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?INOEOOO AIAeIE?EIIEIA?? OA IAI?IO ?A*IAEI

?I. A.I.EII?NA?AIEA AII OE?A?IE

E?EAE* Iaoae?y A?eoi??aia

OAeE
616.441-002-073.66

O?iii?eoie ooieoe?? ueoiaeaeii? caeice, aene?aoi?ii? ooieoe?? ie?ie
o oai?eo ia aeeoociee oiene/iee cia oa a?iioe?aic

14.01.14 – aiaeie?eiieia?y

A a o i ? a o a ? a o

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa iaaee/ieo iaoe

Ee?a – 1998

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia a Aoeiaeinuee?e aea?aeaai?e iaaee/i?e aeaaeai?? IIC
Oe?a?ie ia eaoaae?? ee?i?/ii? ?ioiieia??, aea?aieia?? oa aiaeie?eiieia??

Iaoeia? ea??aieee(

Aeieoi? iaaee/ieo iaoe, i?ioani? I?oae Aaneeue Iaaeiae/,

Aoeiaeinueea aea?aeaaia iaaee/ia aeaaeai?y, caa?aeoaa/ eaoaae?e
iaaee/ii? a?ieia?? oa aaiaoeee

Aeieoi? iaaee/ieo iaoe, i?ioani? Ie?eiee Aaea??e Aiaoie?eiae/,

?inoeooo aiaeie?eiieia?? oa iai?io ?a/iaei ?i. A.I.Eii?na?aiea AII
Oe?a?ie, caa?aeoaa/ a?aeae?eo caaaeueii? aiaeie?eiii? iaoieia??

Io?oe?ei? iiiiaioe:

Aeieoi? iaaee/ieo iaoe, i?ioani?, /e.-ei?aniiiaeaio AII Oe?a?ie

Aiooaei Ian?e Aieiaeeie?iae/, ?inoeooo aiaeie?eiieia?? oa iai?io ?a/iaei
?i. A.I.Eii?na?aiea AII Oe?a?ie, caa?aeoaa/ eaai?aoi??? ooieoe?iiaeueii?
ae?aaiinoeee

Eaiaeeaeao iaaee/ieo iaoe Naa?ai Ieaia Aaea???aia, Oe?a?inueeee
iaoeiai-i?aeoe/iee oeaio? aiaeie?eiii? o??o?a?? ? o?ainieaioaoe??
i?aai?a oa oeaiei, caa?aeoaa/ a?aeae?eo ee?i?/ii? aiaeie?eiieia??

I?ia?aeia onoaiiaa: Euea?anueeee aea?aeaaiee iaaee/iee oi?aa?neoao,
eaoaae?a aiaeie?eiieia??.

Caoeno a?aeaoaeaoueny (17( eenoiiaaea 1998 ?. i 1300 aiae. ia
can?aeaii? niaoe?ae?ciaaii? A/aii? ?aaee Ae.26.558.01 c aiaeie?eiieia??
a ?inoeooo? aiaeie?eiieia?? oa iai?io ?a/iaei ?i. A.I. Eii?na?aiea AII
Oe?a?ie (254114, Ee?a-114, aoe. Aeoai?iaenueea 69)

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? ?inoeoooo
aiaeie?eiieia?? oa iai?io ?a/iaei ?i. A.I. Eii?na?aiea AII Oe?a?ie
(254114, Ee?a-114, aoe. Aeoai?iaenueea 69)

Aaoi?aoa?ao ?ic?neaiee ( 13 ( aeiaoiy 1998 ?.

A/aiee nae?aoa? niaoe?ae?ciaaii? a/aii?

?aaee, aeieoi? a?ieia?/ieo iaoe
E.I.Eaeeinueea

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue ?iaioe: Oee?eaaeiaii? ?eoie aeeiiothoue aaaeeeao
ooieoeith neio?iiicaoei? oiciieiai/ieo i?ioeania. ?o ia?aiao?e iiaeooue
aooe naiinoieiith oa?aeoa?enoeeith oiciieiai/iiai noaio i?aaiicio. A
inoaii? ?iee, o ca’yceo c aeieia?/ii-iani?eyoeeaeie oiiaaie aeiae?eey,
c??n ??aaiue aiaeie?eiieo caoai?thaaiue, iniaeeai iaoieia?? ueoiaeaeii?
caeice (I.A.Aiooaei oa ni?aao., 1994, A.A.Ie?eiee, 1997). Oiio
i?iaaaeaiiy o?iiia?ieia?/ieo aeine?aeaeaiue a aiaeie?eiieia?? iaaoaa?
iniaeeai? aeooaeueiino? (O.E.Eiia?ia, 1989). Iaeiie/aii aaaaoi
na?ae/aiue uiaei aea/aiiy a?i?eoi?a ai?iiiaeueii? aeoeaiino? ?yaeo caeic
aiaeie?eiii? nenoaie, a naia a?iio?ca (A.A.Ie?eiee oa ni?aaao., 1997,
H.de Boer et al, 1989), iaaeie?iee?a ( M.Vance, 1989), noaoaaeo caeic
(S.Dias et al., 1989), oeoeiiiae?aiiai o?ea (A.I.I?oae, 1985,
D.Cardinali, 1989) oa ?io?. I?ioa, o?iii?eoie ueoiaeaeii? caeice,
o?iiiae?aaiinoeea oa o?iiioa?iaeieia?y ?? caoai?thaaiue aea/ai?
iaaeinoaoiuei, a ?nioth/? aeai? a?ae??ciythoueny aaeeeith
aa??aaaeuei?noth.

Caaaeueiia?aeiii, ui ie?eai aeanoeaa iaeaieueo /ioea ?eoii/ia
aeiyeueiinoue a i?aai?ci?, oiio aecia/aiiy ?a/iaei, ui aene?aoothoueny
ieie, aeicaiey? io?eiaoe oyao i?i ?eoii/io i?aaiicaoeith ?icieo nenoai
(O.E.Eiia?ia, 1989). Aieueoinoue aaoi?ia aeieoee aeniiaeo, ui ?iaioa
ie?ie a?aeaeca?eaeth? ooieoe?th ?aaoeyoi?ieo nenoai (A.Ae.Neaiooeei,
A.A.Iaoeineee, 1982, I.E.Aneaiyi e niaao., 1989). Aeaneio?iiic
o?c?ieia?/ieo nenoai i?aaiicio aeyaey?oueny ia ?aiiio noaaeiyo
caoai?thaaiiy i cia/ii ia?aaeo? ii?oieiai/iei ciiiai. Oe?ai?aeii ai?iiie
ni?e/eiythoue i?yio aeith ia ie?ee, ia ui aeaco? iayaiinoue a ieo
niaoeeoi/ieo ?aoeaioi?ia aei oe?ieneio oa o?eeiaeoe?iiiio
(I.H.Oppenheimer, 1983, L.D.Kohn et al., 1985). I?ioa, ie ia cono??/aee
aeaieo i?i no?oeoo?io i?aai?caoe?th o?iii?eoi?a aene?aoi?ii? ooieoe??
ie?ie o oai?eo c ii?ooaiiyi ooieoe?? ueoiaeaeii? caeice.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie: Aeena?oaoe?y ? o?aaiaioii
ieaiiaiai iaoeiaiai aeine?aeaeaiiy Aoeiaeinueei? aea?aeaaii? iaaee/ii?
aeaaeai?? “Aeine?aeeoe aieea e?ia?/ii? nenoaie iiceo oa aae?aia?a?/ieo
no?oeoo? ia ii?oiooieoe?iiaeueiee noai aiaeie?eiieo i?aai?a c iaoith
anoaiiaeaiiy aca?iica’yce?a i?ae ieie a caeaaeiino? a?ae a?eo oa
a?ieia?/ieo ?eoi?a” (iiia? aea?aeaaii? ?a?no?aoe?? 01.9.10010096 a?ae
19.04.1991?.)

Iaoa ?iaioe: Aeyaeoe iniaeeaino? o?iii?eoi?a ooieoe?? ueoiaeaeii?
caeice, o?iii?eoi?a aene?aoi?ii? ooieoe?? ie?ie o oai?eo ia aeeoociee
oiene/iee cia ? a?iioe?aic oa ?o aca?iica’ycie.

Caaaeaiiy aeine?aeaeaiiy:

1. Anoaiiaeoe iniaeeaino? oee?eaae?aiieo ?eoi?a ooieoe?? ueoiaeaeii?
caeice i?e aeeoociiio oiene/iiio cia? oa a?iioe?aic?.

2. Aea/eoe noai aene?aoi?ii? ooieoe?? ie?ie o oai?eo ia aeeoociee
oiene/iee cia ? a?iioe?aic.

3. Aeine?aeeoe o?iii?eoie aene?aoi?ii? ooieoe?? ie?ie o iaoe??io?a c
ii?ooaiiyi ooieoe?? ueoiaeaeii? caeice.

4. Aeyaeoe iiaeeeaee aieea o?iii?eoi?a ooieoe?? ueoiaeaeii? caeice ia
o?iii?eoie aene?aoi?ii? ooieoe?? ie?ie a ii?i? ? i?e iaoieia??
ueoiaeaeii? caeice.

5. ?ic?iaeoe i?eioeeie o?iiiae?aaiinoeee aeine?aeaeaiiy ai?noo
oe?ai?aeieo ai?iii?a o oai?eo c ii?ooaiiyi ooieoe?? ueoiaeaeii? caeice.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a: Aeyaeaiee aeaneio?iiic
ooieoe?? ueoiaeaeii? caeice i?e ?? iaoieia??, ia i?aenoaa? /iai aia?oa
?ic?iaeai? ?aeiiaiaeaoe?? uiaei o?iiiae?aaiinoeee aeeoociiai oiene/iiai
ciao oa a?iioe?aico.

Aia?oa aeine?aeaeaii o?iii?eoie aene?aoi?ii? ooieoe?? ie?ie o oai?eo ia
aeeoociee oiene/iee cia ? a?iioe?aic. Anoaiiaeaii, ui i?e ii?ooaii?
ooieoe?? ueoiaeaeii? caeice oee?eaae?aiiee ?eoi aene?aoi?ii? ooieoe??
ie?ie ci?ith? naith no?oeoo?o. Aeiaaaeaii aieea o?iii?eoi?a ooieoe??
ueoiaeaeii? caeice ia o?iii?eoie aene?aoi?ii? ooieoe?? ie?ie.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a: Ia i?aenoaa? aeine?aeaeaiiy
oee?eaae?aiieo ?eoi?a ooieoe?? ueoiaeaeii? caeice, o?iii?eoi?a
aene?aoi?ii? ooieoe?? ie?ie o oai?eo ia aeeoociee oiene/iee cia ?
a?iioe?aic iaoeiai iaa?oioiaai?, ?ic?iaeai? ? ai?iaaaeaeai? a i?aeoeeo
i?eioeeie o?iiiae?aaiinoeee ii?ooaiue ooieoe?? ueoiaeaeii? caeice.

Iniiai? iaoeia? iieiaeaiiy, ui aeianai? ia caoeno:

O?iii?eoie ooieoe?? ueoiaeaeii? caeice aieeaathoue ia oee?eaae?aii?
?eoie aene?aoi?ii? ooieoe?? ie?ie.

O iaoe??io?a c aeeoociei oiene/iei ciaii a?aeaoaa?oueny cnoa ae?ioac
ai?noo oe?ai?aeieo ai?iii?a a ne?iaaooe? e?ia? a a?e aeaiieo ? aa/??i?o
aiaeei, aeoiaey/e c /iai caa?? e?ia? aeey i?iaaaeaiiy oeeo
aeine?aeaeaiue a?eueo aeioe?eueii i?iaiaeeoe a ae?oa?e iieiaei? aeiy.

O oai?eo ia a?iioe?aic ae?ioace ai?noo oe?ai?aeieo ai?iii?a a
ne?iaaooe? e?ia? i?eiaaeathoue ia ?aieiaee, aaoeoace – ia aeaiiee
ia??iae aeiae, ia i?aenoaa? /iai ?o aeine?aeaeaiiy e?aua i?iaiaeeoe
aaeaiue.

Oee?eaae?aii? ?eoie aene?aoi?ii? ooieoe?? ie?ie o oai?eo ia aeeoociee
oiene/iee cia ? a?iioe?aic ci?iththoue naith no?oeoo?o ? iiaeooue
aeei?enoiaoaaoenue ye naiino?ei? ae?aaiinoe/i? oanoe.

Iniaenoee aianie iiooeoaa/a: Aaoi?ii naiino?eii i?iaaaeaii
aeine?aeaeaiiy oee?eaae?aiieo ?eoi?a ooieoe?? ueoiaeaeii? caeice oa
aene?aoi?ii? ooieoe?? ie?ie, noaio aiaeii-nieueiai? ??aiiaaae o oai?eo
ia aeeoociee oiene/iee cia oa a?iioe?aic, aeeiiaia noaoenoe/ia ia?iaea ?
iaoeiaa ?ioa?i?aoaoe?y io?eiaieo aeaieo.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??: Iniiai? iieiaeaiiy
aeiiia?aeaeenue ia iaoeiai-i?aeoe/i?e eiioa?aioe?? “Aeooaeuei? i?iaeaie
aeeoy/i? oa i?aee?oeiai? aiaeie?eiieia??” (Oa?e?a, 1995?.), ia
i?aeia?iaeieo neiiic?oiao “O??o?a?/i? i?iaeaie ? aeieia?y” (*a?i?aoe?,
1995?., 1996?.), “Aeooaeuei? ieoaiiy iaaee/ii? aeiiiiiae ianaeaiith”
(*a?i?aoe? – Aiaaeueoieuei, 1996?.).

Ioae?eaoe?? oa ?io? oi?ie ai?iaaaeaeaiiy:

Ca oaiith ?iaioe iioae?eiaaii 16 ?ia?o. Na?aae ieo 4 noaoo? o
oeaio?aeueieo oaoiaeo aeaeaiiyo, 4 ?io? iaoeia? noaoo?, 8 oac.

?acoeueoaoe aeine?aeaeaiue ai?iaaaeaeai? o i?aeoeeo ?iaioe a?aeae?eaiue
*a?i?aaoeueeiai iaeaniiai aiaeie?eiieia?/iiai aeeniaina?o, iniiai?
iieiaeaiiy aeei?enoiaothoueny i?e i?iaaaeaii? i?aeoe/ieo caiyoue c
aiaeie?eiieia?? a Aoeiaeinuee?e aea?aeaai?e iaaee/i?e aeaaeai??.

Ia’?i ? no?oeoo?a aeena?oaoe??: Aeena?oaoe?y aeeeaaeaia ia 200
noi??ieao, i?noeoue anooi, ?icae?ee “Iaeyae e?oa?aoo?e”, “Iaoiaee
aeine?aeaeaiiy”, “Ee?i?/ia oa?aeoa?enoeea ianoaaeaieo oai?eo”, “Io?eiai?
?acoeueoaoe”, “Iaaiai?aiiy io?eiaieo ?acoeueoao?a”, aeniiaee oa
i?aeoe/i? ?aeiiaiaeaoe??, iieaae/ee e?oa?aoo?e. ?iaioa ?ethno?iaaia 40
oaaeeoeyie oa 22 ?enoieaie. Nienie e?oa?aoo?e i?noeoue 289 aeaea?ae.

CI?NO ?IAIOE

Caaaeueia oa?aeoa?enoeea oai?eo ? iaoiaee ?o ianoaaeaiiy

A iniiao ?iaioe iieeaaeaiee iaoa??ae eiiieaeniiai aeine?aeaeaiiy
oee?eaae?aiieo ?eoi?a ooieoe?? ueoiaeaeii? caeice, aene?aoi?ii? ooieoe??
ie?ie o 78 oai?eo ia aeeoociee oiene/iee cia (AeOC), 25 – ia a?iioe?aic,
ye? ciaoiaeeeenue ia e?eoaaii? a *a?i?aaoeueeiio iaeaniiio
aiaeie?eiieia?/iiio aeeniaina?? i?ioyaii 1995 – 1996 ??., oa 20
i?aeoe/ii caei?iaeo in?a, ui neeaee eiio?ieueio a?oio. A a?oio
ianoaaeaieo oa?eoee iaoe??ioe c oe?aioieneeicii ? ia?aeiiei a?iioe?aicii
a noaae?? aeaeiiiainaoe?? aac noiooi?o caoai?thaaiue oa ae?aaeaii?
iaaeinoaoiino? e?iaiia?ao. Aaaee?noue oe?aioieneeico oa a?iioe?aico
anoaiiaethaaeenue ca nooiaiyie i?iyao ee?i?/ieo neiioii?a.

Na?aae oai?eo ia AeOC 26 aoee c oe?aioieneeicii eaaeiai nooiaiy (Oe),
32 – c oe?aioieneeicii na?aaeiueiai nooiaiy aaaeeino? (On), 20 – c
aaaeeei oe?aioieneeicii (Oa); 8 oai?eo ia a?iioe?aic -c eaaeei nooiaiai
aaaeeino? caoai?thaaiiy (Ae), 6 – c na?aaei?i (An), 11 – c aaaeeei
a?iioe?aicii (Aa). A?eueo?noue iaoe??io?a neeaaeaee ae?iee (65 ae?iie oa
13 /ieia?e?a c AeOC oa 20 ae?iie ? 5 /ieia?e?a c a?iioe?aicii).

Aeine?aeaeaiiy ooieoe?iiaeueiiai noaio ueoiaeaeii? caeice i?iaiaeeee ca
aeiiiiiaith aecia/aiiy ai?noo a ne?iaaooe? e?ia? oe?ieneio (O4),
o?eeiaeoe?ii?io (O3), oe?aio?iiiiai ai?iiio (OOA), oe?aiaeiaoe?io (OA).
??aaiue oe?ieneio aecia/aee iaoiaeii ?aae?i?ioiiiai aiae?co ca
aeiiiiiaith iaai?o ??i-O4-?I?, o?eeiaeoe?ii?io – ??i -O3-?I?,
oe?aiaeiaoe?io – ??i – OA 125J ?c aeei?enoaiiyi oe?aiaeiaoe?io, i?/aiiai
eiaeii – 125 (A?ei?onue), oe?aio?iiiiai ai?iiio-?EA-OOA (*ao?y) ia
?aae?i?ioiiiio aiae?caoi?? “Aaiia-12”.

Eiioeaio?aoe?th ?ii?a iao??th ? eae?th a ieaci? e?ia?, a?eo?ioeeoao oa
na/? aecia/aee iaoiaeii oioiiao??? iieoi’y ia “OIE -1”, e?aaoei?io – ca
?aaeoe??th c i?e?eiiaith eeneioith (N.E.?yaia e niaao., 1979) c
?a?no?aoe??th iieaciee?a aenoeioe?? ia oioieaei?eiao?? “EOE – 2” ?
niaeo?ioioiiao?? “NO – 46” (A.E.Ia?cii, 1974).

Oaeaee?noue eeoai/eiai? o?eueo?aoe?? ?ic?aoiaoaaee ca ee??ainii
aiaeiaaiiiai e?aaoei?io. Ae?aoiaoaaee eaiaeueoeaaee o?ainii?o ?ii?a.
Aeey ioe?iee o?ainii?oo aeaeo?ie?o?a a ie?eao aeei?enoiaoaaee iieacieee
ee??aino, aene?aoi?io o?aeoe?th ?ii?a iao??th. ?ic?aoiaoaaee aene?aoe?th
e?aaoei?io, ?ii?a iao??th ? eae?th ca 3, 12, 24 aiaeeie. Ioe?ieo
eaiaeueoeaaiai o?ainii?oo aiaee i?iaiaeeee ca iieacieeaie ?aaani?aoe??
aiaee oa ?? aene?aoi?ii? o?aeoe??.

Oee?eaae?aii? ?eoie ooieoe?? ueoiaeaeii? caeice aea/aee ca aeiiiiiaith
aecia/aiiy ai?noo O3, O4, OOA a ne?iaaooe? e?ia? o?e/? ai?iaeiaae aeiae
(08.00; 16.00; 24.00).Oee?eaae?aii? ?eoie aene?aoi?ii? ooieoe?? ie?ie
aeine?aeaeoaaee oeyoii aecia/aiiy ia’?io na/?, ai?noo ?ii?a iao??th ?
eae?th, e?aaoei?io a 3-aiaeeiieo ii?oe?yo na/? i?ioyaii aeiae.

A aeii ianoaaeaiue oai?? aeio?eioaaeenue noaiaea?oeciaaiiai ?aaeeio
nii-aaaeuei??noue, 4-?aciaiai ciioaiiai ?aoeiiio aac iaiaaeaiiy o
i?eeiii ?iaeeie oa c iiii?iei (aei 10a/aeiao) aaeeaaiiyi eooiiii? niei.

Io?eiai? aeai? ia?iaeai? ca aeiiiiiaith ein?ii?-aiae?co. Aea/aee oae?
ia?aiao?e ?eoio: iaci? (na?aaeiueiaeiaiaee ??aaiue aeine?aeaeoaaiiai
iieacieea), aiie?ooaeo (??cieoeth iaeneiaeueiiai a?aeoeeaiiy a?ae
iaci?o), ae?ioaco (iiiaio /ano, a yeee a?aei?/aeinue iaeneiaeueia
cia/aiiy iieacieea) ? aaoeoaco (iiiaio /ano, a yeee niinoa??aaeinue
i?i?iaeueia cia/aiiy iieacieea).

?acoeueoaoe aeine?aeaeaiue ia?iaeai? ia AII IBM PC 486 DX 4-100 ca
aeiiiiiaith i?ia?aie Microsoft Excel.

O?iii?eoie ooieoe?? ueoiaeaeii? caeice o oai?eo ia aeeoociee oiene/iee
cia oa a?iioe?aic

A oiae? aeine?aeaeaiiy aeyaeaiee aeaneio?iiic ooieoe?? ueoiaeaeii?
caeice o iaoe??io?a c AeOC oa a?iioe?aicii (oaaeeoey 1).

O oai?eo c Oe ca?eueooaaeenue iaci?e oa aiie?ooaee O3 oa O4 (?0,05). A?aei?/aany cnoa ae?ioac O3 (ia
4aiae. 44oa.), O4 (ia 8aiae. 40oa.) a a?e aeaiieo oa aa/??i?o aiaeei oa
aaoeoac O3 (ia 8aiae. 34oa.), O4 (ia 8aiae. 48oa.) a a?e ?aieiaeo
aiaeei. Ae?ioaca oa aaoeoaca OOA nooo?ai ia a?ae??ciyeanue a?ae
eiio?ieth.

I?e On iaci?e O3 oa O4 c?inoaee a??ia?aeii ?noioi?oa, i?ae i?e Oe
(?0,05 a?aeiia?aeii).
A?aei?/aany cnoa ae?ioac O3 (ia 5aiae. 11oa.) oa O4 (ia 10aiae. 52oa.) a
a?e aeaiieo oa aa/??i?o aiaeei, i?e/iio aeey O4 a?eueo ae?aaeaiee (ia
2aiae. 12oa.), i?ae i?e Oe, a oaeiae ci?uaiiy aaoeoac iieaciee?a (O3 –
ia 9aiae. 23oa., O4 – ia 9aiae. 28oa.) a a?e ?aieiaeo aiaeei. Ae?ioaca
OOA nooo?ai ia ci?ieeany.

O iaoe??io?a c Oa a?aei?/aeinue nooo?aa ca?eueoaiiy iaci??a O3 (P0,05) ia a?ae??ciyeeny a?ae iieaciee?a
i?e ?ioeo nooiaiyo aaaeeino? caoai?thaaiiy. A?aei?/aany cnoa ae?ioac O3
(ia 4aiae. 01oa.) oa O4 (ia 12aiae. 18oa.) uiaei eiio?ieth a a?e aeaiieo
oa aa/??i?o aiaeei, yeee aeey O4 aoa a?eueo ae?aaeaiei, i?ae i?e ?ioeo
oi?iao caoai?thaaiiy (ia 3aiae. 38oa. ii??aiyii c Oe oa ia 1aiae. 19oa.
ii??aiyii c On). A?aeaoaaany cnoa aaoeoac (O3 – ia 8aiae. 17oa. oa O4 –
ia 10aiae. 02oa.) a a?e ?aieiaeo aiaeei uiaei eiio?ieth. Ae?ioaca OOA ia
a?ae??ciyeanue a?ae eiio?ieth.

C?inoaiiy iaci??a O3 oa O4 oe?eeii caeiiii??ii, ine?eueee a?aeaoaa?oueny
ca?eueoaiiy ??aiy oe?ai?aeieo ai?iii?a. Aeieea? ieoaiiy, /iio aiie?ooaea
O4 c?inoa? i?yii, a O3 – iaa?iaii i?iii?oe?eii nooiaith aaaeeino?
caoai?thaaiiy. Oaea iaocaiaeaeai?noue i?ae ?eoiaie iiaea caeaaeaoe a?ae
eiiaa?n?? oeeo ai?iii?a, ui iaoiiaeth?oueny aeoeai?noth 5/-aeaeiaeeiace.
Oea i?eiouaiiy i?aeoaa?aeaeo?oueny ?iaioaie M.Muracami et al. (1988,
1989), a yeeo aeyaeai? o?iii?eoie aeoeaiino? eiaeoe?ii?i –
5/-aeaeiaeeiace c ae?ioacith ii?aii/?. Ciaioaiiy iaci?o OOA
iiynith?oueny cieaeaiiyi ??aiy inoaiiueiai a ieaci? e?ia? i?e ia?aeiiiio
oe?aioieneeic? ca i?eioeeiii cai?ioiiai ca’yceo.

I?e aiae?c? ?eoio ooieoe?iiaeueii? aeoeaiino? a?iio?ca?ii-oe?ai?aeii?
nenoaie i?eaa?oa? oaaao ?ica?aei?noue ca /anii oaeeue iaeneiaeueii?
nae?aoe?? OOA oa oe?ai?aeieo ai?iii?a. A?eueoa oiai, ae?ioaca OOA ia
ia?aaeo? ae?ioacai nae?aoe?? O3 ? O4. I/aaeaeii, oee?eaae?aiiee ?eoi
ai?noo a ieaci? e?ia? O3 ? O4 caeaaeeoue ia o?eueee a?ae OOA-aeoeaiino?.

Cnoa ae?ioac iieaciee?a, ia iaoo aeoieo, iia’ycaiee c ii?ooaiiyi
oe?ai?aeii-ai?o?ca?ieo a?aeiinei i?e i?aeaeuaii? ooieoe?? ueoiaeaeii?
caeice. A?aeiii, ui oe?aioieneeic aeeeeea? ??cea ca?eueoaiiy
eiioeaio?aoe?? iaeaoii?io (ca aeaieie E.A.Aiiaea?aiei (1991) – a 8,5
?ac?a). A naith /a?ao, aeoiaey/e c i?eioeeio cai?ioiiai ca’yceo, ci?ie a
oeoeiiiae?aiiio o?e?, ye? a?aeaoeeny i?e oe?aioieneeic?, aeeeeeaee
aeieeiaiiy aeaneio?iiico ooieoe?? ueoiaeaeii? caeice.

A?aoiaoth/e aeuacacia/ai? ci?ie, iiaeia ae?eoe aeniiaeo, ui caa?? e?ia?
aeey aeine?aeaeaiiy ai?noo oe?ai?aeieo ai?iii?a o iaoe??io?a c AeOC
a?eueo ?ioi?iaoeaii i?iaiaeeoe a ae?oa?e iieiaei? aeiy. Ee?i?/ia
aeine?aeaeaiiy a?aioe? aaaaea?ii iaei?ioi?iaoeaiei, ine?eueee o oai?eo
ia oe?aioieneeic ia ?aieia? aiaeeie i?eiaaeathoue aaoeoace iieaciee?a.

sse aeaeii c oaaeeoe? 1, o iaoe??io?a c Ae niinoa??aaeiny cieaeaiiy ii
a?aeiioaiith aei eiio?ieth iaci?o oa aiie?ooaee O3 (?0,05).
A?aeaoaaany cnoa ae?ioace O3 (ia 6aiae. 17oa.) ia ?aieia? aiaeeie,
aaoeoace – ia aeaii? aiaeeie (ia 6aiae. 36oa.), ae?ioace OOA (ia 3aiae.
15oa.) a a?e aa/??i?o aiaeei, iii?oi?oee, i?ae i?e Ae, aaoeoace – a a?e
?aieiaeo aiaeei (ia 4aiae. 00oa.).

O iaoe??io?a c Aa niinoa??aaeinue a?eueoa ciaioaiiy iaci??a O3 (P0,05). Aiie?ooaea OOA cieaeoaaeanue i?iii?oe?eii ?c aaaee?noth
caoai?thaaiiy (P0,05). ??aaiue
?ii?a iao??th a a?eo?ioeeoao aeuee, i?ae o caei?iaeo in?a (a?aeiia?aeii
23,9+0,46iIieue/e ? 19,4+0,59 iIieue/e, P0,05). Eiioeaio?aoe?y eae?th a a?eo?ioeeoao cieaeoaaeany
(85,2+0,76iIieue/e – oai??; 96,1+1,07 iIieue/e – caei?ia?, P0,05,
17,8+1,27iI/oa., P>0,05) oa cieaeoaaeenue i?e Oa (119,6+1,75iI/24 aiae.,
P0,05 ?
37,2+1,55iI/24aiae., P>0,05). I?e Aa c?inoa? i?/iee ? ciaioo?oueny
aeaiiee eae?eo?ac.

Ia iaoo aeoieo, iaeiei c iaoai?ci?a aiaeii-nieueiaiai aeenaaeaino i?e
a?iioe?aic? ? iaaeeeoeiaa iaeiie/aiiy o aioo??oi?o i?aaiao eeneeo
ae?eicai?iiae?eai?a, ui iineeththoue a?ae?io?euei?noue niieo/ii? oeaieie
oa aeeeeeathoue cao?eieo iao??th ca iniioe/iei a?aae??ioii.

Ia cao?eieo a oeaieiao ?ii?a iao??th ? aiaee oaeiae iiaea aieeaaoe
iaaeeeoie aacii?aneio. Ia iaoo aeoieo, naia c oeei iia’ycaii a??ia?aeia
ciaioaiiy ae?o?aco aaeaiue ? aeieeiaiiy iiaiai i?eo aene?aoe?? aii/?,
yea a?aei?/aii o iaoe??io?a c a?iioe?aicii.

Ci?ie aiaeii-nieueiaiai iai?io i?e a?iioe?aic? oaeiae iaoiiaeai?
cieaeaiiyi ??aiy IIOA (N.L.Wong et al., 1992). Ioaea, cieaeaiiy
iao??eo?aco iiaea aooe iia’ycaii oaeiae ?c oeei oaeoi?ii.

O?iii?eoie aene?aoi?ii? ooieoe?? ie?ie o oai?eo ia aeeoociee oiene/iee
cia oa a?iioe?aic

Io?eiai? iaie aeai? noiniaii ?eoi?/ii? ae?yeueiino? ie?ie i?e AeOC oa
a?iioe?aic? na?ae/aoue i?i aeaneio?iiic ?o aene?aoi?ii? ooieoe??
(oaaeeoey 2).

O oai?eo ia Oe caieueooaaeenue iaci?e oa c?inoaee aiie?ooaee
na/iaeae?eaiiy (P0,05),
a oaeiae c?inoaiiy iaci?o i aiieiooaee aene?aoei? ?ii?a eae?th (P0,05) oa e?aaoei?io?aco (P>0,05), ciaioaiiy aiie?ooae an?o
iieaciee?a, iniaeeai ae?aaeaia aeey ae?o?aco (POaaeeoey 1 O?iii?eoie ai?noo O3, O4 oa OOA a ne?iaaooe? e?ia? o oai?eo ia aeeoociee oiene/iee cia oa a?iioe?aic Iieacieee Eiio?ieue (n=18) Oe (n=13) On (n=14) Oa (n=14) Ae (n=6) An (n=6) Aa (n=7) O3 Iaci? 1,12(0,018 2,61(0,029* 2,84(0,033* 3,34(0,072* 1,06(0,022* 0,57(0,082* 0,41(0,033* (iI\e) Aiie?ooaea 0,29(0,004 2,12(0,032* 1,52(0,018* 0,52(0,014* 0,21(0,006* 0,07(0,001* 0,05(0,004* Ae?ioaca 12051/ 17035/ 18002/ 16052/ 5018/ 6034/ 6010/ Aaoeoaca 23048/ 8022/ 9011/ 8005/ 15055/ 17012/ 16028/ O4 Iaci? 108,60(0,77 152,54(1,621* 189,63(2,890* 209,17(3,931* 55,74(0,972* 27,96(3,494* 16,05(3,526* (iI\e) Aiie?ooaea 21,85(0,180 31,22(0,598* 18,29(0,262* 26,91(2,839* 13,46(0,016* 7,32(0,927* 1,36(0,482 Ae?ioaca 5058/ 14038/ 16050/ 18016/ 7019/ 6028/ 6014/ Aaoeoaca 16010/ 7022/ 6042/ 6008/ 16025/ 16016/ 16005/ OOA Iaci? 2,37(0,098 1,40(0,249* 0,78(0,048* 0,63(0,071* 6,48(0,454* 9,41(0,674* 12,98(0,723* (iIA\e) Aiie?ooaea 2,69(0,111 1,74(0,434 1,45(0,101* 1,23(0,134* 1,31(0,093* 2,82(0,201 1,78(0,098* Ae?ioaca 1024/ 1029/ 0056/ 0038/ 22042/ 22009/ 21004/ Aaoeoaca 12010/ 12030/ 12056/ 12049/ 11010/ 8010/ 6042/ * - a??ia?aei? ci?ie uiaei eiio?ith (? Oaaeeoey 2 O?iii?eoie ae?o?aco, aene?aoe?? Na+, E+ oa e?aaoei?io o oai?eo ia aeeoociee oiene/iee cia oa a?iioe?aic Iieacieee Eiio?ieue (n=20) Oe (n=26) On (n=32) Oa (n=20) Ae (n=8) An (n=6) Aa (n=11) Ae?o?ac Iaci? 135,1(2,32 186,2(4,89* 172,8(2,37* 107,2(3,25* 113,3(1,75* 97,6(1,75* 97,2(1,59* (ie\3 aiae) Aiie?ooaea 51,0(1,06 108,3(3,13* 64,3(1,07* 45,0(1,17* 22,63(0,411* 31,1(0,61* 17,9(0,34* Ae?ioaca 15005/ 14048/ 10005/ 9035/ 19038/ 20025/ 00034/ Aaoeoaca 4000/ 5030/ 2050/ 24000/ 8000/ 8043/ 12000/ Na+ Iaci? 18,42(0,282 23,42(0,352* 17,29(0,523 11,27(0,269* 15,25(0,420* 11,73(0,442* 10,52(0,284* (iI\3 aiae) Aiie?ooaea 11,74(0,188 16,63(0,319* 11,88(0,149 7,24(0,194* 3,02(0,097* 4,71(0,219* 1,72(0,052* Ae?ioaca 15029/ 12000/ 10042/ 6000/ 17014/ 3002/ 2044/ Aaoeoaca 5000/ 1030/ 21030/ 21000/ 7010/ 8030/ 12000/ E+ Iaci? 4,48(0,054 7,18(0,079* 7,73(0,068* 5,53(0,073* 4,74(0,151 4,61(0,218 5,04(0,172* (iI\3 aiae) Aiie?ooaea 3,75(0,048 6,63(0,071* 7,29(0,057* 3,09(0,056* 2,16(0,082* 1,32(0,068* 1,11(0,042* Ae?ioaca 14014/ 12000/ 10053/ 9027/ 13031/ 2020/ 5059/ Aaoeoaca 2030/ 24000/ 21030/ 21026/ 3005/ 18000/ 18008/ E?aaoei?i Iaci? 1,26(0,017 2,03(0,023* 1,82(0,212 1,08(0,096 1,10(0,074 0,71(0,041* 1,08(0,084 (ieI\3aiae) Aiie?ooaea 0,67(0,093 1,34(0,018* 1,24(0,013* 0,26(0,054* 0,46(0,016 0,57(0,037 0,44(0,021 Ae?ioaca 17000/ 15054/ 12000/ 11053/ 24000/ 1027/ 2040/ Aaoeoaca 5048/ 3050/ 24000/ 21025/ 9020/ 12005/ 12000/ * - a??ia?aei? ci?ie uiaei eiio?ieth (?

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