Ee?anueeee oi?aa?neoao ?iai? Oa?ana Oaa/aiea

OAeE 582, 287; 577, 156

IIEIOIIA IEUeAA IEAENAIAe?IAIA

?aaoeyoeiy aeoeaiinoi aecii?ioa?iac iieieicniaeaeueii?

ae?? ooaiia Hirschioporus laricinus (Karst.) Ryv

Niaoe?aeuei?noue — 03.00.04 aiioiiiy

Aaoi?aoa?ao

ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa aiieiai/ieo iaoe

Ee?a – 1999

Aeena?oaoe??th ? ?oeiien

Aeena?oaoe?y aeeiiaia o Aeiiaoeueeiio aea?aeaaiiio oiiaa?neoao?

Iaoeiaee ea??aiee — Aieei Ieoaeei Iaaiiae/, aeieoi? a?ieia?/ieo iaoe,

i?ioani?, caa?aeoaa/ eaoaae?e Aeiiaoeueeiai aea?aeaaiiai oi?aa?neoaoo.

Io?oe?ei? iiiiaioe — Aa?aaiaoeue Ethaeieea Aeieo?iaia, aeieoi?
a?ieia?/ieo

iaoe, ?inoeooo iie?iaiieia?? i ai?onieia?? ii.Ae.E.Caaieioiiai IAI
Oe?a?ie.

Oeaeiye Naioeaia Aieiaeeie?iaia, aeieoi? a?ieia?/ieo

iaoe, Ee?anueeee oi?aa?neoao ?iai? Oa?ana Oaa/aiea.

I?ia?aeia onoaiiaa — Oa?eianueeee aea?aeaaiee oiiaa?neoao, eaoaae?a
o?c?ieia??

oa aiioiii? ?ineei, i.Oa?e?a.

Caoeno a?aeaoaeaoueny 25 o?aaiy 1999 ?ieo i 13 aiaeei? ia cn?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae 26.001.24 Ee?anueeiai oi?aa?neoaoo
?iai? Oa?ana Oaa/aiea ca aae?anith: 252127, i?. aeaae.Aeooeiaa, 6)

Aaoi?aoa?ao ?ic?neaiee 22 ea?oiy 1999 ?ieo.

C aeena?oaoe??th iiaeia iciaeiieoeny a a?ae?ioaoe? Ee?anueeiai
oi?aa?neoaoo ?iai? Oa?ana Oaa/aiea (252033, i.Ee?a, aoe.Aieiaeee?nueea,
58)

A/aiee nae?aoa?

Niaoe?ae?ciaaii? a/aii? ?aaee

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeueiinoue i?iaeaie. ?ic?iaea oa inai?iiy iiaeo aenieiaoaeoeaieo
oaoiieiaie, yei ni?eythoue eiiieaeniiio aeei?enoaiith iieiea, — aaaeeeaa
ii?iaeii-ainiiaea?nueea caaaeaiiy. Ai?iaaaeaeaiiy aei oaoiieiai?
ae?iaieoeueoaa oa?iaioieo i?aia?aoia iie?iaiiai oa a?eaiiai iioiaeaeaiiy
iiaea nooo?ai aieeaaoe ia aeineoue aeiaai ianeaii noaie iaea?aeaiiy
iiei/ieo i?iaeoeoia. Oa?iaioe iiaeiii oe?iei aea/aoeny oa aaeeaaoeny c
iaoith iiaeaeuaiiy yeinoi i?iaeoeoei?, io?eiaiy i?eioeeiiai iiaeo
i?iaeoeoia oa iiaeaeuaiiy ?aioaaaeueiinoi ae?iaieoeoaa.

Ne/oaeiee oa?iaio, ca o?aaeeoe?th, ae?iaeythoue ic ne/oaa yaiyo oa
oaeyo. I?e oeueiio io?eio?oueny ia aeineoue aeoeaiee oa?iaio c aaeeeei
aiinoii aaeanoiaeo ?a/iaei. ?icoe?aiiy ae?iaieoeoaa ne?ia oa iioeo
iiei/ieo ae?iaia iio?aaothoue iiooeo caiiiieeia ne/oaeieo oa?iaioia.

Oa?iaioii nenoaie aeueo aaceaeiaeueieo a?eaia ca?ac aea/athoueny
aieueo oe?iei oa ieaiiii?ii, aea iioeaeiaeueii a?eae o oeueiio aniaeoi
aea/aii ua ia aeineoue aeia?a. Aaaeeeainoue iiaeiaieo aeineiaeaeaiue
aacia?a/ia, oiio ui ye oiciieiai/ii, oae i oa?iaioaoeaii oa?aeoa?enoeee
/enoeo iioeaeiaeueieo eoeueoo?, ?o ia?aeiii oa aoi?eiii i?ioeane
iaoaaieicio oa aiaeiiaiaeii i?iaeoeoe yaii aeacothoue ia oieueee ia
iaoeiaa cia/aiiy iai?yieo, aea i ia eiai ia?niaeoeaiinoue aeey
i?aeoe/iiai aeei?enoaiiy.

Ii?iaiyeueia i?inoioa eoeueoeaoaaiiy a?eaia, ?o aeniea oiciieiai/ia oa
aaiaoe/ia ieanoe/iinoue aiaee?eaathoue aaeeei iiaeeeainoi coiniaii
i?iieneiai? oa?iaioieiai? oa naeaeoei? ooaiia-i?iaeooeaioia.

Io?eiaiiy oeiiieo a?eaieo iaoaaieioia iia(ycaii c iioeiicaoei?th oiia
aeey ?o aiineioaco iiae /an ooo/iiai eoeueoeaoaaiiy oa ? iaeii?th c
iniiaieo caaea/, yea iio?aao? ae?ioaiiy. Ai?iaaaeaeaiiy aei noa?e
i?iieneiaiai aeei?enoaiiy ye iiaeia aieueoi? eieueeinoi aaceaeiiiioeaoia
i?eaaaea aei caieueoaiiy ei?enieo ?ano?nia iaoi? aea?aeaae.

Iaoa i caaaeaiiy aeineiaeaeaiue. Iaoith ?iaioe aoei iiaeaeuaiiy neioaco
i?ioa?iac iieieicniaeaeueii? aei? ooaiia I-81, A-031, A-032 i ?-323 H.
laricinus oeyoii aeei?enoaiiy ?icieo oaeoi?ia.

Aeey aeinyaiaiiy oei?? iaoe ae?iooaaeeny oaei caaaeaiiy :

c(ynoaaiiy caeaoiinoi ooaiia I-81, A-031, A-032 i ?-323 aei neioaco
iieieicniaeaeueieo i?ioa?iac;

aea/aiiy aieeao oa iiaeai? iaee?aueo aeieiai/ieo oaeoi?ia: niiniao
eoeueoeaoaaiiy, oaiia?aoo?iiai, eeneioiiai ?aaeeiia ?inoo; oa
iniaeeainoae aiineioaco aeineiaeaeoaaieo ooaiia;

aeyaeaiiy niaoeeoiee aoaeaoeaaiai oa acioiiai aeeaeaiiy i?iaeooeaioia
iieieicniaeaeueieo i?ioa?iac;

anoaiiaeaiiy ?iei ?icieo aiieiai/ii-aeoeaieo ?a/iaei : aoeneio oa
aiaa?aeiio, aeno?aeoo ?inoeia ea?oiiei, ni?aiai ai?ioia, yei
aeiaeaaaeeny aei aeeaeeueiiai na?aaeiaeua;

aeineiaeaeaiiy iiaeeeainoi ae?iuoaaiiy i?iaeooeaioia ia aeaoaaeo oa
aeinooiieo noano?aoao;

aeyaeaiiy aeeiaiiee ?inoo, aiineioaco i eiiaoe/ieo ia?aiao?ia i?e
eoeueoeaoaaiii ooaiia ia ?iciiiaiioieo aeeaeeueieo na?aaeiaeuao.

Iaoeiaa iiaecia ?iaioe. Aeineiaeaeaii iiiaae 120 na?aaeiaeu. Io?eiaii
iiai aiaeiiinoi i?i ?iaaiue iieieicniaeaeueii? aeoeaiinoi e ?inoo ooaiia
ia oeeo na?aaeiaeuao. Iiaeia?aii iioeiaeueii iieacieee eeneioiiai oa
oaiia?aoo?iiai ?aaeeiia eoeueoeaoaaiiy, oiia noa?eeicaoei? na?aaeiaeu,
niinia aa?aoei?. Aia?oa aecia/aii eiinoaioe iiiicaoei?, yei
eiio?ieththoue aeoeaiinoue iieieicniaeaeueieo i?ioa?iac o eiaeiiai c
aeineiaeaeaiieo ooaiia i oaiaeaioe?? ciiithaaiiy iieieicniaeaeueii?
aeoeaiinoi oa eeneioiinoi na?aaeiaeua o aeeiaieoei.

Iiae /an aea/aiiy acioiiai oa aoaeaoeaaiai aeeaeaiiy aia?oa aeyaeaii
aeaiaa?oeiiee oa?eoa? neioaco oa?iaioo c iieieicniaeaeueieie
aeanoeainoyie o H. laricinus ia aeaea?aeao aoaeaoeth, yei eaaei
canaiththoueny. Aeiaaaeaii, ui oieueee i?enooiinoue aieea o na?aaeiaeui
eoeueoeaoaaiiy ni?ey? aeoeaiiio neioaco iieieicniaeaeueieo i?ioa?iac.
Oeyoii iiaeai?o eiioeaio?oeie oeoe?ia o aeeaeeueiiio na?aaeiaeui
aeinyaiooi caieueoaiiy aeoeaiinoi iieieicniaeaeueiiai oa?iaioo oa
caieueoaiiy eiai aeoiaeo a /enoiio aeaeyaei a 5 ?acia. Na?aaeiaeua c
iiaeaeuaiei aiinoii oeoe?ia caiaoaioiaaii o Aea?aeiaoaioi Oe?a?ie.

Aeei?enoaiy iaoiaeia iaoaiaoe/iiai ieaioaaiiy aeicaieeei aeieeaaeii
aea/eoe oa?aeoa? caeaaeiinoi iieieicniaeaeueii? aeoeaiinoi e ?inoo H.
laricinus aiae eiioeaio?aoe?? iiei/ii? ne?iaaoee oa ieaiiai nonea.
Ae?aoiaaii ooieoe?? aeathoue iiaeeeainoue i?iaiicoaaoe iiaaaeiieo
i?iaeooeaioo o aoaeue-yeie oi/oei i?inoi?o, iaiaaeaiiai ?iaiyie
oaeoi?ia. Caaaeyee oeueiio aoei iiaeia?aii iaee?aua na?aaeiaeua
eoeueoeaoaaiiy, yea iinoeoue iiei/io ne?iaaoeo oa ieaia nonei. Ia oea
na?aaeiaeua io?eiaii ?ioaiiy i?i aeaea/o iaoaioo ia aeiaoiae.

C(yniaaii, ui oni aeineiaeaeoaaii ooaie H. laricinus iathoue caeaoiinoue
ooai?thaaoe iicaeeioeiii i?ioa?iace c aenieith iieieicniaeaeueiith
aeoeaiinoth. Aeiaaaeaia aeniea ieanoe/iinoue ooaiia. Eoeueoo?a ooaia
A-032 H. laricinus caiaoaioiaaia o Aea?aeiaoaioi Oe?a?ie, ye aeoeaiee
i?iaeooeaio i?ioa?iac iieieicniaeaeueii? ae??. Oni aeineiaeaeaii ooaie
ia aeyaeee ieiaeiiioaiiy o eoeueoo?i.

I?aeoe/ia oeiiiinoue ?iaioe. Ooaie H. laricinus iiaeia aeei?enoiaoaaoe
ye i?iaeooeaioe iieieicniaeaeueieo oa?iaioia, ui iiaeooue ciaeoe
canoinoaaiiy o ae?iaieoeoai ne?ia.

Iiaeeeainoue eoeueoeaoaaiiy H. laricinus ia aeaoaaeo noano?aoao
aeicaiey? c?iaeoe i?ioean ae?iaieoeoaa iieieicniaeaeueiiai oa?iaioo
aieueo aeaoaaoei, oiio ui iiei/ia ne?iaaoea, yea ? aiaeoiaeii
ae?iaieoeoaa iiei/ieo i?iaeoeoia, cie?aia ne?o, iiaea aiae?aco
aeei?enoiaoaaoenue aeey io?eiaiiy iieieicniaeaeueiiai oa?iaioo.

Aiaenooiinoue ieiaeiiioaiiy o eoeueoo?i oa aeniea oiciieiai/ia
ieanoe/iinoue i?iaeooeaioia oa?aeoa?eco? ?o ye aaaeeeai ia(?eoe
aiioaoiieia??, a?aoiaoth/e oa, ui eenei oa neaaieenei i?ioaieioe/ii
i?aia?aoe o ian acaaaei ia ae?iaeythoueny.

Ca(ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie.

I?aaenoaaeaii aeineiaeaeaiiy — oea /anoeia ?iaioe, ui i?iaiaeeoueny ia
eaoaae?i oiciieia?? ?ineei Aeiiaoeueeiai aea?aeaaiiai oiiaa?neoaoo a
?aieao aea?aeathaeaeaoii? oaie ? 97-1aa/16 “Iaea?aeaoe i ai?iaoaaoe iiai
aenieiaeoeaii ooaie aaceaeiaeueieo a?eaia i ?ic?iaeoe ia ?o iniiai
oaoiieiaith ae?iaieoeoaa oeiiieo aiii?aia?aoia i oa?/iaeo i?iaeoeoia”,
yea eii?aeeio?oueny ieaiii iaoeiai-aeineiaeieo oai Iiiinoa?noaa inaioe
Oe?aiie.

Iniaenoee aianie caeiaoaa/a. ?iaioa aeeiiaia iniaenoi aaoi?ii iiae
ea?iaieoeoaii aeie.aiie. iaoe, i?ioani?a, N.O. Iaa?ooeueeiai i aeie.
aiie. iaoe, aeioeaioa I.I. Aieei. Iniaenoa o/anoue aeena?oaioa iieyaaea
o ieaioaaiii oa i?iaaaeaiii eaai?aoi?ieo aeineiaeaeaiue, aiaeici,
ocaaaeueiaiii iaea?aeaiieo aenia?eiaioaeueieo aeaieo, niianoaaeaiii io c
eioa?aoo?ieie iaoa?iaeaie. Aaoi? i?eiineoue ue?o aaey/iinoue anii
niia?iaioieeai eaoaae?e oiciieiaii ?ineei Aeiiaoeueeiai aea?aeaaiiai
oiiaa?neoaoo.

Ai?iaaoeiy ?iaioe. I?i iniiaii iieiaeaiiy aeena?oaoe?? aeiiiaiaeaeiny ia
iiaeia?iaeiie eiioa?aioe??, i?enay/aiie i?iaeaiai ei?aiaaeo oa eiieaaeo
aieeae (O?aioeiy, 1997), iiaeia?iaeiie eiioa?aioe?? “Iaoiaeieiae/aneea
iniiau iiciaiey iniaaiiinoae a?eaia — i?iaeooeaioia oeceieiae/anee
aeoeaiuo aauanoa e ieuaauo i?iaeoeoia” (Aeiiaoeuee, 1997), eiioa?aioe??
iieiaeeo a/aieo “Aeooaeueii i?iaeaie oiciieia?? ?ineei oa aaiaoeee”
(Ee?a, 1996), O c(icaei Oe?aiinueeiai aioaii/iiai oiaa?enoaa (Iieoaaa,
1997), ia iaoeiaeo eiioa?aioeiyo Aeii AeO (Aeiiaoeuee, 1994-1997),
Anaoe?aiinueeeo nooaeaionueeeo eiioa?aioeiyo (Aeiiaoeuee, 1992-1997).

O caaa?oaiiio aa?iaioi aeena?oaoeiy aeiiiaiaeaeanue ia caniaeaiiyo
eaoaae?e oiciieia?? ?ineei Aeii AeO.

Ioaeieaoe??. Ca iaoa?iaeaie aeena?oaoe?? iioaeieiaaii 11 iaoeiaeo
i?aoeue: 4 noaooi a iaoeiaeo aeo?iaeao, 2 iaoaioe i 5 oac eiioa?aioe??.

No?oeoo?a oa ianya ?iaioe. Aeena?oaoeiy aeeeaaeaia a 150 noi?iieao oa
neeaaea(oueny c anooio, 7 ?icaeieia, aeniiaeia, ?aeiiaiaeaoeie, nieneo
eioa?aoo?e ( 198 aeaea?ae ). I?aoey iethno?iaaia 28 oaaeeoeyie oa 54
iaethieaie, ia( 5 aeiaeaoeia.

CIINO ?IAIOE

Iaeyae eioa?aoo?e

Iaeyae eioa?aoo?e neeaaea?oueny ic 3 ?icaeieia, a yeeo aenaioeaii
ieoaiiy ?aaoeyoeii neioaco iicaeeioeiieo oa?iaioia iie?ii?aaieciia,
iiaeaia eeaneoieaoeiy i?ioaiiac oa i?eioeeie canoinoaaiiy io o
i?iieneiainoi, a oaeiae ioa?aeoa?eciaaii i?ioean oa?iaioii? eiaaoeyoei?
iieiea.

Ia(?eoe i iaoiaee aeineiaeaeaiue

Ia(?eoaie iaoeo aeineiaeaeaiue aoee ooaie H. laricinus I-81, A-031,
A-032 i ?-323. Aeei?enoiaoaaeenue /enoi eoeueoo?e, aeaeieaii c ieiaeiaeo
oie ia eaoaae?i oiciieia?? ?ineei AeiiAeO. Ooai A-032 oa I-81
caaeaiiiiaaii o eieaeoeith Iinoeoooo Aioaiiee ii. I.A. Oieiaeiiai IAI
Oe?aiie. Ia ooai I-81 H. laricinus niia?iaioieeaie eaoaae?e io?eiaii
iaoaio Oe?aiie ( Aieei, Iaa?ooeeee, Ie?ioie/aiei oa ii., 1994 ).

A?eae ae?iuoaaee ia ?iaeeeo aeeaeeueieo na?aaeiaeuao i?e oaiia?aoo?i
300N (Aieei,1996). Iniiaiei na?aaeiaeuai aoei aetheici-iaioiiia c
aeiaeaoeii iiia?aeueieo ?a/iaei (Oaaei?iaa, Oea?eia, 1974). Iiieoethaaee
na?aaeiaeua 7-ie aeiaiaei iioeaei?i, yeee c?inoaa ia nonei-aaa?i.

Iieieicniaeaeueio aeoeaiinoue (ICA) aecia/aee ca iaoiaeii Eaaai oa Ioeai
(Kawai, Mukai, 1970). ?ic?aooiie ICA i?iaiaeeee caiaeii c oi?ioeith:

40 . 100 . E

INAEO= ––––––––––– iae/ie,

I

aea: E — eiaoioei?io ?acaaaeaiiy eoeueoo?aeueiiai oieueo?aoo; I — /an,
ca yeee ci 10 ie iieiea i?e aeiaeaaaiii 1 ie eoeueoo?aeueiiai oieueo?aoa
ooai?th(oueny uieueiee naonoie, o oaeeeiao; 40 — na?aaeiie /an cniaeaiiy
iieiea i?e ae?iaieoeoai ne?ia, o oaeeeiao.

Eeneioiinoue na?aaeiaeua ?a?no?oaaee iioaioeiiiao?e/iei iaoiaeii,
iaeiie/aiiy aiiiane — aaaiaei (Iaoa?ao?aneee, 1966). Caeeoeiao
eieueeinoue oeoe?ia aecia/aee ca iaoiaeii Oaaoeiaa oa Eoe(yiaiei
(A?iaeia, 1987), a eieueeinoue aiiiieeneio — oioiiao?e/iei iaoiaeii ca
aecia/aiiyi aiiiiiai acioo (A?iaeia, 1987). Aiino aieeo aecia/aee ca
iaoiaeii A?aaeoi?aea (Bredford, 1976).

Iaea?aeaiiy oa?iaioiiai i?aia?aoo ne/oaeii? ae?? caeienithaaee oeyoii
aeiaeaaaiiy o eoeueoo?aeueio ?iaeeio ni?/aiieeneiai aiiiith o
eieueeinoi, yea aiaeiiaiaea? 80% iane/aiiy (Aieei, 1996).

Noaoenoe/io ia?iaeo ?acoeueoaoia caeienithaaee ca aeiiiiiaith
iaeiioaeoi?iiai aeenia?nieiiai aiaeico ii?iaiyiiy na?aaei?o a?eoiaoe/ieo
aaee/ei iaoiaeii iiiaeeiieo ii?iaiyiue ca e?eoa?iyie Aeoieaia oa
Ouethee, oa e?eoa?ith Nouethaeaioa (Eaeei, 1980; Iaa?ooeeee, Oeeue/aeia,
1984).

Iioeiicaoeith na?aaeiaeua c ieaiei noneii oa iiei/iith ne?iaaoeith
caeienithaaee ca aeiiiiiaith iiaiiai oaeoi?iiai aecia?eiaioo IOA — 52
(Iaeneiia, Ieiaiiaa, A?a/ooeeia, 1976). Aiaeic aeaieo i?iaiaeeee caiaeii
c aeai?eoiii E?eona. Ca aeiiiiiaith eiii(thoa?ieo i?ia?ai caeienithaaee
iiaoaeiao iiaa?oiae aiaeaoeo.

Oni yenia?eiaioe aeeiioaaeeny iaiaio iiae o 3-o iiaoi?iinoyo.

?acoeueoaoe aeineiaeaeaiue oa io iaaiai?aiiy

1.Oiceei-oiii/ii oiiae eoeueoeaoaaiy Hirschioporus laricinus

A ?iaioi aoei i?iaaaeaii aea/aiiy aieeao oaiia?aoo?e ciaiioiueiai
na?aaeiaeua, ?I, aa?aoe??, a oaeiae ?aaeeiia coa?eeicaoe?? na?aaeiaeua
ia ?iaaiue ICA oa iaeiie/aiiy aiiiane ooaiaie H. laricinus.

Anoaiiaeaii, ui ciiia oaiia?aoo?e a iaaeao 28 0N-32 0N ia aiaeaeaa?oueny
ia oa?aeoa?i neioaco iieieicniaeaeueiiai oa?iaioa o aeineiaeaeoaaiee
ia?iiae (30 aeia) (?en.1), a oaeiae ia iaeiie/aiii aiiiane. Aeey
eoeueoeaoaaiiy H. laricinus iiaeia aeei?enoiaoaaoe oaiia?aoo?o 28 0N-32
0N.

?en. 1. Iieacieee ICA ooaio I-81 H. laricinus iiae /an eoeueoeaoaaiiy

a ?icieo oaiia?aoo?ieo ?aaeeiao.

C?iaeaii i?eiouaiiy, ui ei?ioei/aniaa aeiy iiaeaeuaieo oa cieaeaieo
oaiia?aoo? ia iioeaeie H. laricinus ia?aae ?a?no?aoei?th iieacieeia ICA
oa aiiiane aoaea aeeeeeaoe ii?ooaiiy iaiiai?iieeiinoi iaia?ai oa
iaiiiieo i?ioeania o eeioeiao, ui i?eaaaea aei caieueoaiiy aeoiaeo
oa?iaioo a eoeueoo?aeueio ?iaeeio.

Aia?oa anoaiiaeaii, ui aeiy oaiia?aoo? 40 0N i 45 0N i?ioyaii aeaio
aiaeei aeeeeea? iiaeaeuaiiy aeoeaiinoi oa?iaioo. Oaiia?aoo?a 450N
caieueoo? ICA a 3,5 ?acia, a oaiia?aoo?a 40 0N — o 1,9 ?acia. Aeiy
oaiia?aoo?e 550N i?oyaii aeaio aiaeei cieaeo? aeoeaiinoue oa?iaioo.
Iiiai?ii, iieieicniaeaeueia i?ioaiica H. laricinus ia oa?iinoaaieueia,
ye aieueoinoue a?eaieo i?ioa?iac. Aieea cieaeaiieo oaiia?aoo? oaeiae
iiaeaeuo? ICA.

Ii?iaiyeueia aea/aiiy oa?iaioaoeaiie aeoeaiinoi H. laricinus a oiiaao
aeeaeiiiai i iiaa?oiaaiai eoeueoeaoaaiiy aeaco?, ui o ?icii n?iee
eoeueoeaoaaiiy ia?aiiooaaiiy ii-?iciiio aiaeaeaa?oueny ia ICA (?en.2).
C(yniaaii, ui i?e eiiiaoiie oaiia?aoo?i e?aua caeienithaaoe
eoeueoeaoaaiiy aeeaeiiei niiniaii.

?en.2. Iieieicniaeaeueia aeoeaiinoue H. laricinus iiae /an ae?iuoaaiiy

iiaa?oiaaei oa aeeaeiiei niiniaii

Aeineiaeaeaiiy eeneioiiai ?aaeeio eoeueoeaoaaiiy i?iaeooeaioa o
aeeiaieoei aeicaieeee c(ynoaaoe, ui ani aeineiaeaeoaaii ooaie
i?yaeythoue caeaoiinoue aei ?aaoethaaiiy eeneioiinoi na?aaeiaeua. Iiae
/an ae?iuoaaiiy i?iaeooeaioia ia na?aaeiaeui c ?I 2,5 iieieicniaeaeueii?
aeoeaiinoi iaanoaiiaeaii. Oa?aeoa?ii oa, ui iioeiaeueii cia/aiiy ?I aeey
iaeiie/aiiy aiiiane oa neioaco iieieicniaeaeueiiai oa?iaioo ia
caiaathoueny, ui naiae/eoue i?i aeioeieueiinoue aeei?enoaiiy
eiiaiiiaaiiiai eeneioiiai ?aaeeio: nii/aoeo iiaeo?eioaaoe eeneioiinoue,
yea ni?ey? ?inoo a?eaia, a iioii — oa?iaioaoe??. Iioeiaeueiith
eeneioiinoth na?aaeiaeua aeey ?inoo ooaiia I-81, A-031, A-032 i ?-323 ?
cia/aiiy ?I 5,0 i aeua. Aeey aiineioaco iieieicniaeaeueieo oa?iaioia
aeoiaeia cia/aiiy eeneioiinoi iiaeiii neeaaeaoe ?I 3,5.

Eiioeaio?aoeiy iiiia aiaeith iiaea ?aaoethaaoe iiiiee noai noano?aoo.
Ciiia eeneioii? ?iaiiaaae aei cia/aiue, yei aeoiaeyoue ca iaaei cia/aiiy
?Ea, aeeeeea? ciiio eiioeaio?aoe?? noano?aoo i oaeaeeinoue neioaco
oa?iaioo.

Niinia iiaeaiiy aenia?eiaioaeueieo aeaieo o eiaa?eoii/ieo eii?aeeiaoao
aea? ciiao aecia/eoe ?Ea. Ia ?en. 3 o eiaa?eoii/ieo eii?aeeiaoao iiaeaii
eiiaoeeo neioaco iieieicniaeaeueiiai oa?iaioa ooaiii I-81 H. laricinus o
caeaaeiinoi aiae ?I eoeueoeaoaaiiy. Caiaeii c oai?i?th ae?? oa?iaioia —
i?e?aeiaiiy iaeiiai i?ioiio aei aeoeaiiai oeaio?o aiaeia?aaea?oueny o ?I
caeaaeiinoi oaeaeeicoi o eiaa?eoii/ieo eii?aeeiaoao ciiiith oaiaaino
eooa iaoeeo aeioe/ii? aei aioie neaii?ae ia iaeeieoeth. Ineieueee
ia?aoiae oaiaaino eooa iaoeeo ( ia ?en. 3 aiae (1,38( aei 0, oi neioac
iieieicniaeaeueii? i?ioa?iace, ca iaoeie aeaieie, eiio?ieth?oueny
iaeii?th iiiiaaiiith a?oiith c ?Ea iaioa iiae 3,5 oa iioith c ?Eb
aeecueei 3,85.

Eiioeaio?aoeiy iiiia aiaeith o ciaiioiueiio na?aaeiaeui e ona?aaeeii
eiiia?oiaioo, aea neioaco?oueny oa?iaio, iia(ycaii eiiieiei
niiaaiaeiioaiiyi:

H+aioo? = ( / ( . H+ciai ,

aea ( i ( — eiiaoe/ii eiaoioei?ioe (Aa?oieiiaaa, Eaethaeiue, 1990).

?en. 3. Oai?aoe/ia caeaaeiinoue eiaa?eoia (oya aiae ?I, ooaia I-81

H. laricinus.

Caiaene, eiee ( ( ( (?en. 3), oi iiaeia c oaiaaiaiinoth noaa?aeaeoaaoe,
ui o eeioeiao ooaio M-81 H. laricinus iiaeo?eio?oueny aieueo eenea
na?aaeiaeua. Oaae naia noino?oueny e iioeo ooaiia.

Oaeei /eiii aeoeaiinoue iieieicniaeaeueii? i?ioa?iace iiae /an
eoeueoeaoaaiiy ia aetheici-iaioiiiiio na?aaeiaeui aeey on?o ooaiia
eiio?ieth?oueny iaeiaeiai — aeaiia iiiiaaiieie a?oiaie. Acaaaei ?I
ciaiioiueiai na?aaeiaeua 3,5 iaeeieoeue aoiaeeoue a oni ieaoi
eiaa?eoii/iiai i?ioieth ooai?aiiy iieieicniaeaeueiiai oa?iaioo ooaiia I.
laricinus i iiaea aooe aecia/aia, ye iioeiaeueia eiioeaio?aoeiy iiiia
aiaeith aeey neioaco iieieicniaeaeueii? i?ioa?iace.

2. Eoeueoeaoaaiiy Hirschioporus laricinus ia na?aaeiaeuao c

aoaeaaiaeaie

Iiae /an eoeueoeaoaaiiy ooaio A-032 ia na?aaeiaeuao c ?icieie oeoe?aie
aeyaeaii aeaiaa?oeiiee oa?aeoa? neioaco iieieicniaeaeueieo i?ioaac ia
na?aaeiaeuao c aetheicith (?en. 4), e?ioiaeai oa iioeiiii, yei eaaei
niiaeeaathoueny i?iaeooeaioii. O aeeiaiioei iaeiie/aiiy aiiiane ia
na?aaeiaeuao c oeeie oeoe?aie aiaeii/a?oueny aeiaoene/iee oa?aeoa? ?inoo
(?en. 5). Noiaea ea?oeia ?inoo i neioaco oa?iaioo niinoa?iaa?oueny
oaeiae i?e eoeueoeaoaaiii ooaio ?-323 ia naoa?ici oa iaeueoici.

?en. 4. Aeeiaiiea iieieicniaeaeueii? aeoeaiinoi ooaiia

laricinus ia na?aaeiaeui c aetheicith.

?en. 5. Aeeiaiiea iaeiie/aiiy aiiiane ooaiaie H. laricinus

ia n?aaeiaeui c aetheicith.

O oeeo aeiaaeeao ia? iinoea eaoaaieioia ?ai?aniy, eiee aeniea
eiioeaio?aoeiy oeoe?ia ia ii/aoeiaiio aoaii eoeueoeaoaaiiy no?eio?
neioac oa?iaioia, yei aneiieththoue iioa aeaea?aei aeeaeaiiy. Ciaioaiiy
eieueeicoi oeoe?o aeicaiey? i?aaiicio i?iaeoeoaaoe iiao oa?iaioio
nenoaio, ui aoaeo? ae?oaee noano?ao, iiaeeeai i?iaeoeoe aeaniiai
iaoaaieicio, aai aeaea?aea acioiiai aeeaeaiiy.

Uiaei ooaio A-031, iiaeia aeiaanoe, ui iiney aeinyaiaiiy aenieeo
cia/aiue ICA o 10-oe aeiaiaiio aioei aiia iaei ciiith?oueny aei eiioey
ia?iiaeo eoeueoeaoaaiiy.

Oaeei /eiii, iiaeia neacaoe, ui ani aea/aii oeoe?e, e?ii a?aaiiice,
aeia?a aeei?enoiaothoueny ooaiaie H. laricinus aeey neioaco
iieieicniaeaeueieo i?ioa?iac.

Eiiaoe/ii iieacieee ?inoo oa ICA ?iciyoueny (oaae.1). sse i?aaeei, aeey
ooaiia A-031 e A-032 iaeneiaeueia oaeaeeicoue iiaeaeuaiiy ICA aeua, iiae
oaeaeeinoue caieueoaiiy nooi? ?a/iaeie. Aeey ooaio ?-323 — iaaiaee. *an
iiaeai?iiy iieacieeia aiiiane ia?aaeuo? /an iiaeai?iiy iieacieeia ICA.
Iiaeeaiaeoaeueiei aeey eiaeiiai na?aaeiaeua oa oeoe?o ? /an iaeaeuiai
aeiiiii/iiai eiaoioei?ioo eoeueoo?e. Ia na?aaeiaeui c a?aaiiicith
aeiiiii/ii eiaoioei?ioe on?o a?eaia aeui, iiae ia na?aaeiaeuao c iioeie
oeoe?aie.

Oaaeeoey 1

Eiiaoe/ii iieacieee ?inoo i aiineioaco iieieicniaeaeueiiai oa?iaioo iiae
/an eoeueoeaoaaiiy ooaiia H. laricinus ia na?aaeiaeui c aetheicith.

Ooai

Iaene-

iaeueia

oaeaeeinoue

?inoo

(max, aiae-1

*an iiae-

ai?iiy

iieacieeia aiiiane

g, aiae

Iaene-

iaeueia

oaeaeeinoue

ICA

(max, aiae-1

*an iiae-

ai?iiy

iieacieeia

ICA

g, aiae

Aeiiiii-

/iee

eiaoi-

oei?io

Y

Iaoaai-

ei/iee

eiaoi-

oei?io

qmax, aiae-1

A-031

A-032

?-323

0,0139

0,0179

0,0149

49,86

38,72

46,51

0,0212

0,0235

0,0128

32,69

29,49

54,14

0,450

0,588

0,980

0,046

0,030

0,029

Aeineiaeaeaiiy iieacaee, ui iiaeai?ii aeaea?ae aoaeaoeaaiai aeeaeaiiy
iiaeia ?aaoethaaoe aiaeiiaiaeiinoue i?ioeania ?inoo oa ciiie ICA ye o
aie i?enei?aiiy neioaco iieieicniaeaeueiiai oa?iaioo, oae i a aie
caieueoaiiy oaiaeeicoi ?inoo. Iaeneiaeueii oaeaeeicoi ?inoo oa ICA
aeyaeythoueny ia ?aii?o noaaeiyo ?icaeoeo eoeueoo?e.

O aeiaaeeo iiaeaeuaiiy eiioeaio?aoe?? oeoe?ia o aeaa ?ace a ii?iaiyiii c
noaiaea?oiei na?aaeiaeuai aiaeaoaa?oueny caieueoaiiy ICA ia on?o
na?aaeiaeuao c i?ioyaii 30-aeiaiaiai eoeueoeaoaaiiy. Iiaeaeuaiiy
eiioeaio?aoe?? aetheice, iaeueoice i e?ioiaeth aeeeeea? cieaeaiiy
cia/aiue aiiiane iai?eeiioei ia?iiaeo eoeueoeaoaaiiy. Oea, iaaooue,
iia(ycaii ci caeaoiinoth oeeo oeoe?ia o cacia/aiie eiioeaio?aoe??
noeioethaaoe neioac oa?iaioia, ui aeeeeeathoue ?oeioaaiiy eeioeiieo
iaieiiie iioeaeith H. laricinus.

Iiaeaeuaiiy aiinoo oeoe?ia o na?aaeiaeui eoeueoeaoaaiiy o 2 ?ace
aeioeieueii, oiio ui oea caieueoo? aeoiae iieieicniaeaeueiiai oa?iaioo o
5 ?acia i iiaeaeuo? eiai aeoeaiinoue

3. ?ieue aeaea?ae acioo aeey eoeueoeaoaaiiy ooaiia

Hirschioporus laricinus

Iiae /an aeineiaeaeaiiy aeaea?ae acioiiai aeeaeaiiy ( NH4NO3, (NH2)2CO,
(NH4)2SO4, CH3COONH4, NaNO3, KNO3 oa iaioii ) ?ino i?iaeooeaioia
aeyaeaii ia an?o ?a/iaeiao, ui iinoyoue acio (?en.6). Aeia?a
aeei?enoiaothoueny aeey iaeiie/aiiy aiiiane H. laricinus na/iaeia e
ni?/aiieeneee aiiiie, aea iieacieee aiiiane a iaio aeiaaeeao ieae/i,
iiae ia na?aaeiaeui, yea iinoeoue iaioii.

?en.6. Iaeiie/aiiy aiiiane ooaiii A-031 iiae /an ae?iuoaaiiy ia

?icieo aeaea?aeao acioo.

Noiniaii ICA iiaeia aiaecia/eoe, ui iio?ao iao?ith oa eaeith, a oaeiae
aoeaoao aiiiith oeieeii iiaiaothoue neioac iieieicniaeaeueiiai oa?iaioo.
Ia ?aooi aeaea?ae ?iaaiue ICA ieae/ee, iiae ia na?aaeiaeui c iaioiiii a
5-27 ?acia. Oaeei /eiii, ooaie H. laricinus aeey ?inoo oa neioaco
iieieicniaeaeueiiai oa?iaioo e?aua aeei?enoiaothoue acio iaioiio.
Iio?ao- iii iai?eaeaoiee aei eoeueoeaoaaiiy ooaiia H. laricinus.

Aea/aiiy aieeao aiiiieeneio: DL-o?eioioai, DL-oaiieaeaiii,
DL-eiceiiiiiaiae?ioei?eae, DL-eaeoeei, DL-iaoiiiii, L-icieaeoeei,
L-ainoeaeeiaiae?ioei?eae, DL-aaeii, L-a?aiiiiaiae?ioei?eae, DL-o?aiiii,
i?iaiaeeee oeyoii aeiaeaaaiiy ?o o na?aaeiaeua o eieueeinoi,
aeaiaaeaioiie 3 a iaioiio ii aiiiieiiio acioo.

Ia na?aaeiaeuao c o?eioioaiii, eiceiiiiiaiae?ioei?eaeii, aaeiiii oa
o?aiiiiii ICA ia aeyaeaia. Ia na?aaeiaeuao, aea aoea ICA, ?iaaiue ??
neeaaeaa aeecueei 2500 iae/ie. Aeey ?inoo H. laricinus e?aua
aeei?enoiaoaaoe eiceiiiiiaiae?ioei?eae. Aeei?enoaiiy aiiiieeneio ooaiaie
i?ioiaeeoue iaiaeiaeiai. E?aua c on?o aiiiieeneio aeei?enoiaothoueny
o?eioioai oa a?aiiii, neaaei — iaoiiiii oa aaeii.

Cia/aiiy ICA ia na?aaeiaeuao c aiiiieeneioaie ia ii?yaeie ieae/a, iiae
ia na?aaeiaeui c iaioiiii, aea cia/aiiy aiiiane ia oeeo na?aaeiaeuao
iacia/ii iinooiathoueny cia/aiiyi aiiiane ia aetheici-iaioiiiiio
na?aaeiaeui. Oea naiae/eoue, ui aiiiieeneioe aeei?enoiaothoueny
i?iaeooeaioii ia?aaaaeii aeey ?inoo i?aaiicio.

Oaeei /eiii, aeey neioaco iieieicniaeaeueiiai oa?iaioo ooaiaie H.
laricinus iaiaoiaeia iayaiinoue o na?aaeiaeui eoeueoeaoaaiiy aieeiaiai
acioo, yeee na?aae on?o aea/aieo aeaea?ae acioiiai aeeaeaiy ( oieueee o
iaioiii.

4. Eoeueoeaoaaiiy Hirschioporus laricinus ia na?aaeiaeuao c

iiei/iith ne?iaaoeith.

Aeiaeaaaiiy iiei/ii? ne?iaaoee o na?aaeiaeua eoeueoeaoaaiiy aeeeeea?
caieueoaiiy oaeaeeinoi ?inoo oa i?iaeoeoaaiiy iieieicniaeaeueiiai
oa?iaioo ((max), aei oiai ae, /ei aieueoa ne?iaaoee o na?aaeiaeui, oei
aeua (max, ioaea nei?i/o?oueny /an iiaeai?iiy iieacieeia (oaae.2).

Oaaeeoey 2

Ia?aiao?e, ui oa?aeoa?ecothoue aeeiaiieo ciiie aiiiane oa ICA

laricinus iiae /an eoeueoeaoaaiiy ia na?aaeiaeuao c iiei/iith

ne?iaaoeith.

Eieueeinoue

iiei/ii?

ne?iaaoee, %

Iieaciee Iaeneiaeueia

oaeaeeinoue, (max

aiae-1 *an iiaeai?iiy

g, aeia

40

20

10

0

ICA

aiiiana

ICA

aiiiana

ICA

aiiiana

ICA

aiiiana

0,023

0,016

0,011

0,016

0,008

0,015

0,007

0,012

1,3

1,8

2,6

1,8

3,9

1,9

4,1

2,4

Caa?oa? ia naaa oaaao i oa, ui aeiaeaaaiiy ne?iaaoee caieueoo?
o?eaaeinoue aeniiiaioeieii? oace ?inoo oa aiineioaco oa?iaioo aei 10-oe
aeia, eiee i?e ?? aiaenooiinoi aeniiiaioeieia oaca neeaaea? 5 aeia.
Neiae aiaecia/eoe, ui iiae /an aeiaeaaaiiy iaeneiaeueii? eiioeaio?aoe??
iiei/ii? ne?iaaoee o aeiaaeeo c ICA aiaeaoaa?oueny iiaeiaaeaiiy
eaa-ia?iiaeo.

Oaeei /eiii, ciiithth/e eieueeinoue iiei/ii? ne?iaaoee o na?aaeiaeui
iiaeia aeinyaoe i?enei?aiiy ye i?ioeania ?inoo, oae i i?iaeoeoaaiiy
iieieicniaeaeueieo i?ioaac o H. laricinus oa ?aaoethaaoe oaeaeeinoue
oeeo i?ioeania ca aeaniei aaaeaiiyi.

C iaoith iioeiicaoe?? aeeaeeueiiai na?aaeiaeua eoeueoeaoaaiiy H.
laricinus, yea iinoeoue iiei/io ne?iaaoeo i iiaia nonei, aoei i?iaaaeaii
na?ith iiaeaeueieo aenia?eiaioia caiaeii c iiaiei oaeoi?iei
aenia?eiaioii 52 (IOA 52). Aeey eiaeiiai aenia?eiaioo ia 5-oo oa 10-oo
aeiao eoeueoeaoaaiiy neeaaeaeeny ?iaiyiiy ?aa?an??, aiaeiiaiaeii yeei ca
aeiiiiiaith eiii(thoa?ieo i?ia?ai aoaeoaaeeny iiaa?oii aiaeaoeo aiiiane
oa ICA. IOA 52 aeicaieea iai aeieeaaeii aeineiaeeoe oa?aeoa?
caeaaeiinoi, aecia/eoe cia/aiiy ooieoe?? a aoaeue-yeie oi/oei i?inoi?o,
iaiaaeaiiai ?iaiyie oaeoi?ia.

Ui noino?oueny aaeineiiaeaiiy na?aaeiaeua, oi aeey aeeeth/aiiy aoaeoia
aenieiai ii?yaeeo oa ocaaaeueiaiiy io?eiaieo aeaieo iaie aoei ciaioaii
eieueeinoue ?iaiia oaeoi?ia aiaeiiaiaeii IOA 32. A ?acoeueoaoi
aenia?eiaioo io?eiaii aeaii, yei iienothoueny ?iaiyiiyie ?aa?an?? (1) i
(2).

y = 48,983 — 1,051o1+3,548o2 — 4,026o12 — 7,86o22-3,269o1o2 +
3,138o1o22 (1)

o = 107,36 + 1,34o1 — 12,32o2 — 8,85o12 — 24,87o22 — 7,755o1o2 (2)

?iaiyiiy (1) iieno? aeaii c ICA, ?iaiyiiy (2) — aeaii c aiiiane, aea o1
— eiaeiaaii cia/aiiy eiioeaio?aoe?? ieaiiai nonea, a o2 — eiaeiaaii
cia/aiiy eiioeaio?aoe?? iiei/ii? ne?iaaoee.

Oa?aeoa? caeaaeiinoi ICA i aiiiane aiae aeineiaeaeoaaiieo oaeoi?ia
iiaeiaiee. Iiiiiaeueii cia/aiiy ooieoe?? cnoiooi aei e?ath, a ieaoi
iioeiaeueieo cia/aiue ciaoiaeeoueny o oeaio?i iiaa?oii aiaeaoeo. Aeniei
cia/aiiy ICA ciaoiaeyoueny a iioa?aaei, aea o1 ( [ -0,2 ; 0,5 ], a o2 (
[ -0,4 ; 0,5 ] (iae. 7). Ui noino?oueny aiiiane, oi ieaoi iioeioio
iaiaaeo?oueny cia/aiiyie o1 ( [ -0,2 ; 0,4 ] oa x2 ( [-0,3 ; 0,5] .

Oaeei /eiii, aeniei cia/aiiy ICA i aiiiane aoee iaea?aeaii ia
na?aaeiaeuao, o yeeo niiaaiaeiioaiiy iiei/ii? ne?iaaoee i ieaiiai nonea
neeaaea? 1:1, aei oiai ae eieueeinoue aiaee, yea aeiaea?oueny o
na?aaeiaeua, oaeiae ? aaaeeeaith, oiio ui o aeiaaeeo aenieeo
eiioeaio?aoeie aeineiaeaeoaaieo oaeoi?ia cia/aiiy ICA oa aiiiane
ciaioothoueny. *anoea aiaee iiaeiia neeaaeaoe o?aoth /anoeio noano?aoo,
ioiae, oaeineiiaeaia na?aaeiaeua aeeth/a? iiei/io ne?iaaoeo, ieaia nonei
oa aiaeo a niiaaiaeiioaiii 1:1:1. Ca ?acoeueoaoaie oeeo aeineiaeaeaiue
Aea?aeiaoaioii Oe?aiie i?eeiyoi ?ioaiiy i?i aeaea/o iaoaioo.

5. Aaaaeaiiy o na?aaeiaeua eoeueoeaoaaiiy Hirschioporus laricinus

aiieiai/ieo aeiaeaoeia

Aaaaeaiiy aenieeo eiioeaio?aoeie aeno?aeoia ?inoeia ea?oiiei nei?i/o?
noaoeiiia?io oaco ?inoo, i?i ui naiae/eoue cieaeaiiy aiiiane iai?eeiioei
ia?iiaeo eoeueoeaoaaiiy. Aeno?aeo ?inoeia ea?oiiei iiaeaeuo? ICA oieueee
a aeniiiaioeieiie oaci ?inoo (oaa.3), i?e iiaeaeueoiio eoeueoeaoaaiii ia
an?o na?aaeiaeuao ?iaaiue aeoeaiinoi oa?iaioo ia aiae?iciy?oueny aiae
?iaiy ICA eiio?ieueiiai aetheici-iaioiiiiai na?aaeiaeua, aea i?e
iaeneiaeueiie eiioeaio?aoe?? yeno?aeoo cia/aiiy ICA ieae/a, iiae o
eiio?iei.

Oaaeeoey 3

Aieea aiinoo aeno?aeoo ?inoeia ea?oiiei ia

iieieicniaeaeueio aeoeaiinoue ooaia I-81 H. laricinus.

Aiino

aeno?aeoo

ie/e

Aie, aeiaa

5

10

15

20

Iieieicniaeaeueia aeoeaiinoue, iae/ie

00,00

61,54

124,62

189,23

5904,4

6887,2

8650,6

8335,2

15738,4

14572,5

16565,4

10051,2

17719,6

18176,1

16530,9

2118,4

15015,4

17158,0

15175,6

659,5

Aeiaeaaaiiy o na?aaeiaeua eoeueoeaoaaiiy ni?aiai a?ioia o eiioeaio?aoe??
0,5% i 1,0% noeioeth?, ye iaeiie/aiiy aiiiane, oae i aiineioac
iieieicniaeaeueiiai oa?iaioo.

Aaaaeaiiy aiaa?aeiio o na?aaeiaeua noeioeth? ICA. I?e eiioeaio?aoe?? 4
ia/e aeoeaiinoue aeua, iiae i?e 2,5 ia/e. Oaea ae caeaaeiinoue
niinoa?iaa?oueny i noiniaii aiiiane. Aeiaeaaaiiy aoeneio ia ?aii?o
noaaeiyo eoeueoeaoaaiiy cieaeo? ICA, iaeii/anii aiaeaoaa?oueny
i?eaii/aiiy ?inoo, aea iiciioa aeoeaiinoue oa?iaioo ?icei caieueoo?oueny
oa aeinyaa? ?iaiy eiio?ieueiiai aetheici-iaioiiiiai na?aaeiaeua, a
cia/aiiy aiiiane iaaioue ia?aaeuothoue cia/aiiy iioeo na?aaeiaeu.

Oaeei /eiii, aeey iiaeo?eioaaiiy aenieeo cia/aiue ICA H. laricinus
aei?aaaeaii aaaaeaiiy o na?aaeiaeua eoeueoeaoaaiiy aiaa?aeiia o
eieueeinoi 4 ia/e, yea caieueoo? ICA a 2-2,5 ?acia.

Aoenei e?aua aeei?enoiaoaaoe o eiioeaio?aoe?? 4 ia/e c iaoith
caieueoaiiy aiiiane iiae /an o?eaaeiai eoeueoeaoaaiiy.

AENIIAEE

Ia iiaenoaai aeineiaeaeaiue ?aaoeyoe?? iieieicniaeaeueieo i?ioa?iac
ooaiia Hirschioporus laricinus ( Karst ) Ryv. iiaeia c?iaeoe oaei
aeniiaee :

Oni aea/aii ooaie I-81, A-031, A-032 i ?-323 H. laricinus iathoue
caeaoiinoue i?iaeoeoaaoe iicaeeioeiii eenei i?ioa?iace c aenieith
iieieicniaeaeueiith aeoeaiinoth. Iaeaieueo aeoeaiei ? ooai A-032. Oeyoii
aeei?enoaiiy ?icieo oiciieiai/ieo oaeoi?ia iiaeeeaa ciiithaaiiy cia/aiue
iieieicniaeaeueii? aeoeaiinoi oa aeinyaiaiiy ?? aenieiai ?iaiy o
aoaeue-yeiai ic ooaiia H. laricinus.

Ciiithaaiiy oaiia?aoo?iiai ?aaeeio eoeueoeaoaaiiy o iaaeao aiae 280N aei
320N ia aiaeaeaa?oueny ia oa?aeoa?i neioaco iieieicniaeaeueieo i?ioa?iac
oa ?inoi H. laricinus. Oeae oaiia?aoo?iee ?aaeei ? iioeiaeueiei aeey
eoeueoeaoaaiiy i?iaeooeaioo. C iaoith caieueoaiiy aeoiaeo oa?iaioa
aeioeieueii ia?aae eiai aeaeieaiiyi iiaeaeaaaoe ioieiaeaeaiith aei
iecueeeo iethniaeo oaiia?aoo? ( 2-50N ) i?ioyaii aeaio aiaeei, a oaeiae
iaa?iaaiiy o iaaeao 40-45 0N i?oyaii aeaio aiaeei, aei oiai ae
iaa?iaaiiy ia? ia?aaaao.

Ooaie H. laricinus caeaoii ?aaoethaaoe ?I na?aaeiaeua, iiae /an ?o
ae?iuoaaiiy ia na?aaeiaeuao c oeoe?aie. Aeoeaiinoue iieieicniaeaeueii?
i?ioa?iace iiae /an eoeueoeaoaaiiy H. laricinus ia aetheici-iaioiiiiio
na?aaeiaeui eiio?ieth?oueny aeaiia iiiiaaiieie a?oiaie. ?I 3,5
iaeeieoeue ? iioeiaeueiith aeey ooai?aiiy iieieicniaeaeueieo oa?iaioia
ooaiaie H. laricinus. Aeey oeeo ooaiia iioeiaeueii cia/aiiy eeneioiinoi
aeey iaeiie/aiiy aiiiane (?I 5,0) oa neioaco oa?iaioa (?I 3,5) ia
caiaathoueny, ui naiae/eoue i?i iiaeeeainoue aeei?enoaiiy eiiaiiiaaiiai
?aaeeio eoeueoeaoaaiiy : nii/aoeo iiaeo?eioaaoe eeneioiinoue, yea ni?ey?
?inoo a?eaia, a iiney oeueiai oa?iaioaoe??.

Iiaeai?ii aeaea?ae aoaeaoeaaiai aeeaeaiiy iiaeia ?aaoethaaoe
niiaaiaeiioaiiy i?ioeania ?inoo oa ciiie iieieicniaeaeueii? aeoeaiinoi
ooaiia H. laricinus ye o aie i?enei?aiiy neioaco iieieicniaeaeueiiai
oa?iaioo, oae i o aie oaeaeeinoi ?inoo. Aeei?enoiaoth/e na?aaeiaeua c
eiiaiiiaaiei aeaea?aeii aoaeaoeth, iiaeia, eiee iaiaoiaeii,
neio?iiicoaaoe i?ioeane ?inoo i aiineioaco. Iaeneiaeueii oaeaeeinoi
?inoo oa aiineioaco aeyaeythoueny ia ia?oeo noaaeiyo ?icaeoeo eoeueoo?.
Aetheica, e?ioiaeue, iioeii, naoa?ica oa iaeueoica aeia?a
aeei?enoothoueny ooaiaie aeey neioaco iieieicniaeaeueiiai oa?iaioa.
Aeioeieueii aeei?enoiaoaaoe iiaeaeuaiiy eiioeaio?aoe?? oeoe?ia o 2 ?ace
o ii?iaiyiii ci noaiaea?oiei aetheici-iaioiiiei na?aaeiaeuai, ui
caieueoo? aeoiae iieieicniaeaeueiiai oa?iaioo a 5 ?acia oa iiaeaeuo?
eiai aeoeaiinoue.

H. laricinus caeaoiee aeei?enoiaoaaoe aeey ?inoo iioeaeith oni
aeineiaeaeaiii aeaea?aea acioo, aea iio?ao eaeith oa iao?ith, a oaeiae
aoeaoao aiiiith oeieeii iiaiaothoue neioac iieieicniaeaeueiiai oa?iaioa.
Ia na?aaeiaeuao c aiiiieeneioaie, NH4NO3, CH4NO2, (NH4)2SO4 ?iaaiue ICA
ieae/ee, iiae ia na?aaeiaeui c iaioiiii o 5-27 ?acia. Aeey aeoeaiiai
neioaco iieieicniaeaeueieo i?ioa?iac H. laricinus iaiaoiaeia iayaiinoue
o na?aaeiaeui eoeueoeaoaaiiy aieeiaiai acioo.

Iiaeaeuaia eieueeinoue aeno?aeoo ?inoeia ea?oiiei o aeeaeeueiiio
na?aaeiaeui iiaiao? ye ?inoiai i?ioeane, oae i neioac oa?iaioa. Aeno?aeo
?inoeia ea?oiiei noeioeth? ICA oieueee ia ia?oeo eoaiao ?inoo
eoeueoo?e.Ni?aa ai?ioii o eiioeaio?aoe?? 0,5-1,0% i?enei?th? ?ino oa
caieueoo? ICA H. laricinus. Aeiaeaaaiiy aiaa?aeiio o na?aaeiaeua o
eieueeinoi 4 ia/e caieueoo? aeoeaiinoue iieieicniaeaeueiiai oa?iaioa a
2-2,5 ?acia. Aianaiiy aoeneio a na?aaeiaeua eoeueoeaoaaiiy o eieueeinoi
4 ia/e aeicaiey? io?eiaoe aieueo aenieee aeoiae aiiiane i?iaeooeaioo.

Aaaaeaiiy iiei/ii? ne?iaaoee o na?aaeiaeua eoeueoeaoaaiiy aeeeeea?
iiaeaeuaiiy oaeaeeinoi ?inoo oa i?iaeoeoaaiiy iieieicniaeaeueiiai
oa?iaioo, a oaeiae caieueoaiiy o?eaaeinoi aeniiiaioeieii? oace ?inoo
i?iaeooeaioo.

Aeei?enoaiiy iaoiaeia iaoaiaoe/iiai ieaioaaiiy aeicaieeei aeieeaaeii
aea/eoe oa?aeoa? caeaaeiinoi ICA i ?inoo I. Laricinus aiae
eiioeaio?aoe?? iiei/ii? ne?iaaoee oa ieaiiai nonea. ?ic?aoiaaii ooieoe??
aeathoue ciiao i?iaiicoaaoe iiaaaeiieo i?iaeooeaioo a aoaeue-yeie oi/oei
i?inoi?o, yeee iaiaaeaiee ?iaiyie oaeoi?ia.

Aie iiniaiiai iioeaeith ia aiaeaeaa?oueny ia ?iaii ICA I.laricinus.

Aieiaii ioaeeeaoe?? ca oaiith aeena?oaoe?? :

Ieeeoeia I. A. Aeeyiea ?I n?aaeu ia iaeiieaiea iieieinaa?ouaathuaai
oa?iaioa ooaiiaie Hirschioporus laricinus // Iie?iaiieiai/iee aeo?iae. —
1998. — ?4.- N. 43-48.

Ieeeoeia I.A. Aeeyiea oeoiai?iiiia ia iieieinaa?ouaathuoth aeoeaiinoue e
?ino Hirschioporus laricinus. // Aaniee Aeiiaoeeiai oieaa?neoaoa. —
1998. — ?1. — N.141-144.

Ieeeoeia I.A. Aeeyiea ?I ia eeiaoeeo neioaca iieieinaa?ouaathueo
oa?iaioia ooaiiia Hirschioporus laricinus // Aanoiee i?iaeai aeieiaee e
iaaeeoeeiu. — Iieoaaa-Oa?ueeia, 1998. — ?11. — N. 18-25.

Ieeeoeia I.A., Aieei I. E., Iaa?ooeeee N.O. ?ieue enoi/ieeia acioa aeey
eoeueoeae?iaaiey ooaiia A-031 Hirschioporus laricinus // Aanoiee i?iaeai
aeieiaee e iaaeeoeeiu. — Iieoaaa-Oa?ueeia, 1998. — ?11. — N. 26-33.

Iao. 96031179. Oe?a?ia. IIE 5 N 12 N 1/38. AEeaeeueia na?aaeiaeua aeey
ae?iuoaaiiy ooaio I-81 Hirschioporus laricinus — i?iaeooeaioo
iieieicaa?oath/iai oa?iaioo / Aieei I. I., Iaa?ooeueeee N. O., Oaaeioia
I. A., Iieioiia I. I. ( Oe?a?ia ).- 1 n. Cayae. 27.03.96; Iioae.
06.02.97. — 2 n.

Iao. 97073489. Oe?a?ia. IIE N 12 N 1/38, 9/14. AEeaeeueia na?aaeiaeua
aeey ?inoo ooaio I-81 Hirschioporus laricinus — i?iaeooeaioo
iieieicniaeaeueiiai oa?iaioo / Iieioiia I. I., Aieei I. I., Iaa?ooeueeee
N. O. ( Oe?a?ia ). — 1 n. Cayae. 2.07.97; Iioae. 10.06.98. — 2 n.

Ai?enaiei I. A. (Ieeeoeia), Iaa?ooeeee N. O., Aieei I. E. Aeeyiea
?acee/iuo aeic iiei/iie nuai?ioee ia ?ino e i?iaeooee?iaaiea
iieieinaa?ouaathuaai oa?iaioa Hirschioporus laricinus // Onoie/eainoue
?anoaiee e no?anniaui yaeaieyi. AeiiAeO; Aeiiaoee,1995. — N.168-173.
Aeai. a AIOA 18.12.95, ?21 — Oe 96.

Ieeeoeia I. A. Aeeiaieea, eeiaoeea ?inoa e neioaca iieieinaa?ouaathuaai
oa?iaioa Hirschioporus laricinus // Iao. I Iaaeaeoia?iaeiie
eiioa?aioeee: Iaoiaeieiae/aneea iniiau iiciaiey aeieiae/aneeo
iniaaiiinoae a?eaia — i?iaeooeaioia oeceieiai/anee aeoeaiuo niaaeeiaiee
e ieuaauo i?iaeoeoia. — Aeiiaoee, 1997. — N.34-37.

Ieeeoeia I.A. Aeeyiea ?aaeeiia noa?eeecaoeee ia iieieinaa?ouaathuoth
aeoeaiinoue Hirschioporus laricinus // Iao. O nuacaea Oe?aeineiai
aioaie/aneiai iauanoaa. — Iieoaaa,1997. — N. 98.

Ieeeoeia I. A. Aieiieeneiou, eae enoi/iee acioa aeey eoeueoae?iaaiey
Hirschioporus laricinus — i?iaeooeaioa iieieinaa?ouaathuaai oa?iaioa //
Iao. aoc. iao/i. eiio. i?ioanni?nei-i?aiiaeaaaoaeueneiai ninoaaa ii
eoiaai iao/iienneaaeiaaoaeueneie ?aaiou. — Aeiiaoee: AeiiAO, 1997. —
N.79-82.

Ieeeoeia I. A. Aeeyiea e?eoe/aneeo oaiia?aoo? ia auoiae
iieieinaa?ouaathuaai oa?iaioa o Hirschioporus laricinus // Oac. aeiee VI
Eiio. iieiaeuo o/aiuo: Aeooaeueiea i?iaeaiu oeceieiaee ?anoaieee
aaiaoeee. — Eeaa, 1996. — N.122-124.

Iieioiia I.I. ?aaoeyoeiy aeoeaiinoi aecii?ioa?iac iieieicniaeaeueii?
ae?? ooaiia Hirschioporus laricinus (Karst.) Ryv. — ?oeiien.

Aeena?oaoeiy ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa aiieiai/ieo
iaoe ca niaoeiaeueiinoth 03.00.04 — aiioiiiy. — Ee?anueeee oiiaa?neoao
iiaii Oa?ana Oaa/aiea, Ee?a, 1999.

Aeena?oaoeiy i?enay/aia aeineiaeaeaiith aieeao ?icieo oiciieiai/ieo
oaeoi?ia: oaiia?aoo?e oa niiniao eoeueoeaoaaiiy, eeneioiinoi
na?aaeiaeua, aoaeaoeaaiai oa acioiiai aeeaeaiiy, a oaeiae aiieiai/ieo
aeiaeaoeia ia iieieicniaeaeueio aeoeaiinoue oa iaeiie/aiiy aiiiane
ooaiaie Hirschioporus laricinus I-81, A-031, A-032 i ?-323 —
i?iaeooeaioaie iieieicniaeaeueieo i?ioa?iac. A aeena?oaoe??
aeineiaeaeaii iiiaae 120 aeeaeeueieo na?aaeiaeu. Aea/aii aeeiaiieo oa
eiiaoeeo neioaco iieieicniaeaeueieo i?ioa?iac ia oeeo na?aaeiaeuao.
Oeyoii iiaeai?o eiioeaio?aoeie oeoe?ia aeinyaiooi caieueoaiiy aeoiaeo
iieieicniaeaeueiiai oa?iaioa a 5 ?acia. Aeineiaeaeaii oa?aeoa?
caeaaeiinoi iieieicniaeaeueii? aeoeaiicoi oa ?inoo a?eaa aiae
eiioeaio?aoe?? iiei/ii? ne?iaaoee oa ieaiiai nonea — aeaoaaeo oa
aeinooiieo neeaaeiaeo aeeaeeueiiai na?aaeiaeua, caaaeyee /iio aoei
iiaeia?aii iioeiaeueia na?aaeiaeua eoeueoeaoaaiiy. Aeaii ?aeiiaiaeaoe??
uiaei i?iieneiaiai eoeueoeaoaaiiy ooaiia H. laricinus. Ca iniiaieie
?acoeueoaoaie i?aoei iaea?aeaii iaoaio ia aeiaoiae oa aeai i?ii?eoaoii
aeiaiaeee.

Eeth/iai neiaa: ne/oaeii oa?iaioe, neeaaeiai na?aaeiaeua, iioeiicaoeiy,
ii-

aa?oiy aiaeaoeo, iieieicniaeaeueia aeoeaiinoue

Ieeeoeia I. A. ?aaoeyoeey aeoeaiinoe yecii?ioaeiac iieieinaa?ouaathuaai
aeaenoaey ooaiiia Hirschioporus laricinus ( Karst. ) Ryv. — ?oeiienue.

Aeenna?oaoeey ia nieneaiea iao/iie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03. 00. 04 — aeioeiey. — Eeaaneee oieaa?neoao
eiaie Oa?ana Oaa/aiei, Eeaa, 1999.

Aeenna?oaoeey iinayuaia aii?inai enneaaeiaaiey aeeyiey ?acee/iuo
oeceieiae/aneeo e aeioeie/aneeo oaeoi?ia: oaiia?aoo?u e niiniaa
eoeueoeae?iaaiey, eeneioiinoe n?aaeu, oaea?iaeiiai e acioiiai ieoaiey, a
oaeaea aeieiae/aneeo aeiaaaie ia iieieinaa?ouaathuoth aeoeaiinoue e
iaeiieaiea aeiiannu ooaiiaie Hirschioporus laricinus I-81, A-031, A-032
e ?-323 . H. laricinus — iiaue ia?niaeoeaiue i?iaeooeaio
iieieinaa?ouaathueo i?ioaeiac, eioi?ua iiaoo eniieueciaaoueny a
i?iecaiaenoaa nu?ia eae caiaieoaee nu/oaeiiai oa?iaioa. Iieo/aiea
oeaiiuo a?eaiuo iaoaaieeoia naycaii n iioeiecaoeeae oneiaee
eoeueoeae?iaaiey a oneiaeyo eneonnoaaiiuo eoeueoo? e yaeyaony iaeiie ec
iniiaiuo caaea/, o?aaothueo ?aoaiey. A ?aaioa enneaaeiaaii aieaa 120
ieoaoaeueiuo n?aae. Eco/aia aeeiaieea e eeiaoeea neioaca
iieieinaa?ouaathueo i?ioaeiac ia yoeo n?aaeao. Auyniaii, /oi ooaiiu H.
laricinus niiniaiu ?aaoee?iaaoue ?I n?aaeu eoeueoeae?iaaiey,
iioeiaeueiua cia/aiey eeneioiinoe aeey iaeiieaiey aeiiannu e
i?iaeooee?iaaiey iieieinaa?ouaathuaai oa?iaioa ia niaiaaeatho. Eciaiaiea
oaiia?aoo?u a i?aaeaeao io 280N aei 320N ia neacuaaaony ia oa?aeoa?a
neioaca iieieinaa?ouaathuaai oa?iaioa e ?inoa H.laricinus.
N?aaieoaeueiia eco/aiea oa?iaioaoeaiie aeoeaiinoe i?iaeooeaioa a oneiaey
aeoaeiiiai e iiaa?oinoiiai eoeueoeae?iaaiey iieacuaaao, /oi a ?aciua
n?iee eoeueoeae?iaaiey ia?aiaoeaaiea ii-?aciiio neacuaaaony ia
iieieinaa?ouaathuae aeoeaiinoe, i?e eiiiaoiie oaiia?aoo?a eo/oa
eniieueciaaoue eoeueoeae?iaaiea aeoaeiiui niiniaii. Enneaaeiaaiey ?iee
?acee/iuo oaeaaiaeia iieacaee, /oi iiaeai?ii enoi/ieea oaea?iaeiiai
ieoaiey ooaiiia H.laricinus iiaeii ?aaoee?iaaoue nii?yaeaiiinoue
i?ioeannia ?inoa e iieieinaa?ouaathuae aeoeaiinoe, eae a noi?iio
oaaee/aiey aeoeaiinoe iieieinaa?ouaathuaai oa?iaioa oae e a noi?iio
oaaee/aiey nei?inoe ?inoa i?iaeooeaioa. Iooai iiaeai?a eiioeaio?aoeee
naoa?ia aeinoeaiooi oaaee/aiea auoiaea iieieinaa?ouaathuaai oa?iaioa a 5
?ac. I?e eco/aiee ?iee enoi/ieeia acioiiai ieoaiey (NH4NO3, (NH2)2CO,
(NH4)2SO4, CH3COONH4, NaNO3, KNO3, iaioiia e 10-oe aieiieeneio:
DL-o?eioioai, DL-oaieeaeaiei, DL-eeceiiiiiaeae?ioei?eae, DL-eaeoeei,
DL-iaoeiiei, L-ecieaeoeei, L-aenoeaeeiaeae?ioei?eae, DL-aaeei,
L-a?aeieiaeae?ioei?eae, DL-o?aiiei ) auyniaii, /oi H. laricinus
eniieuecoao aeey ?inoa ieoeaeey ana enneaaeiaaiiua enoi/ieee
i?aaie/aneiai e iai?aaie/aneiai acioa, ii aeey aeoeaiiai neioaca
iieieinaa?oeaathueo i?ioaeiac iaiaoiaeeii iaee/ea a n?aaea
eoeueoeae?iaaiey aaeeiaiai acioa, eioi?ue n?aaee anao eco/aiiuo aauanoa
niaea?aeeony oieueei a iaioiia. Aaaaeaiea iiei/iie nuai?ioee a
eoeueoo?aeueioth n?aaeo H. laricinus aucuaaao iiauoaiea nei?inoe ?inoa e
i?iaeooee?iaaiey iieieinaa?ouaathuaai oa?iaioa, a oaeaea oaaee/aiea
aeeeoaeueiinoe yeniiiaioeeaeueiie oacu ?inoa. I?e eniieueciaaiee
iaoiaeia iaoaiaoe/aneiai ieaie?iaaiey aeaoaeueii eco/ai oa?aeoa?
caaeneiinoe iieieinaa?ouaathuae aeoeaiinoe e aeiiannu i?iaeooeaioa io
eiioeaio?aoeee iiei/iie nuai?ioee e ieaiiai nonea — aeaoaauo e
aeinooiiuo eiiiiiaioia ieoaoaeueiie n?aaeu. Au/eneaiiua ooieoeee
iicaieytho i?iaiice?iaaoue iiaaaeaiea i?iaeooeaioa a ethaie oi/ea
i?ino?ainoaa, ia?aie/aiiiai o?iaiyie oaeoi?ia. Aaaaeaiea aeaaa?aeeeia a
n?aaeo eoeueoeae?iaaiey H.laricinus noeioee?oao ia?aciaaiea
iieieinaa?ouaathuaai oa?iaioa. Aeiaaaeaiea aoeneia ia ?aiieo noaaeeyo
eoeueoeae?iaaiey iiaeaaeyao neioac iieieinaa?ouaathuaai oa?iaioa e
aeiiannu. Aeaiu ?aeiiaiaeaoeee aeey i?iiuoeaiiiai eoeueoeae?iaaiey
ooaiiia H. laricinus. Ii ?acoeueoaoai ?aaiou iieo/ai iaoaio ia
ecia?aoaiea e aeaa i?ei?eoaoiua ni?aaee ia ecia?aoaiea.

Eeth/aaua neiaa: nu/oaeiua oa?iaiou, eiiiiiaiou n?aaeu, iioeiecaoeey,
iiaa?oiinoue ioeeeea, iieieinaa?ouaathuay aeoeaiinoue

Nikitina O. A. Regulation of the activity of milk coagulating
exoproteases of Hirschioporus laricinus ( Karst.) Ryv. cultures. —
Manuscript.

The thesis of Candidate of sciences ( Biology) on speciality 03.00.04.
— biochemistry. — Kyiv state University named after Taras Shevchenko,
Kyiv, 1999.

The thesis is devoted to the problems of different physiological
factors — temperature and means of cultivation, the acidity of the
medium, carbon and nitrogen nutrition and also some biological additions
— influencing milk coagulating activity and the accumulation of biomass
by M-81, A-031, A-032 and P-323 Hirschioporus laricinus — the producents
of milk coagulating proteases. There have been investigated more then
120 nutrition mediums in the present paper. The 5 times increasing of
milk coagulating ferment(s output is achieved by means of sugars(
concentrations( selection.

The character of dependence of milk coagulating activity and grows on
milk serum(s and beer wort — cheap and easy-of-access components of
nutrition medium is investigated in detail; on the basis of this the
optimal cultivation medium was selected. Some recommendation concerning
industrial cultivation of Hirschioporus laricinus cultures are given. As
a result of the work carried out the invention patent and 2 priority
certificates of invention are obtained.

Key words: rennet ferments, medium components, optimization, response
surface, milk coagulating activity

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NH4NO3 (NH2)2CO (NH4)2SO4 CH3COONH4 NaNO3 KNO3
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