeii/aniiai i?aeaeuaiiy
no?eeino? aei oai?ia iaoiiaeth? aenieo aaiaoe/io a?aceea?noue iiaeo
ni?o?a oa a?a?eae?a.

Ca?ac o ae?iaieoeoa? ae?iuothoueny aenieia?iaeaei? aaoa?iceni?
a?a?eaee, aea aeey ?o noai?aiiy aeei?enoiao?oueny iaaaeeea e?euee?noue
e?i?e. Oiio i?aeaeuaiiy o?iaeaeiino? ca ?aooiie ?icoe?aiiy aaiaoe/ii?
iniiae a?a?eae?a ? iaei??th c iniiaieo caaea/ naeaeoe??. Aaaeeeaei
caniaii noaa?e?caoe?? aenieeo o?iaea?a ? naeaeoe?y ia ?ioi?oao aei
oai?ia . Aeey noai?aiiy aenieii?iaeoeoeaieo a?a?eae?a eoeo?oaece c
oe?iieie aa?iiii?/ieie iciaeaie, no?eeeo i?ioe iniiaieo oai?ia
iaiao?aeia iayai?noue aea/aiiai ca an?ia iciaeaie, a oiio /ene? ?
?ioiieia?/ieie aeanoeainoyie, aeo?aeiiai iaoa??aeo.

Ana oea iaoiiaeei aea?? oaie aeena?oaoe?eii? ?iaioe ? aecia/eei
iniiai? iai?yiee iaoeo aeine?aeaeaiue.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie.
Iaoeia? ?ic?iaee ca oaiith aeena?oaoe?? aoee neeaaeiaith /anoeiith
aeine?aeaeaiue eaai?aoi??? naeaeoe?? ? ian?iieoeoaa eoeo?oaece i?e
eaoaae?? aaiaoeee, naeaeoe?? oa ian?iieoeoaa Oiainueei?
n?euenueeiainiiaea?nueei? aeaaeai?? – ? aeeiioaaeenue ca?aeii c
aea?aeaaiith iaoeiai-oaoi?/iith i?ia?aiith “I?iaeiaieuenoai-95”, i?iaeo
“Ca?ii” ? oaiaoe/iiai ieaio eaai?aoi??? : “Naeaeoe?y ? ian?iieoeoai
aenieia?iaeaeieo a?a?eae?a eoeo?oaece aeey oiia E?ninoaio ? Iie?nny
Oe?a?ie” (iiia? aea?aeaaii? ?a?no?aoe?? 0194V038349).

Iaoa ? caaea/? aeine?aeaeaiue. Aieiaiith iaoith ?iaioe aoei
aea/aiiy ?ioiieia?/ieo oa?aeoa?enoee naiicaieeaieo e?i?e eoeo?oaece ?
caeiiii??iinoae oniaaeeoaaiiy no?eeino? aei iniiaieo oai?ia a?a?eaeaie
F1 a oiiaao E?ninoaio Oe?a?ie.

A?aeiia?aeii c iinoaaeaiith iaoith ae??ooaaeenue ne?aeoth/?
caaea/?:

— aea/eoe ia aeaio oiiao (aacci?iiee iin?a ? a oiiaao n?aici?ie)
anth ?iai/o eieaeoe?th e?i?e ? a?a?eae?a ca no?ee?noth aei iooe?/anoi? ?
eaoth/i? naaeee, noaaeiaeo aieeae oa ooca??ico ea/ai?a;

— i?ae?a?aoe eiio?anoi? ca no?ee?noth aei eiaeii? oai?iae e?i?? ?
i?iaanoe ae?aeaeuei? no?auoaaiiy ii /ioe?ueio noaiao;

— aecia/eoe no?ee?noue aei cacia/aieo oai?ia o e?i?e ? a?a?eae?a o
o?ueio ioieoao aei?iaoaaiiy (iaa/aeueii-aeine?aeia ainiiaea?noai
“?iaeiee?aea” Oiainueei? NAA, naeaeoe?eia iiea aeaaeai??, Iane?anueeee
?aaeaini-oaoi?eoi Ee?anueei? iaeano?);

— aeine?aeeoe oa?aeoa? oniaaeeoaaiiy no?eeino? aei oai?ia i?e
naeaeoe?? aaoa?icenieo a?a?eae?a;

— aeyaeoe ei?aeyoe?ei? aca?iica’ycee i?ae iniiaieie
ainiiaea?nueei-a?ieia?/ieie iciaeaie ? no?ee?noth aei oai?ia oa
eeaneo?eoaaoe aeo?aeiee iaoa??ae c iaoith iiaeaeueoiai eiai
naeaeoe?eiiai aeei?enoaiiy;

— aeynieoe oeiaei/eii?noue oai?ia ia e?i?yo ? a?a?eaeao a oiiaao
n?aici?ie, i?iaieaoe?eiiai oiio oa o ca’yceo c iioeiaeaeaiiyi ?ineei
oe?aeieeaie;

— ?aeiiaiaeoaaoe aeey i?aeoe/ii? naeaeoe?? e?i?? ? a?a?eaee ye
aeaea?aea no?eeino? aei iniiaieo oai?ia.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Aia?oa a oiiaao
E?ninoaio Oe?a?ie ia ??cieo oiiao i?iaaaeaii eiiieaenia aea/aiiy
no?eeino? naiicaieeaieo e?i?e, a o?i /ene? iiaeo e?i?e a?o/eciyii?
naeaeoe?? oa a?a?eae?a eoeo?oaece aei iniiaieo oai?ia (iooe?/anoa ?
eaoth/a naaeea, noaaeia? aiee? ? ooca??ic ea/ai?a ). Aeyaeai?
ei?aeyoe?ei? ca’ycee i?ae no?ee?noth aei oai?ia ? oe?iieie
ainiiaea?nueei-a?ieia?/ieie iciaeaie ?ineei. Aeae?eai? e?i?? c
eiiieaeniith no?ee?noth aei iniiaieo oai?ia. Aeine?aeaeaii oa?aeoa?
oniaaeeoaaiiy no?eeino? e?i?e aei oai?ia a a?a?eaeao F1- oea
eiaeii?ioaaiiy. ?aeiiaiaeiaai? aeaea?aea no?eeino? aei ie?aieo oai?ia, a
oaeiae e?i??, ye? ii?aeiothoue aenieo CEC ca no?ee?noth aei oai?ia ?
o?iaea?i ca?ia. Aecia/aia oeiaei/eii?noue iniiaieo oai?ia a oiiaao
E?ninoaio Oe?a?ie.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. Aeey i?aeoe/ii?
naeaeoe?? iiaeo a?a?eae?a eoeo?oaece i?iiiio?oueny aeei?enoaoe : aea/ai?
iaoiaee ioe?iee aeo?aeiiai iaoa??aeo ca no?ee?noth aei iniiaieo oai?ia,
ei?aeyoe?ei? ca’ycee i?ae ie?aieie oai?iaaie ? aaaeeeaeie
ainiiaea?nueei-a?ieia?/ieie iciaeaie, caeiiii??iino? oniaaeeoaaiiy
no?eeino? e?i?e aei oai?ia a a?a?eaeao F1, e?i?? ye aeaea?aea no?eeino?
aei iooe?/anoi? naaeee (*E 915, Ni151, Ni 125), eaoth/i? naaeee (I346cI,
OeA 10cI, F 115), noaaeiaeo aieeae (I 346cI, I502, *E 929, F2),
ooca??ico ea/ai?a (Oo 719, *E 218IA, Ia 21).

Iniaenoee aianie caeiaoaa/a. Caeiaoaa/ iniaenoi a?aa o/anoue a
ieaioaaii? oa caeeaaeaii? iieueiaeo aeine?ae?a, i?iaaaeaii?
?ioiieia?/ieo ioe?iie oa aiae?c?a, a ia?iaoe? iaoa??ae?a ? i?aeaioiaoe?
?o aei ae?oeo. Aaoi?ii cae?eniaii aiae?c ? ocaaaeueiaiiy e?oa?aoo?ieo
aeaieo ca oaiith aeena?oaoe?? oa iaea?aeaieo aenia?eiaioaeueieo aeaieo.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iniiai? ?acoeueoaoe
aeine?ae?a aeiiia?aeaeenue ia iaoeiaeo ?aaeao aa?iiii?/iiai oaeoeueoaoo
Oiainueei? n?euenueeiainiiaea?nueei? aeaaeai?? (Oiaiue, 1996-1998 ??.),
iaoeiai-iaoiaee/ieo eiioa?aioe?yo iieiaeeo a/aieo ? niaoe?ae?no?a,
(?inoeooo caiea?ianoaa OAAI, 1999 ??e), ia eiioa?aioe?? “No/ani?
i?iaeaie ?ineeiieoeoaa ? ei?iiae?iaieoeoaa “ (Oiaiue, 1998),
iaoeiai-i?aeoe/i?e eiioa?aioe?? ONAA (Oiaiue 1998??e).

Ioae?eaoe??. Ca iaoa??aeaie aeine?aeaeaiue iioae?eiaaii 5
iaoeiaeo i?aoeue, o yeeo i?aaenoaaeai? iniiai? iieiaeaiiy aeena?oaoe?? .

Ianya ? no?oeoo?a aeena?oaoe??. Aeena?oaoe?eia ?iaioa aeeeaaeaia
ia 130 noi??ieao iaoeiiieniiai oaenoo, aeeth/a? anooi, 5 ?icae?e?a,
aeniiaee, ?aeiiaiaeaoe?? aeey naeaeoe?eii? i?aeoeee, nienie aeei?enoaieo
aeaea?ae ? aeiaeaoee. A iniiai?e /anoei? ai?uaii 34 oaaeeoe?, 9
?enoie?a.

Nienie aeei?enoaieo aeaea?ae aeeth/a? 211 iaeiaioaaiue, c yeeo
10 ?iicaiiith iiaith.

Ci?no ?iaioe

NO*aniee noai naeaeoe?? eoeo?oaece ia no?ee?noue aei oai?ia

(iaeyae e?oa?aoo?e)

Ca?aeii c e?oa?aoo?ieie aeaieie iieacaii, ui ia oeae /an aeineoue
aeaoaeueii aea/ai? ieoaiiy iaoiaaiaco aeine?aeaeoaaieo oai?ia ,
a?ieia?/i? iniaeeaino? caoaeiee?a, ca’ycie no?eeino? aei iniiaieo oai?ia
c ie?aieie ii?oieia?/ieie iciaeaie ? caeiiii??iino? oniaaeeoaaiiy
no?eeino? a?a?eaeaie F1. Oi/ ?aeeii? aeoiee , iniaeeai a ieoaiiyo
aaiaoe/ieo iaoai?ci?a oniaaeeoaaiiy no?eeino? aei aeaieo oai?ia ia
?nio?. Ia aeaeue, a?eueo?noue ae?iuoaaieo a?a?eae?a iathoue ua
iaaeinoaoith no?ee?noue aei oai?ia. Oiio iaiao?aei?noue noai?aiiy
no?eeeo aei iaea?eueo oeiaei/eiieo oai?ia a?a?eae?a aeey oiia E?ninoaio
Oe?a?ie ? iniiaith aeey iaiieaaeeaeo aeine?aeaeaiue a oeueiio iai?yieo.

IAOA??AE, OIIAE, IAOIAeEEA AeINE?AeAEAIUe

Aenia?eiaioaeuei? aeine?aeaeaiiy i?iaaaeai? c 1996 ii 1998 ??e a
o?ueio ioieoao: AeI IAeA “?iaeiee?aea” Oiainueei? NAA, Iane?anueeee
?aaeaini-oaoi?eoi Ee?anueei? iaeano? ? ia naeaeoe?eiiio iie? Oiainueei?
NAA.

Iiaiaei? oiiae ca ?iee aea/aiiy a?ae??ciyeenue cia/iith
i?ieea?noth, iniaeeai ia aoaiao iioiaaiaco eoeo?oaece, ui aeicaieeei
iaea?aeaoe ia’?eoeai? aeniiaee. Ni?eyoeea? oiiae aeey ?icaeoeo
caoaeiee?a noaaeiaeo aieeae neeaeenue a 1996 ?ioe?, a aeey iooe?/anoi?
naaeee a 1998 ?ioe?.

Aeo?aeiei iaoa??aeii aeey i?iaaaeaiiy aeine?aeaeaiue aoee 24
ae?oi? naiicaieeai? e?i?? eoeo?oaece ??ciiai iioiaeaeaiiy c eiio?anoiith
no?ee?noth aei iniiaieo oai?ia. No?auoaaiiyi aeaieo e?i?e ca /ioe?ia
ae?aeaeueieie noaiaie aoei iaea?aeaii i?ino? a?a?eaee, ye? aea/aeenue ca
no?ee?noth aei iaea?eueo oeiaei/eiieo oai?ia a oiiaao i?e?iaeiiai ?
i?iaieaoe?eiiai oii?a. Ia no?ee?noue aei iooe?/anoi? ? eaoth/i? naaeee,
noaaeiaeo aieeae oa ooca??ico ea/ai?a aea/aeenue e?i?? ? a?a?eaee ia
ae?eyieao a?a?eaeecaoe?? ? ?iciiiaeaiiy. Anueiai ioe?iaii 145 iiaeo ?
?aeiiiaaieo a?a?eae?a oa 188 naiicaieeaieo e?i?e.

Iin?a i?iaiaeeee a?o/io ai?caeiaei niiniaii (70o35ni) ii o?e
ian?ieie a ai?caei, a iioeiaeuei? aeey ciie no?iee. Ae?eyiee iaeii- oa
aeai?yaeeia? aeiaaeeiith 7,2i, 19 – ai?caei?, iae?eiaa ieiua 5-10i2.
Iiaoi?i?noue a aeine?aeao o?e?aciaa .

Ioe?ieo c?ace?a ia no?ee?noue aei iooe?/anoi? ? eaoth/i? naaeee,
ooca??ico ea/ai?a i?iaiaeeee ca iaoiaeeeaie O.?. Iaie??iea,
A.A.A?enaiea, ?.?.N?aeaiea (1972) ? I.A.Aeeeiai? , A.A.?aauaiea oa ?i.
(1989), noaaeiaeo aieeae – I.A.Iaa?ioeueei?, I.A.?iaeeea (1977).

?ioiieia?/io oa?aeoa?enoeeo c?ace?a eoeo?oaece ?iaeee ca
?acoeueoaoaie o?ueio??/ieo ioe?iie ca iaeneiaeueiei a?aenioeii
o?aaeaiiy.

Ainiiaea?nuee?, a?ieia?/i? oa ii?oieia?/i? iciaee eoeo?oaece
aea/aee ca iaoiaeeeith ?inoeoooo ?ineeiieoeoaa OAAI.

Aiae?c caaaeueii? oa niaoeeo?/ii? eiia?iaoe?eii? caeaoiino?
e?i?e ca no?ee?noth aei iooe?/anoi? ? eaoth/i? naaeee, noaaeiaeo aieeae
oa ooca??ico ea/ai?a a nenoai? ae?aeaeueieo no?auoaaiue aeeiiaiee ca
iaoiaeeeith A??oo?iaa (iiaeaeue 1, iaoiae 4) a iiaeeo?eaoe??
A.A.Aieueoa, I.I.E?ooia oa ?i. (1980). Ioe?ieo i?iyao iciaee no?eeino?
aei iooe?/anoi? naaeee a F1 i?iaiaeeee ca aeiiiiiaith eiao?oe??ioa
nooiaiy aeii?iaioiino? (hp) ( Ae.A?thaaeea?, 1966).

Iaoaiaoe/io ia?iaeo aeaieo aenia?eiaioaeueieo aeine?aeaeaiue
i?iaiaeeee ca iaoiaeaie aeenia?n?eiiai, ei?aeyoe?eiiai oa ?aa?an?eiiai
aiae?c?a , aeeeaaeaieo A.I.Aeini?oiaei (1985) c aeei?enoaiiyi
ia?niiaeueiiai eiii’thoa?a Pentium 166.

?ACOEUeOAOE AEA*AIIss E?I?E ?IAI*I? EIEAEOe?? CA NO?EE?NOTH AeI INIIAIEO
OAI?IA

Ia i?e?iaeiiio oa i?iaieaoe?eiiio ?ioaeoe?eieo oiiao aea/aii 24
ae?oi? naiicaieeai? e?i??. Aea/aiiy aeo?aeiiai iaai?o naiicaieeaieo
e?i?e ca no?ee?noth aei iniiaieo oai?ia aeicaieeei aeae?eeoe ?yae
no?eeeo e?i?e aei naaeeiaeo oai?ia: *E915, Ni 125, An 073IA, Ni 151, Ia
21, I346cI, F 115, I 502, Oo 719, *E4282, *E 218IA, An 28A, *E 929;
noaaeiaeo aieeae : An 28A, *E 915, I 346cI, I 502, *E929,F 2 oa aei
ooca??ico ea/ai?a : S 72, *E 915, I 346cI, F 115, I 502, *E 929, Ia 21,
Oo 719.

Iniaeeai oe?iieie aeey naeaeoe?? ia a?oiiaee ?ioi?oao aei oai?ia
aeyaeeenue ne?aeoth/? e?i?? :*E 915, *E 929, I346cI, I502 (oaae. 1).
E?i?? Ia 21, F115 oa Oo 719 i?iyaeee aenieo no?ee?noue aei naaeeiaeo
oai?ia ? ooca??ico ea/ai?a oa iii??io no?ee?noue aei noaaeiaeo aieeae.
A?oiiao no?ee?noue aei naaeeiaeo oai?ia ? noaaeiaeo aieeae i?iyaeee
e?i?? An 28A oa F2. Iaea?eueo ni?eeiyoeeaeie aei oai?ia aeyaeeenue e?i??
A?? 44 oa A 246.

A i?ioean? aea/aiiy an? e?i?? eeaneo?eiaaii ca no?ee?noth aei
iniiaieo oai?ia. A?aei?/aii, ui o e?i?e, ye? aa?eoee a a?oio na?aaeiuei
oa neeueiio?aaeoaaieo no?ee?noue neeueii aa??thaaea ii ?ieao ? ioieoao
aei?iaoaaiiy, oiae? ye o no?eeeo e?i?e oey iciaea aoea a?eueo
noaa?eueiith.

A i?e?iaeieo oiiaao ca?aaeaiiy c 145 ioe?iaieo iiaeo ? ?aeiiiaaieo
a?a?eae?a aeiey oeo, ye? o?aaeoaaeenue iooe?/anoith naaeeith neeaaeaea
8,3%. Na?aae ianoaaeaieo 188 naiicaieeaieo e?i?e a?aenioie o?aaeaieo
noaiiaea 2,6-7,2. Eaoth/ith naaeeith aoea o?aaeaia o?eueee e?i?y I502 a
iacia/i?e

i???. O?aaeaiiy e?i?e ? a?a?eae?a noaaeiaeie aieeyie neeueii aa??thaaei
ii ?ieao.

Aea/aany ca’ycie no?eeino? e?i?e ? a?a?eae?a aei iniiaieo oai?ia
? c ii?oieia?/ieie iciaeaie oa a?ieia?/ieie aeanoeainoyie. Aeyaeaii
na?aaei?e cai?ioi?e ei?aeyoe?eiee ca’ycie i?ae nooiaiai o?aaeaiiy
iooe?/anoith naaeeith ? aenioith i?ee??ieaiiy ea/aia (r = -0,67) oa
neaaeee cai?ioi?e ca’ycie i?ae o?aaeaiiyi noaaeiaeie aieeyie ? ianith
1000 ca?ai(r = -0,39). I?ae ?ioeie ii?oieia?/ieie iciaeaie ? no?ee?noth
aei oai?ia ?noioieo ca’yce?a ia aeyaeaii.

Anoaiiaeaii, ui no?ee?noue eoeo?oaece aei an?o aea/aieo oai?ia
caeaaeeoue a?ae aaiioeiiaeo iniaeeainoae e?i?e, aeieia?/ieo oiia,
no?ie?a n?aae ? ?o nei?inoeaeino?. A?eueoo no?ee?noue aei oai?ia
i?iyaeee oi?ie c o?eaae?oei aaaaoaoe?eiei ia??iaeii.

Oaaeeoey 1

?ioiieia?/ia oa?aeoa?enoeea e?i?e eoeo?oaece ca no?ee?noth aei iniiaieo
oai?ia, 1996-1998 ??.

Iacaa e?i?e O?aaeai?noue, %

Iooe?/anoa naaeea Eaoth/a naaeea Noaaeia? aiee? Ooca??ic ea/ai?a

Ni 125 0,5 0 41,4 5,7

An 073 IA 1,0 0 41,6 7,7

Ii 236 9,8 0 37,8 14,4

Ni 151 1,9 0 15,8 13,4

S 72 3,3 0 14,3 4,3

AeAa 42 3,9 0 11,3 10,8

An 28 A 0,2 0 0,6 10,3

*E 915 0 0 0 0,9

I346cI 0,4 0 0 3,2

OeA 10cI 3,1 0 34,3 11,9

*E 209 1,9 24,1 14,7 10,3

A 246 6,9 13,7 8,7 13,4

E?I 052 6,0 16,6 46,5 8,0

F 115 1,6 0 9,1 0,9

F 2 1,6 0 1,6 9,3

THA 24 5,4 0 13,1 5,2

O 23 3,2 0 68,2 19,6

I502 1,6 0 0,7 1,5

*E 929 0,5 0 0 1,3

*E 218IA 1,0 0 42,2 8,1

A?? 44 9,2 11,6 59,2 5,4

*E 4282 0,6 0 23,6 5,3

OO 719 0,7 0 11,1 3,2

Ia 21 0,8 0 12,2 2,0

ONIAAeEOAAIIss NO?EEINO? E?I?E AeI OAI?IA A A?A?EAeAO F1

Oniaaeeoaaiiy no?eeino? e?i?e aei naaeeiaeo oai?ia

Uia iiai?oa aeei?enoiaoaaoe iioaioe?ei? iiaeeeaino? eoeo?oaece
iaiao?aeii i?aeaeuoaaoe no?ee?noue iiaeo ?? a?a?eae?a aei iaea?eueo
oeiaei/eiieo oai?ia. Aeey oeueiai aaaeeeai ciaoe ia o?eueee ?ioiieia?/i?
oa?aeoa?enoeee aeo?aeieo aaouee?anueeeo oi?i, aea ? caeiiii??iino?
oniaaeeoaaiiy no?eeino? a a?a?eaeao ia?oiai iieie?iiy .

*?oei? caeiiii??iino? a oniaaeeoaaii? no?eeino? aei iooe?/anoi?
naaeee a iaoeo aeine?aeao ia niinoa??aaeinue. I?e aiae?c? no?eeino?
a?a?eae?a, io?eiaieo oeyoii no?auoaaiiy no?eeeo ? iano?eeeo e?i?e
oa?aeoa? oniaaeeoaaiiy a a?eueoino? aeiaaee?a iinea i?ii?aeiee oa?aeoa?,
oi/a niinoa??aaany aieea a?eueo ni?eeiyoeeai? aaouee?anueei? oi?ie. I?e
no?auoaaii? no?eeeo e?i?e i?ae niaith io?eiaii 75% no?eeeo a?a?eae?a.
Iaeneiaeueia o?aaeaiiy iooe?/anoith naaeeith a?aei?/aia i?e no?auoaaii?
no?eeeo e?i?e c iano?eeeie. A a?a?eaeieo eiia?iaoe?yo OeA 10cI o E?I 052
? A?? 44 o Ia 21 o?aaeaiiy ?ineei iooe?/anoith naaeeith aeinyaaei aei
20% a on?o ioieoao aei?iaoaaiiy.

Ioe?ieo i?iyao iciaee no?eeino? aei U. zeae a F1 ?iaeee ca
aeiiiiiaith eiao?oe??ioa nooiaiy aeii?iaioiino? (hp) ( oaae. 2).

Oaaeeoey 2

Oniaaeeoaaiiy iieueiai? no?eeino? aei U. zeae a?a?eaeaie F1

A?oia no?eeino? aaoue-

e?anueeeo oi?i E?euee?noue no?auoaaiue
Nooi?iue aeii?ioaaiiy (hp)

< - 1 a?ae –1 aei –0,5 a?ae –0,5 aei –0 0 a?ae –0 aei +0,5 a?ae+0,5 aei +1 > +1

_ _

e-oue % e-oue % e-oue % e-oue % e-oue % e-oue % e-oue %

N N 29 4 13,8 6 20,7 2 6,9 12 41,4 2 6,9 2 6,9 1 3,4

N N.N 25 5 20,0 4 16,0 2 8,0 6 24,0 0 0 5 20,0 3 12,0

N I 12 0 0 2 16,7 1 8,3 0 0 4 33,3 4 33,3 1 8,3

N.N N 13 5 38,5 2 15,4 0 0 2 15,4 1 7,7 0 0 3 23,1

N.N N.N 6 3 50,0 1 16,7 1 16,7 0 0 0 0 0 0 1 16,7

I N 5 0 0 3 60,0 0 0 0 0 0 0 1 20,0 1 20,0

I N.N 2 0 0 1 50,0 1 50,0 0 0 0 0 0 0 0 0

I?ei?oea : hp< -1 – iaaeaeii?ioaaiiy no?eeino?; 0.5+1–iaaeaeii?ioaaiiy ni?eeiyoeeaino?, N- no?eea, N.N-
na?aaeiueino?eea, I- iano?eea; e-oue – e?euee?noue no?auoaaiue.

Ca ?aaeoe??th a?a?eae?a F1 aei U.zeae nooi?iue aeii?iaioiino?
aa??th? a?ae i?ion aei iethn iane?i/aiiino?, oiaoi niinoa??aaeenue
aeiaaeee iaaaoeaiiai iaaeaeii?ioaaiiy (a?ae’?iiee aaoa?icen),
iaaaoeaiiai aeii?ioaaiiy, i?ii?aeiee i?iya no?eeino?, iiceoeaia
aeii?ioaaiiy ? iiceoeaia iaae-

aeii?ioaaiiy (iiceoeaiee aaoa?icen) no?eeino? aei iaoiaaio.

Ioaea, ioe?iea no?eeino? aei iooe?/anoi? naaeee i?inoeo
a?a?eae?a ia?oiai iieie?iiy ca nooiaiai aeii?iaioiino? iieacaea
ia?aaaaeii i?ii?aeiee oei oniaaeeoaaiiy i?e no?auoaaii? no?eeeo e?i?e ?
i?iya ??cieo oei?a aeii?ioaaiiy a on?o ?ioeo eiia?iaoe?yo no?auoaaiiy.
Oaeee oei oniaaeeoaaiiy oa?aeoa?iee aeey iie?aaiiiai eiio?ieth
no?eeino?. Oea ua ?ac i?aeoaa?aeaeo?, ui no?ee?noue aei U. zeae iineoue
iie?aaiiee oa?aeoa?.

Aeey i?aeoe/ii? naeaeoe?? aaaeeeai aecia/eoe oa?aeoa? oniaaeeoaaiiy
aeanoeainoae no?eeino? ? ni?eeiyoeeaino? aei naaeeiaeo oai?ia. I?i
i?eaeaoi?noue no?eeeo e?i?e aeey aeei?enoaiiy a yeino? aaiaoe/ieo
aeiii??a i?e naeaeoe?? ia ?ioi?oao noaeyoue ca ?acoeueoaoaie
eiia?iaoe?eii? caeaoiino? a ae?aeaeueieo no?auoaaiiyo.

A nai?o aeine?aeao ae?aeaeueiei iaoiaeii ie i?iaiae?coaaee a?n?i
naiicaieeaieo e?i?e, eiio?anoieo ca ?aaeoe??th aei Ustilago. zeae: Ii
236,Ni 151, AeAa 42, An 073IA,S 72, Ni 125, *E 915 ? An 28A ? o?noue
aei Sorosporium. reilianum : E?I 052,I 346cI,OeA 10cI, F 115, *E 209 ? A
246. Aea/aee o?eueee i?yi? eiia?iaoe??, oae ye ca e?oa?aoo?ieie aeaieie
o a?a?eae?a niinoa??aa?oueny ia cia/iee ?aoeei?ieiee aoaeo ca no?ee?noth
aei naaeeiaeo oai?ia.

Aeenia?n?eiee aiae?c ?acoeueoao?a aei?iaoaaiiy a?a?eae?a ia no?ee?noue
aei iooe?/anoi? ? eaoth/i? naaeee, a oaeiae caaaeueii? ? niaoeeo?/ii?
eiia?iaoe?eii? caeaoiino? (CEC ? NEC) ca oeeie iciaeaie o an?o
aeine?aeao iieacaa aenieee ??aaiue ?o aeinoia??iino? (Fo ( F05).
Iaee?aueie ca CEC ii no?eeino? aei U. zeae aeyaeeenue e?i?? *E 915, Ni
151 ? Ni 125 (oaae.3), a aei S. reilianum -I 346cI, F 115 ? OeA 10cI
(oaae.4).

Oaaeeoey 3

Caaaeueia eiia?iaoe?eia caeaoi?noue e?i?e eoeo?oaece ca no?ee?noth aei
iooe?/anoi? naaeee a ??cieo ioieoao aea/aiiy (1998 ?.)

E?i?y “?iaeiee?aea” “Iane?aea”

Aoaeo

CEC Aa??aina

NEC ?aia

e?i?? Aoaeo

CEC Aa??aina

NEC ?aia

e?i??

Ii 236

Ni 151

AeAa 42

An 073IA

S 72

Ni 125

*E 915

An 28A

I??05 1,51

-0,64

0,23

-0,83

1,89

-0,59

-2,49

0,93

0,49 10,41

5,48

2,26

8,29

6,73

6,21

1,5

7,05 7

3

5

2

8

4

1

6 1,05

-2,51

1,79

1,34

3,32

1,02

-4,09

-1,92

0,89 21,39

3,10

7,98

28,71

21,07

12,6

5,42

16,24 5

2

7

6

8

4

1

3

I?ei?oea: iaea?eueoa a?ae’?iia aaee/eia oa?aeoa?eco? iaeaeu? aoaeoe
CEC ca no?ee?noth.

E?i?? eoeo?oaece oa?aeoa?ecothoueny ??cieie aeanoeainoyie
eiia?iaoe?eii? caeaoiino? ca iciaeith no?eeino? aei U. zeae . CEC ? NEC
cia/ii aa??th? i?ae aieeaii aeieia?/ieo oaeoi??a, aea ia a on?o oi?i a
iaeiaeia?e i???. Aenie? ? noaa?euei? aeanoeaino? CEC i?iyaeee e?i?? *E
915, Ni 151 ? Ni 125. E?i?y S 72 a iaio ioieoao aei?iaoaaiiy iieacaea
iaeaeu? aeanoeaino? CEC ca ni?eeiyoeea?noth. A?aeiinii ?ioeo e?i?e
a?aei?/aii i?iya aca?iiae?? aaiioei-na?aaeiaeua.

E?aueie ca CEC ii no?eeino? aei eaoth/i? naaeee aeyaeeenue e?i??
I 346cI ? OeA 10cI, ye? ca oe??th iciaeith ?noioii a?ae??ciyeenue a?ae
?ioeo e?i?e, ? iaee iecuee? cia/aiiy aa??aine NEC, oiio aeai? e?i??
iaea?eueo oe?ii? aeey naeaeoe?? ia no?ee?noue aei eaoth/i? naaeee, oae
ye aiie noaa?eueii ia?aaeaaaoeiooue iioiinoao iciaeo no?eeino?. Aeniea
aa??aina NEC e?i?? F 115 iieaco?, ui ie?ai? eiia?iaoe?? c aeaiith
e?i??th iiaeooue cia/ii a?aeoeeyoenue ye a noi?iio no?eeino? oae ?
ni?eeiyoeeaino?. Aenieee ??aaiue NEC iathoue eiia?iaoe?? i?ae e?i?yie c
aenieeie ? iecueeeie cia/aiiyie CEC.

Oaaeeoey 4

Ioe?iea aoaeo?a caaaeueii? ? eiinoaio niaoeeo?/ii? eiia?iaoe?eii?
caeaoiino? oa aa??ain NEC ca no?ee?noth aei eaoth/i? naaeee
(aacci?iiee iin?a, 1998 ?.)

E?i?? Eiinoaioe NEC

Aoaeo

CEC

Aa??aina

NEC

I 346cI OeA 10cI F 115 *E 209 A 246

E?I 052 -1,24 -0,24 7,15 -0,43 -5,24 2,41 20,07

I 346cI

-0,49 -1,23 7,42 -4,45 -4,14 19,56

OeA 10cI

-2,03 3,56 -0,78 -3,34 4,43

F 115

-12,45 8,57 -2,60 71,33

*E 209

1,90 7,07 56,65

A 246

0,61 31,24

I??05

1,88 1,33

I?ei?oea: iaea?eueo? a?ae’?ii? aaee/eie oa?aeoa?ecothoue iaeaeu? aoaeoe
CEC ? eiinoaioe NEC ca no?ee?noth aei eaoth/i? naaeee.

Ioaea, aeey naeaeoe?? eoeo?oaece ia no?ee?noue aei iooe?/anoi? naaeee
ie ?aeiiaiaeo?ii aeei?enoiaoaaoe e?i?? *E 915, Ni 151 ? Ni 125, ye
aeiii?e no?eeino?, a aei eaoth/i? naaeee – e?i?? I 346cI, OeA 10cI ? F
115. Aeai? e?i?? i?iyaeee aenieo no?ee?noue aei naaeeiaeo oai?ia ?
iathoue aeia?o EC ca no?ee?noth aei aeaieo oai?ia.

Oniaaeeoaaiiy no?eeino? e?i?e aei noaaeiaeo aieeae .

Iaea?eueo iioe?aieie ia eoeo?oaec? ? noaaeia? aiee?, ye?
aeeeeeathoueny a?eaaie ?c ?iaeo Fusarium . Ca aeaieie A.A. ?aauaiea ?
I.I.A?eaeiueiai? (1995) aeii?ioth/a i?noea a a?oi? ooca???a ia? aeae F.
moniliforme.

A iaoeo aeine?aeao eiia?iaoe?eio caeaoi?noue naiicaieeaieo e?i?e
ie aecia/aee iaoiaeii ae?aeaeueieo no?auoaaiue ca iaoiaeeeith A??oo?iaa-
iaoiae 4, iiaeaeue 1 .

Aiae?c aoaeo?a caaaeueii? eiia?iaoe?eii? caeaoiino? ca no?ee?noth aei
noaaeiaeo aieeae iieacaa, ui ca oe??th iciaeith aeae?eeeenue e?i?? I
346cI, I 502, *E 929 ? F2 (oaae. 5).

Oaaeeoey 5

Caaaeueia eiia?iaoe?eia caeaoi?noue e?i?e eoeo?oaece ca no?ee?noth aei
noaaeiaeo aieeae, 1997–1998 ??.

Ioieo aei?iaoaaiiy

Iacaa Oiainueea NAA (?iaeiee?aea( (Iane?aea(

e?i?? Aoaeo CEC ?aia

E?i?? Aoaeo CEC ?aia

e?i?? Aoaeo CEC ?aia

e?i??

THA 24 6,77 4 6,33 5 7,43 5

F2 — — -7,79 4 -6,06 4

OeA 10cI 12,04 5 11,24 7 16,35 7

O 23 19,87 6 19,38 8 20,71 8

I 502 -19,21 1 -12,61 2 -14,23 2

*E 929 -6,65 3 -10,5 3 -13,34 3

*E 218IA — — 8,72 6 12,6 6

I 346cI -12,82 2 -14,82 1 -18,97 1

I??05 2,45

1,25

2,75

I?ei?oea: Iaea?eueoa a?ae’?iia aaee/eia oa?aeoa?eco? iaeaeu? aoaeoe CEC
ca no?ee?noth aei noaaeiaeo aieeae.

E?i? I 346cI ? *E 929 ii?aeiothoue aenieo CEC ca no?ee?noth aei
noaaeiaeo aieeae c aenieeie aoaeoaie CEC ca o?iaeaei?noth, oiio aiie ?
iniaeeai oe?iieie a naeaeoe??. E?i?y F2 oaeiae aeoaea oe?iia aeey
naeaeoe?? a?a?eae?a ia no?ee?noue aei noaaeiaeo aieeae, oiio ui aiia
?aiiioaioia, a aeniea o?c?ieia?/ia ?aaeoeai?noue ?aiiioaioieo e?i?e
no?eio? ?icaeoie aaaaoueio oei?a aieeae. A iaoeo aeine?aeao a?eueo?noue
(84,6%) a?a?eae?a ca o/anoth e?i?? F2 i?iyaeee aenieo no?ee?noue aei
noaaeiaeo aieeae a on?o ioieoao aei?iaoaaiiy.

No?ee?noue a?a?eae?a aei noaaeiaeo aieeae caeaaeeoue a?ae e?eueeino?
no?eeeo aaouee?anueeeo oi?i. Oae, ?c 23 a?a?eae?a, ui a?aeianai? a a?oio
aenieino?eeeo, 70% iaee no?ee? iaeaea? aaouee?anuee? oi?ie, aai iaeio ?c
ieo. No?ee? a?a?eaee a iniiaiiio io?eiai? i?e no?auoaaii? no?eeeo e?i?e
?c ni?eeiyoeeaeie (87%). No?auoaaiiy na?aaeiueino?eeeo e?i?e i?ae niaith
aea?, ye i?aaeei, ? na?aaeiueino?ee? a?a?eaee. Aeey noai?aiiy no?eeeo
a?a?eae?a aaaeaii aeei?enoiaoaaoe iaeaea? no?ee? oi?ie , aai oi/ iaeio
no?eeo, yea a aeei?enoiaoaaeanue a ?ie? iaoa?einueei? oi?ie.

Oaeei /eiii, aeey naeaeoe?? eoeo?oaece ia no?ee?noue aei noaaeiaeo
aieeae a oiiaao E?ninoaio Oe?a?ie ie ?aeiiaiaeo?ii aeei?enoiaoaaoe e?i??
I 346cI, I 502, *E 929 ? F2, ye aeiii?e no?eeino?. Aeai? e?i?? i?iyaeee
aenieo no?ee?noue aei noaaeiaeo aieeae a ??cieo aeieia?/ieo oiiaao,
iathoue aenieo eiia?iaoe?eio caeaoi?noue ca no?ee?noth aei oe???
oai?iae, a e?i?? I 346cI ? *E 929 iathoue ua e aenie? aoaeoe CEC ca
o?iaeaei?noth.

A oniaaeeoaaii? no?eeino? aei noaaeiaeo aieeae aieiaio ?ieue
a?ae?a?athoue aaie c aaeeoeaieie

aoaeoaie.

No?ee?noue aei noaaeiaeo aieeae aeia?a oniaaeeiao?oueny, i?i ui
na?ae/eoue iacia/ia ??cieoey i?ae eiao?oe??ioaie oniaaeeoaaiiy a
oe?ieiio ? aocueeiio ?icoi?ii?.

Aeey iciaee no?eeino? aei noaaeiaeo aieeae io?eiai? iaeaea ??ai?
eiao?oe??ioe oniaaeeoaaiiy a oe?ieiio ? aocueeiio ?icoi?ii? a ??cieo
ioieoao aei?iaoaaiiy.

Oea aiai?eoue i?i oa, ui a aaiaoe/iiio eiio?ie? oe??? iciaee
iniiaio ?ieue a?ae?a?athoue aaeeoeai? aoaeoe aai?a ? aaeeoeaia
i?ieea?noue a caaaeuei?e oaiioeiia?e i?ieeaino? a?aeiinii aaeeea a
ii??aiyii? c na?aaeiaeuiith.

Oniaaeeoaaiiy no?eeino? e?i?e aei ooca??ico ea/ai?a

Aea/aiiy oniaaeeoaaiiy no?eeino? aei ooca??ico ea/ai?a a
a?a?eaeao F1 a nenoai? ae?aeaeueieo no?auoaaiue iaie i?iaiaeeeinue
i?ioyaii aeaio ?ie?a a ??cieo ioieoao . A noaio no?auoaaiiy aoei
aeeth/aii a?n?i e?i?e: Ia 21, Oo719, F2, *E 4282, Ni 125, OeA 10cI, *E
218IA oa A?? 44.

Aeenia?n?eiee aiae?c aeaieo iieacaa aeinoia??io ??cieoeth i?ae
a?a?eaeaie ca o?aaeaiiyi ooca??icii.

Ia iniia? ii??aiyiiy na?aaei?o eaaae?ao?a caaaeueii? ?
niaoeeo?/ii? EC iiaeia c?iaeoe aeniiaie, ui aieea aai?a aaeeoeaiiai oeio
(aecia/ath/eo CEC) cia/ii aeuee aieeao aai?a iaaaeeoeaiiai oeio ae??
(aecia/ath/eo NEC). *ei cia/i?o? eieeaaiiy, aeeeeeai? CEC, oei a?eueoee
aieea aai?a aaeeoeaiiai oeio.

Cia/ia aa??thaaiiy iieacieea CEC/NEC (1,9 – 6,4), aeaco? ia neeaaeio
nenoaio oniaaeeoaaiiy no?eeino? aei ooca??ico ea/ai?a ? ia iiaeeea?noue
cia/iiai aieeao iaaaeeoeaieo aoaeo?a (aeii?ioaaiiy oa ai?noaco). Ia oea
aeaco? oaeiae cia/ia i?ieea?noue eiao?oe??io?a oniaaeeoaaiiy a oe?ieiio
? aocueeiio ?icoi?ii? (I2 = 0,78 – 0,95 ? h2 = 0,21-0,60).

Na?aae aea/aieo e?i?e iaeaeuo CEC ca iciaeith no?eeino? aei
ooca??ico ea/ai?a iathoue e?i??: Ni 125,*E 218IA,OO 719 oa Ia 21(oaae.
6).

Oaaeeoey 6

Caaaeueia eiia?iaoe?eia caeaoi?noue e?i?e eoeo?oaece ca no?ee?noth aei
ooca??ico ea/ai?a (1998 ??e)

Ioieo aei?iaoaaiiy

Iacaa Oiainueea NAA (?iaeiee?aea( (Iane?aea(

e?i?? Aoaeo CEC ?aia

E?i?? Aoaeo CEC ?aia

e?i?? Aoaeo CEC ?aia

E?i?? Ni 125 oa *E 218IA c aenieeie cia/aiiyie CEC oa iecueeith
aa??ainith NEC noaa?eueii ia?aaeathoue iciaeo no?eeino? iioiinoao a
??cieo eiia?iaoe?yo no?auoaaiiy.

Iecueea cia/aiiy CEC iaea e?i?y OeA 10cI ca no?ee?noth aei
ooca??ico ea/ai?a ? noaaeiai? aieeecie. A?a?eaee ca o/anoth aeaii? e?i??
iaea?eueoa o?aaeoaaeenue noaaeiaeie aieeyie ? ooca??icii a on?o ioieoao
aea/aiiy. Oea aeaco? ia iiaeeea?noue aeeoociiai o?aaeaiiy ca?ai
ooca??icii /a?ac noa?aeaiue ea/aia ii noaeeiii-aieieienoeo io/eao
n?i’yi?aeee. I?i aeiaaeee aeeoociiai o?aaeaiiy eoeo?oaece ooca??icii
iia?aeiieyeinue ? ?ai?oa.

Oaeei /eiii, i?e oniaaeeoaaii? no?eeino? aei ooca??ico ea/ai?a
ia?aaaaeio ?ieue a?ae?a?athoue aaie c aoaeoaie aaeeoeaii? ae?? , aea
cia/ia aa??thaaiiy iieacieea CEC/NEC, oa eiao?oe??io?a oniaaeeoaaiiy a
oe?ieiio oa aocueeiio ?icoi?ii? aeaco? ia iiaeeea?noue cia/iiai aieeao
iaaaeeoeaieo aoaeo?a.

No?ee?noue a?a?eae?a caeaaeeoue a?ae no?eeino? aaouee?anueeeo
oi?i, a oaeiae a?ae ?o i?iaeoeoeaiino?. Na?aae aea/aiiai iaai?o
naiicaieeaieo e?i?e aeuo CEC ca no?ee?noth aei ooca??ico ea/ai?a
i?iyaeee e?i?? Ni 125, *E 218IA, Oo 719 oa Ia 21. Aeai? e?i?? iiaeia
aeei?enoiaoaaoe a yeino? aeiii??a no?eeino? aei aeaii? oai?iae.

OEIAeI*EII?NOUe INIIAIEO OAI?IA O NO*ANIEO A?A?EAe?A ? ?O AAOUeE?ANUeEEO
OI?I

Iaeaea a on?o ?aeiiao ae?iuoaaiiy eoeo?oaece ia ca?ii cia/io
oeiaeo ni?e/eiythoue iooe?/anoa , eaoth/a naaeea, ei?aiaa? ? noaaeia?
aiee? oa ooca??ic ea/ai?a. Oeiaei/eii?noue oeeo oai?ia ia iaeiaeiaa.

A ca’yceo c oei, ui naeaeoe?y ia no?ee?noue eoeo?oaece a
iniiaiiio aaeanue aei iooe?/anoi? naaeee, ca?ac oeiaei/eii?noue oe???
oai?iae ia aeniea. I?iaaaeaia aea/aiiy iieacaei, ui ??aaiue no?eeino?
?aeiiiaaieo ? iiaeo e?i?e oa a?a?eae?a aeinoaoi?e aeey oo?eiaiiy ??aiy
iaaeiai??a o?iaeath a?ae o?aaeaiiy iooe?/anoith naaeeith ieae/a ii?iao
oeiaei/eiiino?.

I?e na?aaeiueiio eiao?oe??io? oeiaei/eiiino? iooe?/anoi? naaeee
0,25% , ao?aoe o?iaeath a iaoeo aeine?aeao ia ia?aaeuoaaee 0,4-0,5% o
e?i?e ? 0,3-0,6% o a?a?eae?a.

A oiiaao n?aici?ie i?ioyaii o?ueio ?ie?a aea/aiiy a?a?eaee ia
o?aaeoaaeenue eaoth/ith naaeeith, a na?aae aea/aieo e?i?e o?eueee e?i?y
I 502 aoea o?aaeaia a iacia/i?e i???.

Aeniea oeiaei/eii?noue eaoth/i? naaeee i?iyaeeany a oiiaao
aacci?iiiai iin?ao a 1998 ?ioe?. O ni?eeiyoeeaeo e?i?e ? a?a?eae?a
ao?aoe a?ae eaoth/i? naaeee iiaeooue aeinyaaoe aei 30%. Oea iia’ycaia c
oei, ui na?aae aea/aieo oai?ia eaoth/a naaeea ia? iaea?eueoee
eiao?oe??io oeiaei/eiiino?. Ca aeaieie A.A.A?enaiea oa ?.E.Aeoaeee a?i
noaiiaeoue 1,2. ? oi/a a oiiaao n?aici?ie aeaia oai?iaa ia i?iyaeeanue,
aea , ye iieacaee aeine?aeaeaiiy, i?e iaeiie/aii? ?ioaeoe?? S. reilianum
a a?oio?, eaoth/a naaeea iiaea aeeeeeaoe cia/i? ao?aoe.

Iaea?eueo oeiaei/eiiith a oiiaao E?ninoaio Oe?a?ie ? noaaeiaa
aieeecia, yea e??i cieaeaiiy o?iaeath iaiineoue aaeeeo iia?/io oeiaeo,
aeeeeeath/e eaie?noue noaaae, ui ca?eueoo? ao?aoe i?e cae?aii?. Ao?aoe
o?iaeath a?ae aieeae noaiiaeee 1,3-8,9% ia eiaei? 10% o?aaeaieo ?ineei.

Ao?aoe o?iaeath a?ae ooca??ico ea/ai?a iiaeooue aeinyaaoe 8,5% o
ni?eeiyoeeaeo a?a?eae?a, aea o a?eueoino? a?a?eaeieo eiia?iaoe?e ao?aoe
a?ae oe??? oai?iae ia?aaoaaee a iaaeao oeiaei/eiiino? ? ia ia?aaeuoaaee
5%.

Cia/ia o?aaeaiiy ea/ai?a ooca??icii niinoa??aaeinue i?e
iioeiaeaeaii? ?o noaaeiaei iaoaeeeii. Aeyaeaii aenieiaeinoia??iee
ca’ycie i?ae nooiaiai o?aaeaiiy ea/ai?a ooca??icii ? iioeiaeaeaiiyi ?o
eoeo?oaecyiei iaoaeeeii. Ia eiaei? 10% i?e?inoo iioeiaeaeaiiy ea/ai?a
oe?aeieeii o?aaeaiiy ooca??icii ca?eueooaaeinue ia 4,5-7,5%. Noia?iee
iaaeia?? o?iaeath a?ae o?aaeaiiy oai?iaaie ? ooeiaeaeaiiy noaaeiaei
iaoaeeeii o ni?eeiyoeeaeo a?a?eae?a iiaea aeinyaoe 11,6%.

Aeniiaee

Aeiaaaeaia iiaeeea?noue io?eiaiiy aeinoia??ii? oa?aeoa?enoeee
naiicaieeaieo e?i?e ? a?a?eae?a eoeo?oaece ca no?ee?noth aei iniiaieo
oai?ia (iooe?/anoa ? eaoth/a naaeea, noaaeia? aiee? ? ooca??ic ea/ai?a)
ia i?e?iaeiiio (i?iaieaoe?eiiio) oii? a ni?eyoeea? aeey ?icaeoeo
iaoiaai?a ?iee . Aeey ia’?eoeaii? ioe?iee naeaeoe?eiiai iaoa??aeo
aeinoaoi? eiai aeai??/ia aea/aiiy a aeaio — o?ueio ioieoao c ??cieie
aeieia?/ieie oiiaaie.

Anoaiiaeaii, ui no?ee?noue eoeo?oaece aei an?o aea/aieo oai?ia
caeaaeeoue a?ae aaiioeiiaeo iniaeeainoae e?i?e, aeieia?/ieo oiia,
no?ie?a n?aae ? ?o nei?inoeaeino?. A?eueoo no?ee?noue aei oai?ia
i?iyaeee oi?ie c o?eaae?oei aaaaoaoe?eiei ia??iaeii.

Aeyaeaii na?aaei?e cai?ioi?e ei?aeyoe?eiee ca’ycie i?ae nooiaiai
o?aaeaiiy iooe?/anoith naaeeith ? aenioith i?ee??ieaiiy ea/aia (r =
-0,67) oa neaaeee cai?ioi?e ca’ycie i?ae o?aaeaiiyi noaaeiaeie aieeyie ?
ianith 1000 ca?ai(r = -0,39). I?ae ?ioeie ii?oieia?/ieie iciaeaie ?
no?ee?noth aei oai?ia ?noioieo ca’yce?a ia aeyaeaii.

Eiiieaenio no?ee?noue aei iniiaieo oai?ia i?iyaeee naiicaieeai? e?i??
na?oiai? eieaeoe?? (I 346cI, I 502, Ia 21, F115,F 2,An 28A) ?
a?o/eciyii? naeaeoe?? (*E 915, *E 929, Oo 719).

Aeine?aeaeaii, ui iaea?eueo noaa?eueii no?eeeie a ??cieo oiiaao
ae?iuoaaiiy aoee a?a?eaee F1 , io?eiai? a?ae no?auoaaiiy no?eeeo e?i?e
i?ae niaith. I?e no?auoaaii? no?eeeo e?i?e c ni?eeiyoeeaeie ? iaaiaee, a
caeaaeiino? a?ae eiie?aoieo aaouee?anueeeo ia? iiaeia io?eiaoe a?a?eaee
F1 a?ae aeniei no?eeeo aei ni?eeiyoeeaeo.

Ioe?iea no?eeino? aei iooe?/anoi? naaeee i?inoeo a?a?eae?a ia?oiai
iieie?iiy ca nooiaiai aeii?iaioiino? iieacaea ia?aaaaeii i?ii?aeiee oei
oniaaeeoaaiiy i?e no?auoaaii? no?eeeo e?i?e ? i?iya ??cieo oei?a
aeii?ioaaiiy a on?o ?ioeo eiia?iaoe?yo no?auoaaiiy.

Aea/aiiy oniaaeeoaaiiy no?eeino? e?i?e aei oai?ia a a?a?eaeao F1
iaoiaeii ae?aeaeueieo no?auoaaiue iieacaei aieiaio ?ieue aai?a c
aaeeoeaiei aoaeoii, ui aeicaiey? ?aeiiaiaeoaaoe aeia?? a naeaeoe?eiiio
i?ioean? ca oaiioeiii.

A ?acoeueoao? aea/aiiy eiia?iaoe?eii? caeaoiino? aeo?aeieo naiicaieeaieo
e?i?e ca no?ee?noth aei iniiaieo oai?ia aeae?eaii aeaea?aea no?eeino?
aei iooe?/anoi? naaeee (*E 915, Ni 151, Ni 125), eaoth/i? naaeee (I
346cI, OeA 10cI, F115), noaaeiaeo aieeae (I 346cI, I 502, *E 929,
F2),ooca??ico ea/ai?a (Oo 719,*E 218IA, Ia 21), ye? iaee aenie? aoaeoe
CEC a on?o ioieoao aei?iaoaaiiy. E?i?? *e 915, I 346cI, *E 929, I 502,
Oo 719, f2 ii?aeiothoue aenieo CEC ca no?ee?noth aei oai?ia ? o?iaea?i
ca?ia.

Iaea?eueo oeiaei/eiiith a oiiaao E?ninoaio Oe?a?ie ? noaaeiaa aieeecia,
yea e??i cieaeaiiy o?iaeath iaiineoue aaeeeo iia?/io oeiaeo, aeeeeeath/e
eaie?noue noaaae, ui ca?eueoo? ao?aoe i?e cae?aii?. Ao?aoe o?iaeath a?ae
aieeae noaiiaeee 1,3-8,9% ia eiaei? 10% o?aaeaieo ?ineei.

Aeyaeaii aenieiaeinoia??iee cay’cie i?ae nooiaiai o?aaeaiiy ea/ai?a
ooca??icii ? iioeiaeaeaiiyi ?o eoeo?oaecyiei iaoaeeeii. Ia eiaei? 10%
i?e?inoo iioeiaeaeaiiy ea/ai?a oe?aeieeii o?aaeaiiy ooca??icii
ca?eueooaaeinue ia 4,5-7,5%. Noia?iee iaaeia?? o?iaeath a?ae o?aaeaiiy
oai?iaaie ? ooeiaeaeaiiy noaaeiaei iaoaeeeii o ni?eeiyoeeaeo a?a?eae?a
iiaea aeinyaoe 11,6%.

?aeiiaiaeaoe?? aeey naeaeoe?eii? i?aeoeee

Aeey aeinoia??ii? ioe?iee naiicaieeaieo e?i?e ? a?a?eae?a eoeo?oaece ca
no?ee?noth aei iniiaieo oai?ia ia i?e?iaeiiio ( i?iaieaoe?eiiio) oii?
iaiao?aeii ?o aea/aoe ia iaioa aeaio ?ie?a a aeaio-o?ueio ioieoao c
??cieie aeieia?/ieie oiiaaie.

A ca’yceo c oei , ui a oniaaeeoaaii? no?eeino? aei iooe?/anoi? oa
eaoth/i? naaeee naaeee, noaaeiaeo aieeae ? ooca??ico ea/ai?a aieiaio
?ieue a?ae?a?athoue aaie c aaeeoeaiei aoaeoii ?aeiiaiaeo?oueny aeia?? a
naeaeoe?eiiio i?ioean? ca oaiioeiii.

I?iiiio?oueny aeei?enoaoe a naeaeoe?? a?a?eae?a eoeo?oaece e?i??, ui ?
aeaea?aeaie no?eeino? aei iooe?/anoi? naaeee (*E 915, Ni 151,Ni125),
eaoth/i? naaeee (I346cI, OeA10cI,F115), noaaeiaeo aieeae (I346cI, I502,
*E929, F2),ooca??ico ea/ai?a (Oo 719,*E 218IA, Ia 21), ye? iieacaee
aenie? aoaeoe CEC a on?o ioieoao aei?iaoaaiiy.

NIENIE ?IA?O IIOAE?EIAAIEO II OAI? AeENA?OAOe??

?oaai ?.I. Aiae?c eiia?iaoe?eii? caeaoiino? naiicaieeaieo e?i??
eoeo?oaece ca no?ee?noth aei noaaeiaeo aieeae // Ca??iee iaoeiaeo
i?aoeue Oiainueei? NAA.- Oiaiue, 1998.- N. 75-78.

?oaai ?.I. Oniaaeeoaaiiy no?eeino? naiicaieeaieo e?i?e eoeo?oaece aei
ooca??ico ea/ai?a a a?a?eaeao ia?oiai iieie?iiy. // Ca??iee iaoeiaeo
i?aoeue Oiainueei? NAA. No/ani? i?iaeaie ?ineeiieoeoaa ?
ei?iiae?iaieoeoaa.- Oiaiue, 1998.- *.2.- N. 94-97.

?oaai ?.I. Naeaeoe?y eoeo?oaece ia no?ee?noue aei noaaeiaeo aieeae a
oiiaao E?ninoaio Oe?a?ie. // Ca??iee iaoeiaeo i?aoeue ?inoeoooo
caiea?ianoaa OAAI.- Ee?a, 1999.- Aei.1-2.- N. 121-124.

?oaai ?.I. Naeaeoe?y eoeo?oaece ia no?ee?noue aei naaeeiaeo oai?ia a
oiiaao E?ninoaio Oe?a?ie. //Atheaoaiue ?inoeoooo
n?euenueeiainiiaea?nueei? i?e?ia?ieia??. *a?i?a?a, 1999.- ?4.- N.17-19.

*o/i?e ?.I., Iiaeieyi A.A., ?oaai ?.I., No?oi?inueeee TH.I. Naeaeoe?y oa
ian?iieoeoai a?a?eae?a eoeo?oaece // A?niee aa?a?ii? iaoee .-
1999.-Niaoe. aeione.- N.61-64.

AIIOAOe?ss

?oaai ?.I. Naeaeoe?y a?a?eae?a eoeo?oaece ia no?ee?noue aei
oai?ia a oiiaao E?ninoaio Oe?a?ie.-?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
n?euenueeiainiiaea?nueeeo iaoe ca niaoe?aeuei?noth 06.01.05 – naeaeoe?y
? ian?iieoeoai. ?inoeooo caiea?ianoaa OAAI, nio. *aaaie, 1999.

I?iaaaeaii aea/aiiy ?ioiieia?/ieo oa?aeoa?enoee 24 naiicaieeaieo
e?i?e eoeo?oaece ??ciiai iioiaeaeaiiy ? caeiiii??iinoae oniaaeeoaaiiy
no?eeino? aei iniiaieo oai?ia a?a?eaeaie F1 a oiiaao E?ninoaio Oe?a?ie.

Aeyaeaii ei?aeyoe?ei? aca?iica’ycee i?ae iniiaieie
ainiiaea?nueei-a?ieia?/ieie iciaeaie ? no?ee?noth aei oai?ia oa
eeaneo?eiaaii aeo?aeiee iaoa??ae ca no?ee?noth aei iniiaieo oai?ia.

Aeae?eaii c?acee c eiiieaeniith no?ee?noth aei naaeeiaeo oai?ia,
noaaeiaeo aieeae oa ooca??ico ea/ai?a. Cai?iiiiiaaii aeei?enoaoe a
naeaeoe?? a?a?eae?a eoeo?oaece e?i??, ui ? aeaea?aeaie no?eeino? aei
ie?aieo oai?ia. Aecia/aii, ui iaea?eueo oeiaei/eiieie a oiiaao E?ninoaio
Oe?a?ie ? noaaeia? aiee?.

Eeth/ia? neiaa: eoeo?oaeca, naiicaieeai? e?i??, a?a?eaee ,
no?ee?noue, oai?iae, oniaaeeoaaiiy, eiia?iaoe?eia caeaoi?noue,
oeiaei/eii?noue.

AIIIOAOeEss

?oaai E.I. Naeaeoeey aea?eaeia eoeo?ocu ia onoie/eainoue e
aieaciyi a oneiaeyo Eaninoaie Oe?aeiu. ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa
naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe 06.01.05 – naeaeoeey e
naiaiiaiaenoai. – Einoeooo caieaaeaeey OAAI, *aaaiu, 1999.

I?iaaaeaii eco/aiea eiioiieiae/aneeo oa?aeoa?enoee 24
naiiiiueaiiuo eeiee eoeo?ocu ?aciiai i?ienoiaeaeaiey e caeiiiia?iinoae
ianeaaeiaaiey onoie/eainoe e iniiaiui aieaciyi aea?eaeaie F1 a oneiaeyo
Eaninoaie Oe?aeiu. Auyaeaii ei??aeyoeeiiiua acaeiinayce iaaeaeo
iniiaiuie oicyenoaaiii – aeieiae/aneeie i?eciaeaie e onoie/eainoueth e
aieaciyi e eeanneoeoee?iaaii enoiaeiue iaoa?eae ii onoie/eainoe e
iniiaiui aieaciyi.

Auaeaeaiu ia?acoeu n eiiieaeniie onoie/eainoueth e aieiaiaaui
aieaciyi, noaaeaaui aieeyi e ooca?eico ii/aoeia. Iniaaiii oeaiiuie aeey
naeaeoeee ia a?oiiiaie eiioieoao yaeythony neaaeothuea eeiee *E 915, *E
929, I346cI, eioi?ua i?iyaeee aunieith onoie/eainoue e eco/aaiui
aieaciyi eae a oneiaeyo anoanoaaiiiai, oae e i?iaieaoeeiiiiai oiiia.

Eeiee Ia 21, F115 e Oo 719 i?iyaeee aunieoth onoie/eainoue e
aieiaia e ooca?eico ii/aoeia e oia?aiioth onoie/eainoue e noaaeaaui
aieeyi. A?oiiiaoth onoie/eainoue e aieiaiaaui aieaciyi e noaaeaaui
aieeyi i?iyaeee eeiee An 28A e F 2. ?aeiiaiaeoaony eniieueciaaoue a
naeaeoeee aea?eaeia eoeo?ocu aeaiiua eeiee

A i?ioeanna enneaaeiaaiee onoaiiaeaii, /oi onoie/eainoue e
eco/aaiui aieaciyi eiio?iee?oaony aaeaeeoeaii – aeiieiaioiie
aaiaoe/aneie nenoaiie. Eco/aiea iniaaiiinoae ianeaaeiaaiey onoie/eainoe
e iocu?/aoie aieiaia, iueueiie aieiaia, noaaeaaui aieeyi e ooca?eico
ii/aoeia iieacaei, /oi a iniiaa aaiaoe/aneiai eiio?iey yoeo i?eciaeia
eaaeao i?aeiouanoaaiii aaiu n aaeaeeoeaiuie yooaeoaie, /oi iicaieyao
?aeiiaiaeiaaoue ioai? a naeaeoeeiiiii i?ioeanna ii oaiioeio.

. A ?acoeueoaoa eco/aiey eiiaeiaoeeiiiie niiniaiinoe enoiaeiuo
naiiiiueaiieo eeiee ii onoie/eainoe e iniiaiui aieaciyi auaeaeaiu
enoi/ieee onoie/eainoe e iocu?/aoie aieiaia (*E 915, Ni 151, Ni 125),
iueueiie aieiaia (I 346cI, OeA 10cI, F115), noaaeaaui aieeyi (I 346cI, I
502, *E 929, F2),ooca?eico ii/aoeia (Oo 719,*E 218IA, Ia 21), eioi?ua
eiaee aunieea yooaeou IEN ai anao ioieoao eniuoaiey. Eeiee *e 915, I
346cI, *E 929, I 502, Oo 719, f2 eiatho aunieoth IEN eae ii onoie/eainoe
e aieaciyi, oae e ii o?iaeath ca?ia.

Ii?aaeaeaii, /oi iaeaieaa a?aaeiiiniie a Eaninoaie Oe?aeiu
yaeyaony noaaeaaay aieeue. Iioa?e o?iaeay io noaaeaauo aieeae ninoaaeee
1,3-8,9% ia eaaeaeua 10% ii?aaeaiiuo ?anoaiee.

Eeth/aaua neiaa: eoeo?oca, naiiiiueaiiay eeiey, aea?eae,
onoie/eainoue, aieaciue, ianeaaeiaaiea, eiiaeiaoeeiiiay niiniaiinoue,
a?aaeiiiniinoue.

ANNOTATION

Ruban I.O. Plant breeding of hybrids of corn for disease
resistance to in conditions of Forest-steppe of Ukraine. The manuscript.

The dissertation on competition of a scientific degree of the
candidate of agricultural sciences on a speciality 06.01.05 — selection
and seeding. — Institute of agriculture UAAS, Chabany, 1999.

The study of immunity of the characteristics 24 inbred of lines
of corn of a different origin and laws of inheritance of stability to
the basic disease by hybrids F1 in conditions of a Forest-Steppe of
Ukraine is carried out. Is revealed correlation interrelations between
basic facilities — by biological attributes and stability to i disease
and initial material by the initial material by an initial material by

Samples with complex stability to smut to disease, stalk
rottenness and Fusarium of ear are allocated. It is recommended to use
in selection of hybrids of corn of a line, that are sources of stability
to separate

disease. Is determined, that most harmful in a Forest-Steppe of Ukraine
is stalk rottenness.

Key words: corn, inbred a line, hybrid, resistance, disease,
inheritance, combinational ability, harmful.

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