2

OE?A?INUeEA AEAAeAI?ss AA?A?IEO IAOE

AeA?AEAAIEE I?E?ONUeEEE AIOAI?*IEE NAAe

EI?NAEIAA NA?OEAIA IAAE?AIA

OAeE 633.812: 665.527. 64 (477.75)

AEIEIAI-A?IEIA?*I? INIAEEAINO? ? AO??IIE?EI?NOUe AEAe?A ?IAeO THYMUS L.
IA I?AAeAIIIIO AA?AC? E?EIO

03.00.05 — Aioai?ea

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa

a?ieia?/ieo iaoe

sseoa — 1998

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia a Aea?aeaaiiio I?e?onueeiio aioai?/iiio naaeo OAAI
(AeIAN) o a?aeae?e? iiaeo oaoi?/ieo ? e?ea?nueeeo eoeueoo? oa
aa?iiaoai?ieia?/i?e noaioe?? «I?e?onueeee naae».

Iaoeiaee ea??aiee — aeieoi? a?ieia?/ieo iaoe ?aaioyaia Aaea??e
Aeieo?iae/, AeIAN, caa. a?aeae?eii iiaeo oaoi?/ieo ? e?ea?nueeeo
eoeueoo?.

Io?oe?ei? iiiiaioe:

— Aeieoi? a?ieia?/ieo iaoe Ei?aeaiaanueeee Aeaaeeneaa A’y/aneaaiae/,
AeIAN,

caa. a?aeae?eii oei?e ? ?ineeiiino?;

— Eaiaeeaeao a?ieia?/ieo iaoe ?ia Aiae??e Aaneeueiae/, E?einueeee
aea?aeaaiee aa?a?iee oi?aa?neoao, aeioeaio eaoaae?e aioai?ee,
o?c?ieia?? ?ineei ? i?e?ia?ieia??.

I?ia?aeia onoaiiaa — ?inoeooo ao??iie?eieo ? e?ea?nueeeo ?ineei
OAAI,

i. N?ioa?iiieue.

Caoeno a?aeaoaeaoueny » 16 » a?oaeiy 1998 ?. i 14 aiae. ia
can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee Ae 53.369.01 c caoenoo
aeena?oaoe?e ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
i?e I?e?onueeiio aioai?/iiio naae? OAAI ca aae?anith: 334267,
Oe?a?ia, E?ei, i. sseoa, Aea?aeaaiee I?e?onueeee aioai?/iee naae.

C aeena?oaoe??th iiaeia iciaeiieoenue o iaoeia?e a?ae?ioaoe? AeIAN

Aaoi?aoa?ao ?ic?neaiee » 13 » eenoiiaaea 1998 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee, e.a.i._______________ N.TH.Naaeiao?nueeee

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. I?e ?ic?iaoe? e?ea?nueeeo cania?a ? i?aia?ao?a
c ?ineeiii? ne?iaeie noai?aiiy ne?iaeiii? aace aeey iaaee/ii?
i?iieneiaino? iaaoaa? iniaeeaiai cia/aiiy. Aaaeeeaa i?noea a oe?e
i?iaeai? caeia? iiooe, anaa?/ia aea/aiiy ? aaaaeaiiy a eoeueoo?o iiaeo
e?ea?nueeeo ?ineei. Iaeiei c ia?niaeoeaieo aeey aeei?enoaiiy a
iaaeeoeei?, ? aeae /aa?aoeue (Thymus L.) c ?iaeeie yniioeiaeo
(Lam?aceae). A ca’yceo c oei, ui ne?iaeia /aa?aoeth aeey oa?iaeiia?
caaioiaey?oueny, ye i?aaeei, c aeeei?ineeo ?ineei ? /anoi ia
a?aeiia?aea? iniiaiei noaiaea?oai, aaaaeaiiy a eoeueoo?o /aa?aoeth ia?
aaeeea i?aeoe/ia cia/aiiy. Eoeueoo?a /aa?aoeth aeicaieeoue iioe?eoe
iiaeeeaino? eiai aeei?enoaiiy.

Aiae?c e?oa?aoo?ieo aeaea?ae iieacaa, ui iiaiiai eiiieaeniiai
aeine?aeaeaiiy aeae?a /aa?aoe?a aein? ia i?iaiaeeeiny. Iaaecae/aeii
iaiaaeaia oaeiae e?euee?noue ie?aieo i?ea?iaeueieo ?ia?o c eiai a?ieia??
? iniaeeai a?io?i??. A?eueoa /anoeia iayaii? e?oa?aoo?e c a?oie ?ineei,
yeo ie ?icaeyaea?ii, iineoue caaaeueiee iaeyaeiaee oa?aeoa?. Ia
aea/aiith ? oaeiae i?iaeaia a?ieia?/ii? i?iaeoeoeaiino?. Oiio noai?aiiy
a?eueo o?iaeaeieo aeae?a ? a?a?eae?a /aa?aoeth c iaeneiaeueiith
e?euee?noth a?ieia?/ii aeoeaieo niieoe a ne?iaei? caeeoa?oueny
iaae??oaiith ? iaaeoi aeooaeueiith i?iaeaiith.

Ae??oaiith iinoaaeaiiai caaaeaiiy a cia/i?e i??? ni?ey?
aeei?enoaiiy ia?niaeoeaieo iaoiae?a noai?aiiy aeo?aeiiai iaoa??aeo,
aeineiiaeeo e?eoa???a eiai ioe?iee oa aoaeoeaieo cania?a a?aeai?o
aenieii?iaeoeoeaieo aaiioei?a, iniiaaieo ia aeeaieiio ? anaa?/iiio
aea/aii? i?iaeoeoe?eiiai i?ioeano ? oaeoi??a na?aaeiaeua, ui
aecia/athoue ??aaiue iaeiie/aiiy ao??ii? ie??.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie.
Aeine?aeaeaiiy aeeiiai? ca oaiaoe/iei ieaiii Aea?aeaaiiai I?e?onueeiai
aioai?/iiai naaeo c 1995 ii 1998 ??. c i?iaeaie 02 «Caeo/eoe, aea/eoe ?
aeae?eeoe ia?niaeoeai? aeey ai?iaaaeaeaiiy o ae?iaieoeoai aeaee
ao??iie?eieo, e?ea?nueeeo, i?yiia?iiaoe/ieo oa e?aneeueieo ?ineei,
aaeineiiaeeoe iaoiaee naeaeoe?? ? noai?eoe aenieii?iaeoeoeai? ni?oe»
(iiia? aea?aeaaii? ?a?no?aoe?? 0197 U004319) ? ieaiii ani??aionueei?
i?aeaioiaee.

Iaoa aeine?aeaeaiiy — iiooe ? aeae?eaiiy ia?niaeoeaieo aeae?a ?
oi?i oeyoii ii??aiyeueiiai aea/aiiy a?iii?oieia?/ieo, o?i?/ieo
iniaeeainoae oa ooee?oa?ieo iciae 12 aeae?a /aa?aoeth a caeaaeiino? a?ae
aeieia?/ieo oaeoi??a.

Caaaeaiiy aeine?aeaeaiiy:

— Aea/eoe iniaeeaino? c?inoaiiy ? ?icaeoeo ?io?iaeoeiaaieo
aeae?a /aa?aoeth ? aeaoe noeneo aioai?/io oa?aeoa?enoeeo oeeo aeae?a o
iiaeo oiiaao ae?iuoaaiiy (I?aaeaiiee aa?aa E?eio), ioe?ieoe aeioeie c
eiiieaeno ii?oia?ieia?/ieo iieaciee?a, aeae?eeoe ia?niaeoeai? ?
?aeiiaiaeoaaoe ?o aeey aaaaeaiiy a eoeueoo?o;

— Anoaiiaeoe eiiiiiaioiee neeaae ao??ii? ie?? 12 aeae?a /aa?aoeth,
aeeiai?eo eiai iaeiie/aiiy a iioiaaiac?, aecia/eoe e?euee?nio
caeaaei?noue iniiaieo eiiiiiaio?a ? ianiai? /anoee ao??ii? ie?? a?ae
aeieia?/ieo oiia a ia??iae oea?o?iiy;

— Aeyaeoe iniiai? no?oeoo?i? aeaiaioe i?iaeoeoeaiino? /aa?aoeth ?
neeanoe ?? iaoaiaoe/io iiaeaeue;

— Ocaaaeueieoe aaaaoi??/i? aa?iiaoai?ieia?/i? aeai? ca ia??iae
aaaaoaoe?? /aa?aoeth, ia ?o iniia? aecia/eoe e?euee?ni? iieacieee
aeiia /aa?aoeth aei oiia ciai?oiueiai na?aaeiaeua ? aeaoe ioe?ieo aieeao
inoaii?o ia i?iaeoeoeai?noue ?ineei.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Aea/aiee nooi?iue
aaeaioaoe?? ?ineei /aa?aoeth aei iiaeo oiia ? c?iaeaii noeneee
aioai?/iee iien ?io?iaeooeaio?a a eoeueoo?? ia I?aaeaiiiio aa?ac? E?eio.

Aeine?aeaeaiee iioiaaiac oeaiiiiioeyoe?e, aecia/aia
aioo??oiueiaeaeiaa i?ieea?noue aiaoiii-ii?oieia?/ieo,
ainiiaea?nueei-oe?iieo ? a?io?i?/ieo iciae.

Aia?oa aecia/aia e?euee?noue o?iiinii o Thymus camphoratus Hoffm.
et L?nk ? T. r?chard?? Pers. Subsp. n?t?dus (Guss), neioaciaaieo
iie?iei?ae?a ? i?aeaeaeiaeo a?a?eae?a.

Aecia/aiee o?i?/iee neeaae ao??ii? ie?? 12 aeae?a, ui aeeth/athoue
15 aeioei?a /aa?aoeth, i?e/iio aeey 8 aeae?a aia?oa; aea/aia aeeiai?ea
iaeiie/aiiy ao??ii? ie?? oa ?? iniiaieo eiiiiiaio?a a iioiaaiac?; ia
iniia? ii??aiyeueiiai aea/aiiy eiiiiiaioiiai neeaaeo ao??ii? ie??
/aa?aoeth eaioi?iiai ? aaeae?ioa aeyaeaia ?o iiae?ai?noue ca o?i?/iei
neeaaeii ? iieacaia ia?niaeoeai?noue aaaaeaiiy a eoeueoo?o /aa?aoeth
eaioi?iiai ye ao??iie?eii? ?ineeie.

Aia?oa ?ic?iaeaia iiaeaeue i?iaeoeoeaiino? /aa?aoeth, iniiaaia ia
?aaeoe?? aaiioeio /aa?aoeth ia aeieia?/i? oiiae, aecia/ai? neeaaeia?
iciaee i?iaeoeoeaiino? ? /anoee aieeao aaiioeio oa aeieia?/ieo oaeoi??a.

Aia?oa aeaia aeaoaeueia oa?aeoa?enoeea aeiia /aa?aoeth aei
iniiaieo oaeoi??a na?aaeiaeua ae?iuoaaiiy, aecia/ai? ? iaoaiaoe/ii
iaa?oioiaai? e?euee?ni? iieacieee iioeiaeueieo ? e?eoe/ieo oiia
oi?ioaaiiy o?iaeath oa a?ineioaco ao??ii? ie??. Anoaiiaeai?
caeiiii??iino? i?ieeaino? e?eueeino? ? yeino? ao??ii? ie??.

Ia i?aenoaa? eiiieaenieo aeine?aeaeaiue aeae?eai? /ioe?e
ia?niaeoeai? aeey aaaaeaiiy a eoeueoo?o aenieii?iaeoeoeai? aeaee
/aa?aoeth: T.camphoratus Hoffm. et Link. (/. eaioi?iee), T.vulgaris L.
(/. cae/aeiee), T.pulegioides L. (/. aeioeiee) ? T.striatus Vahl. (/.
oaeaaaiii?ecaieoaaoee).

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. Noai?aia iaoaiaoe/ia
iiaeaeue i?iaeoeoeaiino? /aa?aoeth, ui iieaco? caeaaei?noue iciaee
i?iaeoeoeaiino? a?ae aaiioeio ? oaeoi??a na?aaeiaeua, aeicaiey?
i?iaiicoaaoe i?iaeoeoeai?noue ?ineei a caeaaeiino? a?ae oaiia?aoo?e
iia?o?y ? noie iiaae?a, ui, a naith /a?ao, ni?ey? i?aeaeuaiith
aoaeoeaiino? naeaeoe?eiiai i?ioeano.

Iaa?oioiaaia ia?niaeoeai?noue aaaaeaiiy a eoeueoo?o /aa?aoeth
eaioi?iiai ye ao??iie?eii? ? e?ea?nueei? ?ineeie, aeneiaeaia i?iiiceoe?y
ae?iuoaaiiy eiai ia i?aaei? Oe?a?ie.

Aeae?eai? aenieii?iaeoeoeai? aeaee /aa?aoeth — /. cae/aeiee, /.
aeioeiee ? /. oaeaaaiii?ecaieoaaoee — ? iiaeie ia?niaeoeaieie
e?ea?nueeeie eoeueoo?aie ? iiaeooue aeei?enoiaoaaoeny ye aeaea?aea ?yaeo
oaiieueieo niieo/aiue aeey oa?iaoeaaoe/ii? i?iieneiaino?. Aeiiae
eoeueoo?e /aa?aoeth aei na?aaeiaeua c?inoaiiy neoaeaoue aeey ioe?iee
aa?iaeieia?/iiai oiio iieueiaeo aeine?ae?a, oiia ? ?acoeueoao?a
ae?iuoaaiiy ne?iaeie o ae?iaieoeoa?. ?ic?iaeai? caniae ia?aeiii?
aa?ioaoi?ee ae?iuoaaiiy /aa?aoeth.

Iniaenoee aianie a io?eiaiiy ?acoeueoao?a aeine?aeaeaiue iieyaa? a
naiino?ei?e ?ic?iaoe? ? aeeiiaii? iaoeiai-aeine?aeieoeueei? i?ia?aie,
ia?iaoe? ? aiae?c? aenia?eiaioaeueieo aeaieo (1995-1998 ??.); a
e?eoe/iiio aiae?c?, nenoaiaoecaoe?? oa ia?iaoe? ia?aeiieo aeaieo
oaiieia?/ieo niinoa?aaeaiue a?aeae?eo Iiaeo oaoi?/ieo ? e?ea?nueeeo
eoeueoo? Aea?aeaaiiai I?e?onueeiai aioai?/iiai naaeo a eiiieaen? c
aaaaoi??/ieie aeaieie aa?iiaoainoaioe?? «I?e?onueeee naae» ca ia??iae
1980-1994 ??., noai?aii? aace aeaieo, i?aeaioiaoe? noaoae,
?aeiiaiaeaoe?e ? ai?iaaaeaeaii? o ae?iaieoeoai.

?acii c iaoeiaei ea??aieeii aeeiiai?: aea?? ia’?eo?a ? iaoiaeee
aeine?aeaeaiiy, ?ic?iaea ? no?oeoo?a aeena?oaoe?eii? ?iaioe.

A?aea?? ? i?aea?? iaoa??ae?a o cai?ao e?ea?nueeeo o?aa aeey
ae?oae, ui iino?aaeaeaee aiane?aeie *i?iiaeeuenueei? eaoano?ioe;
aecia/aiiy aioei?e?iaii? aeoeaiino? ao??ii? ie?? /aa?aoeth i?iaiaeeee
?acii c Oe?a?inueeei IAe? aeeoy/i? eo?i?oieia?? ? o?c?ioa?ai??,
Ia?oio?aaiaaei ?aeiiiei oa?eoi??aeueiei iaaee/iei ia’?aeiaiiyi (I?OII)
(/anoea iniaenoi? o/ano? — 10 %).

Ai?iaaoe?y ?acoeueoao?a. Iaoa??aee aeine?aeaeaiue aoee
iia?aeiieai? ia can?aeaiiyo a?aeae?eo iiaeo oaoi?/ieo ? e?ea?nueeeo
eoeueoo?, a/aii? ?aaee AeIAN (1995-1998 ??.); ia iaoeiai-i?aeoe/i?e
eiioa?aioe?? iieiaeeo a/aieo «I?iaeaie aeaiae?ieia??, ea?oieoeoaa,
ieiae?aieoeoaa» (sseoa, 1996, 1997); 6 I?aeia?iaei?e iaoeiai-i?aeoe/i?e
eiioa?aioe?? «Iao?aaeeoe?eia ?ineeiieoeoai, aeieia?y ? caei?ia’y
(Aeoooa, 1997); 4 I?aeia?iaei?e eiioa?aioe?? c iaaee/ii? aioai?ee (Ee?a,
1997); iaoeiai-i?aeoe/i?e eiioa?aioe?? aei 110-??//y a?ycueie?ea?i?
«Iieiaee» (?aiaoi??y, 1997), ia i?aeia?iaeieo eo?nao c iiaeaethaaiiy
«Crop Weather Modell?ng» a ?c?a?e? (1998 ?.).

Ioae?eaoe??. Ca oaiith aeena?oaoe?? iioae?eiaaii 12 iaoeiaeo
?ia?o, a ?o /ene?: o?e — o ca??ieeao iaoeiaeo ?ia?o, n?i — o iaoa??aeao
eiioa?aioe?e, iaeei ?ioi?iaoe?eiee eenoie ? iaoiaee/i? ?aeiiaiaeaoe??.

No?oeoo?a oa ianya aeena?oaoe??. Aeena?oaoe?y neeaaea?oueny c
anooio, naie ?icae?e?a, aeniiae?a ? ?aeiiaiaeaoe?e ae?iaieoeoao,
aeeeaaeaia ia 226 noi??ieao iaoeiiieniiai oaenoo, i?noeoue 17 oaaeeoeue,
45 ?enoie?a ? 18 aeiaeaoe?a. A nieneo aeei?enoaieo aeaea?ae iaaaaeaii
217 iaeiaioaaiue, a ?o /ene?, 92 caei?aeiii?.

INIIAIEE CI?NO ?IAIOE

CI?NO AeINE?AeAEAIUe, OIIAE ? IAOIAeEEA ?O I?IAAAeAIIss

Aeine?aeaeaiiy i?iaiaeeeeny a 1995-1998 ??. o a?aeae?e? iiaeo
oaoi?/ieo ? e?ea?nueeeo eoeueoo? Aea?aeaaiiai I?e?onueeiai aioai?/iiai
naaeo (AeIAN) oa aa?iiaoai?ieia?/i?e noaioe?? «I?e?onueeee naae»,
?icoaoiaaii? iiaeeco a?ae aeine?aeii? ae?eyiee. I?e aiae?c? iniiaieo oac
?icaeoeo aeei?enoiaoaaeenue aaaaoi??/i? aeai? c oaiieia?? a?aeae?eo a
eiiieaen? c aaaaoi??/ieie aeaieie aa?iiaoainoaioe?? «I?e?onueeee naae».
Iniaeeao aaey/i?noue ca i?iiiceoe?? c iie?auaiiy ?iaioe ? oe?ii?
caoaaaeaiiy ca ci?noii aeena?oaoe?? aaoi? aeneiaeth? ia/aeueieeo
aa?iiaoainoaioe?? «I?e?onueeee naae», eaiaeeaeaoo aaia?ao?/ieo iaoe
Ae.?.Oo?na.

sse ia?eoe aeine?aeaeaiue aoee aeei?enoai? 12 aeae?a, ui
aeeth/athoue 15 aeioei?a /aa?aoeth, io?eiaieo ii aeaeaeoonai oa
?io?iaeoeiaaieo a AeIAN: Thymus vulgar?s L. — ?io?iaeoeoe?eiee iiia?
14232, AeIAN; T. vulgar?s L. — 53777, I?ia//eia; T. vulgar?s L. — 16275,
Oaaeoea??y; T. str?atus Vahl. — 32589, ?oae?y,; T. puleg?o?des L. —
32089, ?oae?y; T. n?tens Lamott. — 33889, O?aioe?y; T. r?chard?? Pers.
Subsp. n?t?dus (Gass) — 25889, *ao?y; T. mast?ch?na L. — 46389,
I?aea?eaiaee; T. transcaucas?cus Ronn. — 16786, A?oc?y; T. kotschyanus
Bo?ss. et Hoken. — 66574, 66974, A??iai?y; T. serpyllum L. — 12786,
Oaaeoea??y; T. caucas?cus W?lld. ex Ronn-16686, A?oc?y; T. camphoratus
Hoffm. et L?nk — 55377, Ii?ooaae?y; T. comosus Heuffel ex Gr?seb. —
21989, ?oioi?y.

Iaoiaeeea iieueiaeo aenia?eiaioaeueieo aeine?aeaeaiue
a?aeiia?aeaea aeiiaai Iaoiaeeee iieueiaeo aeine?ae?a, i?eeiyoi? o ?AE?
(N?ioa?iiieue, 1977). Ia?iaeo aeaieo i?e?iee?iaoe/ieo ?
aa?iiaoai?ieia?/ieo niinoa?aaeaiue i?iaiaeeee ca iaoiaeeeaie
E.N.Eaeue/aanueei? (1971), I.?.N?i?oeeii? (1973). I?e iieno aeeoo?aeo
oi?i, iniaeeainoae iaaiiiooai?aiiy o ??cieo aeae?a /aa?aoeth
aeei?enoiaoaaee ?iaioe Ie.A.Oaaei?iaa (1962), ?.A.Na?aa?yeiaa ?
O.?.Na?aa?yeiai? (1969), A.I.Aieoa?aa ( 1968, 1973), sseana (J.Jalas,
1972), I.A.Eeieiaa (1973), TH.E.Iaieoeueeiai (1978), A.A.Aia?iie (1990).

Oaiieia?/i? niinoa?aaeaiiy ca ?inoii ? ?icaeoeii ia’?eo?a
aeine?aeaeaiue i?iaiaeeee ca iaoiaeeeith ?.I.Aaeaeaiaia (1974) c
aeayeeie ci?iaie ? aeiiiaiaiiyie noiniaii aei eoeueoo?e, aea/aiiy
a?ieia?/ieo iniaeeainoae oea?o?iiy — ca iaoiaeeeith A.I.Iiiiia?ueiaa
(1960).

Noaoenoe/ia ia?iaea iaea?aeaieo ?acoeueoao?a i?iaaaeaia iaoiaeaie
ei?aeyoe?eiiai, ?aa?an?eiiai oa o?ueiooaeoi?iiai aeenia?n?eiiai aiae?c?a
ia ia?niiaeueiiio eiii’thoa?? ?BM Pent?um 166 MMX (Aeini?oia, 1968;
Iao??iue, E??ia, 1971; Caeoeaa, 1973; Ao?o?, 1982; Ai?i?ae/, 1984;
Ni??y?a, Aioiai, 1985; Aethe, 1997).

A?IEIA?*I? INIAEEAINO? *AA?AOeTH IA I?AAeAIIIIO AA?AC? E?EIO

Eiiieaenia aea/aiiy aeieiai-a?ieia?/ieo iniaeeainoae aeae?a
/aa?aoeth a iiaeo oiiaao ae?iuoaaiiy aeicaieeei anoaiiaeoe, ui
?io?iaeooeaioe i?ioiaeyoue iiaiee oeeee ?icaeoeo (a?ae ii/aoeo
aaaaoaoe?? aei aeic??aaiiy ian?iiy) a na?aaeiueiio ca 88 — 121 aeaiue.
?acoeueoaoe aeine?aeaeaiue a?aeia?aaeai? ca aeiiiiiaith niieo/aiiai
oaiiniaeo?a . A caeaaeiino? a?ae o?eaaeino? oac iioiaaiaco ? no?ie?a
oea?o?iiy, an? aea/ai? aeaee ?icae?eai? ia /ioe?e oaiia?oie:
?aiiueiea?oo/?, na?aaeiueiea?oo/?, i?ciueiea?oo/? ? ie?aii a /aoaa?oo
a?oio aeae?eaiee /aa?aoeue eaioi?iee, ui a?ae??ciy?oueny a?eueo i?ci?ie
no?ieaie oea?o?iiy ? aeic??aaiiy ian?iiy. Aiae?c aeaieo iieacaa, ui
ii/aoie oea?o?iiy iaea?eueo ?aiiueiea?oo/eo aeae?a niinoa??aa?oueny a
ia?oeo /eneao o?aaiy, i?e i?aeaeuaii? na?aaeiueiaeiaiai? oaiia?aoo?e
iia?o?y aei 12-13 N (T. r?chard?? ssp. n?t?dus, T. caucas?cus).
I?ci?oa, o ae?oa?e aeaeaae? o?aaiy, eiee iia?o?y i?ia??aa?oueny aei 16 N
? aeua, caoea?oathoue T. vulgar?s, T. str?atus, T. mast?ch?na, T.
transcaucas?cus, T. kotschyanus, T. serpyllum, T. comosus, T.
puleg?o?des. I?e e?ioe? o?aaiy — ia ii/aoeo /a?aiy, i?e
na?aaeiueiaeiaia?e oaiia?aoo?? iia?o?y 19-20 N, ii/eia?oueny ?ice?eaaiiy
ia?oeo ea?oie o T. n?tens, T. camphoratus. Aeiaiaee ?eoi ?icioneaiiy
ea?oie /aa?aoeth ia? iaeei iaeneioi — a aeaa’youe — aeanyoue aiaeei
?aieo. Iioeiaeueieie oiiaaie ? oaiia?aoo?a iia?o?y 19-20 N ? a?aeiinia
aieia?noue 55-65 %.

Ianiaa oea?o?iiy ?aiiueiea?oo/eo aeae?a ianoa? i?e iaeiie/aii? noi
aeoeaieo oaiia?aoo? iia?o?y aeua 5 N aeecueei 708 N, na?aaeiueiea?oo/eo
— 871 N, i?ciueiea?oo/eo — 966 N ? /aa?aoeth eaioi?iiai — 1233 N.
Iiiiaeaiiy aaaaoaoe?? a on?o aeine?aeaeoaaieo aeae?a ianoa? i?e
i?aeaeuaii? na?aaeiueiaeiaiai? oaiia?aoo?e iia?o?y aei 5 N.
O?c?ieia?/iee i?i?ioi oace aooii?caoe?? aeey /aa?aoeth cae/aeiiai neeaa
10.5 N, a aeey /aa?aoeth eaioi?iiai — 11.8 N. Ieaeiy iaaea aeey ii/aoeo
oea?o?iiy oeeo aeae?a a?aeiia?aeii neeaea 14.9 N ? 16.8 N. I?iaaaeai?
aeine?aeaeaiiy i?aeoaa?aeaeothoue a?ieia?/io iio?aao /aa?aoeth
eaioi?iiai a a?eueo aenieeo ??aiyo oaiia?aoo?, iaiao?aeieo aeey
ii?iaeueiiai ?icaeoeo ? ianoaaaiiy oac aooii?caoe?? ? oea?o?iiy, ui
iaiao?aeii a?aoiaoaaoe i?e ?aeiioaaii? oeueiai aeaeo. Aeai? na?aaei?o
aaaaoi??/ieo noi aeoeaieo oaiia?aoo? iia?o?y aeua 5 N iiaeooue aooe
aeei?enoai? i?e neeaaeaii? i?iaiico ianoaaaiiy iniiaieo oac ?icaeoeo
/aa?aoeth.

Aeey aeyaeaiiy e?eoe/ieo ia??iae?a iio?aae /aa?aoeth aei aieiae
aoa aeei?enoaiee iaoiae ?aa?an?eiiai aiae?co, ia iniia? yeiai iaea?aeaii
??aiyiiy ca’yceo:

Y=0,35+0,007 X1+0,005 X2,
(1)

Aea Y — o?iaeae ne?iaeie c 1 i2; O1 — noia iiaae?a ca eeiaiue ? na?iaiue
(ii); O2 — noia iiaae?a ca o?aaaiue (ii) c eiao?oe??ioii ei?aeyoe??
R=0.830 0.080 ? anoaiiaeaii aeaa e?eoe/ieo ia??iaee, a?aeiia?aeaeueieo
ca oi?ioaaiiy o?iaeath ne?iaeie: e?oi? a?ae?inoaiiy iaaii?a ae?oai?
aaia?aoe?? a eeii?-na?ii? ? ia??iae ?ioaineaiiai ?inoo aaniith, a
o?aai?.

A?IO?I?*IA OA?AEOA?ENOEEA AO??II? IE?? *AA?AOe?A IA

I?AAeAIIIIO AA?AC? E?EIO

Ie?y /aa?aoeth eieae?ciaaia o ainueieee?oeiieo, /ioe?e-, aeai- oa
iaeiiee?oeiieo caeiceao ii an?o i?aaiao ?ineeie: noaae?, eenoeao,
a?ii/eao, /aoa/oe?, i?eoea?oeao. Ia?aaaaeia cia/aiiy iathoue aaeee?
ainueieee?oeii? caeicee, ui yaeythoue niaith a?n?i ie??ooai?thaaeueieo
ee?oei. A?ae??ciyany aaeeeith e?euee?noth oeeo caeicie /aa?aoeue
eaioi?iee. Na?aaei?e ?ici?? caeicie o a?eueoino? aeae?a noaiiaea 60-80
iei. Iaea?eueoa /enei caeicie aeyaeaii o i?ae?aaa?’? /aoa/ee, iaeiaioa —
ia iiaa?oi? noaaea. ?nio? o?niee ca’ycie i?ae ianiaith /anoeith ao??ii?
ie?? oa e?euee?noth ao??iie?eieo caeicie o i?ae?aaa?’? /aoa/ee ? ia
iiaa?oi? eenoiai? ieanoeiee. Aeaee c a?eueoith e?euee?noth caeicie
oa?aeoa?ecothoueny a?eueo aenieith ianiaith /anoeith ao??ii? ie??. O
aeae?a c iacia/iei /eneii caeicie a?aecia/aii iecueeee ai?no ianea.
Io?eiai? ?acoeueoaoe aeia?a ocaiaeaeothoueny c aeaieie I. A. Ianoa?aiei
(1935) ii ?icia?eio ? A. Ae. ?aaioyaiaa (1983) ii eaaaiae?. Iaea?eueoa
e?euee?noue ie?? neioaco?oueny a nooea?ooyo, aeaui iaioa ?? o eenoeao ?
i?aeoe/ii a?aenooiy a noaaeao. O a?eueoino? aea/aieo aeae?a nooea?ooy
neeaaeaee a?eueoo /anoeio ea?oeiai? ne?iaeie. A?ae??ciyeeny a?ae ?ioeo
aeae?a /aa?aoeue cae/aeiee (53777) ? /aa?aoeue oaeaaaiii?ecaieoaaoee, a
yeeo /anoea eenoe?a noaiiaeea 40-50 %. E??i oiai, a eenoeao /aa?aoeth
eaioi?iiai oa /aa?aoeth oaeaaaiii?ecaieoaaoiai aoei a?aecia/aii aeineoue
aenieee ai?no ao??ii? ie?? (4-5 % a?ae nooi? ?a/iaeie), ui aeeeeea?
?ioa?an aeey naeaeoe??.

Ianiaa /anoea ao??ii? ie?? /aa?aoeth aa??th? a oe?ieeo iaaeao —
a?ae 0.4 % o /aa?aoeth iiaco/iai aei 10.1 % o /aa?aoeth eaioi?iiai.
Ei?aeyoe?eiee aiae?c ia aeyaea ca’yceo i?ae o?iaeaei?noth oa ianiaith
/anoeith ao??ii? ie?? (r=0.01), ui na?ae/eoue i?i iacaeaaeia
oniaaeeoaaiiy oeeo iciae ? iiaeeeaino? iaea?aeaiiy aaiioei?a, ui
ii?aeiothoue aenieo o?iaeaei?noue c aenieei ai?noii ianiai? /anoee
ao??ii? ie??. I?ioa, aeyaeaiee i?yiee ei?aeyoe?eiee ca’ycie i?ae
i?iaeoeoeai?noth ? ianiaith /anoeith ao??ii? ie?? (r=0.55), ui aeicaiey?
aanoe naeaeoe?eiee a?aea?? ?ineei ca aenieei ai?noii ao??ii? ie?? o
aaiioei?a. Aeae?eaii /ioe?e aeaee, ui iathoue iaaio oe?ii?noue (oaae. 1)
— /. eaioi?iee, /. oaeaaaiii?ecaieoaaoee, /. aeioeiee ? /. cae/aeiee
(53777).

Aiane?aeie a?io?i?/ieo aeine?aeaeaiue a ao??i?e ie?? /aa?aoeth
?aeaioeo?eiaaii 25 eiiiiiaio?a. Aeii?ioth/eie ? oeiie, ea?aae?ie,
e?iaeiie ? 1.8-oe?iaie (oaae. 2). Anoaiiaeaii, ui iaeneiaeueiee aeo?ae
oeiieo a?aei?/aiee ia ii/aoeo oea?o?iiy. A aeeiai?oe? ai?noo e?iaeiieo
aeae?eai? aeaa iaeneioio — ia ii/aoeo ? a e?ioe? oea?o?iiy. Iaeneiaeueia
e?euee?noue 1.8-oe?iaiea a?aecia/aia o oac? ianiaiai oea?o?iiy.
Aecia/ai? o?ceei-o?i?/i? iieacieee ao??ii? ie?? 12 aeae?a /aa?aoe?a a
oiiaao

Oaaeeoey 1

Oa?aeoa?enoeea aeae?a /aa?aoeth c iniiaieo ooee?oa?ieo iciae (1995-1997
??.)

?io?iaeooe?eiee iiia?

AeaeIioiaeaeaiiyO?iaeae ne?iaeie c 1 ?ineeie, aIaniaa /anoea ao??ii?
ie??, % a?aeI?iaeoe oeai?noue1 ?ineeie, aAi?no iniiaiiai eiiiiiaioo
ao??ii? ie??, %ne?i? ianeaanietho-ii nooi?
iane14232T.vulgarisAeIAN200-6800,25-0,410,66-2,060,5-2,8Oeiie
59,853777T.vulgarisI?ia//eia142-6900,30-0,591,03-2,220,4-4,1Oeiie aei
80,016275T.vulgarisOaaeoea??y98-3000,20-0,640,57-2,820,2-1,9E?iaeiie aei
90,032589T.striatus?oae?y180-5400,34-1,251,03-4,030,6-6,8Oeiie aei
74,032089T.pulegioides?oae?y230-8700,36-0,901,09-3,180,8-7,8Oeiie aei
73,033889T.nitensO?aioe?y200-3800,42-0,761,24-2,170,8-2,9Oeiie
66,125889T.richardii
ssp.nitidus*ao?y90-1100,88-1,243,00-6,200,8-1,4Oeiie
55,746389T.mastichinaI?aea?eaiaee110-2000,35-0,601,25-2,330,4-1,2Ea?aae?
ie
59,316786T.transcau-casicusA?oc?y30-840,45-1,000,83-3,130,1-0,8Ea?aae?ie
62,566574T.KotschyanusA??iai?y200-4000,34-0,541,03-1,760,7-2,2Oeiie
69,966974T.kotschyanusA??iai?y160-2440,48-0,601,35-2,120,8-1,5Oeiie
66,512786T.serpyllumOaaeoea??y56-1200,02-0,100,04-0,370,0-0,1Oeiie
28,316686T.caucasicusA?oc?y160-2000,34-0,640,97-2,700,5-1,3E?iaeiie
83,655377T.camphoratusIi?ooaae?y80-5340,74-4,562,07-10,10,6-10,1,8-Oe?ia
ie aei 80,021989T.comosus?oioi?y200-3500,38-0,981,27-2,970,8-3,4Oeiie
74,4

I?aaeaiiiai aa?aaa E?eio. I?e ?icaeyae? o?ce/ieo eiinoaio ie??,
iaea?aeaieo ia ??cieo noaae?yo iioiaaiaco, niinoa??aaeany caaaeueia
caeiiii??i?noue o ci?ithaaii? ie?? — i?aeaeuaiiy u?eueiino? ?
eiao?oe??ioa ?ao?aeoe?? a?ae oace aooii?caoe?? aei aeic??aaiiy ian?iiy.

Aeine?aeaeai? aeioeie /aa?aoe?a ca?oiiaai? ca neioacii
oa?iaii?ae?a. Ia?oa a?oia aeae?a — oeiie-ea?aae?ieueia, neeaaea?oueny c
aeaio oaiioei?a: oeiieueiiai (neioaco? oeiie a iaaeao a?ae 48 aei 80 %)
oa ea?aae?ieueiiai (ea?aae?iea aei 63 %). Ae?oaa a?oia — e?iaeiieueia
(e?iaeiiea a?ae 70 aei 90 %). ? o?aoy — oe?iaieueia (e?euee?noue
1.8-oe?iaiea aeinyaa? 70-80 %). Oe?eaai a?aecia/eoe, ui na?aae c?ace?a
/aa?aoeth cae/aeiiai aeyaeaii aea? oaii?ane, ui neioacothoue oeiie aei
80% ? e?iaeiie aei 90 %, ui na?ae/eoue i?i iacaeaaeia oniaaeeoaaiiy oeeo
iciae (oa?iaii?ae?a) ? iiaeeeaino? noai?aiiy iiaeo aaiioei?a ?ineei c
caaeaiei o?i?/iei neeaaeii.

Oaaeeoey 2

Ai?no iniiaieo eiiiiiaio?a ao??ii? ie?? aeae?a /aa?aoe?a

o oac? ianiaiai oea?o?iiy

A e ae e / a a ? a oe
thEiiiiiaioeT.vulgarisT.striatusT.pulegioidesT.nitensT.richardiiT.mastic
hinaT.transcaucasicusT.kotschyanusT.serpyllumT.caucasicusT.camphoratusT.
comosus123 -I?iai0,70,90,40,71,01,20,73,10,70,30,2Eaioai
0,40,10,30,60,20,90,30,70,10,3-I?iai 0,20,20,10,30,20,20,20,41,0Naa?iai
0,10,10,11,30,20,20,11,10,92,0I??oeai
0,10,10,10,10,10,90,50,5 -Oa?i?iai11,20,10,81,21,21,60,82,50,80,3Eeiiiai
0,30,30,20,30,30,20,30,50,51,8-Oe?iaie
0,30,40,11,01,61,51,21,01,61,01,773,51,2-Oa?i?iai8,49,91,17,711,16,814,4
6,29,47,61,80,32,1n-Oeeiie
13,912,50,811,619,89,59,04,64,25,05,60,20,76,7Eaioi?a
1,30,80,31,00,10,31,0E?iaeiie
1,81,976,02,93,12,82,32,40,73,04,183,63,63,2Ea??io?eai
1,92,40,93,40,71,31,01,01,012,91,20,2Oa?i?i.-4-ie
0,31,10,61,01,60,51,20,31,41,1-Oa?i?iaie0,20,66,4Ai?iaie
0,60,31,21,80,51,40,21,81,84,80,60,7Oeiie
59,859,116,758,248,066,155,716,22,269,928,311,81,374,4Ea?aae?ie
3,13,90,92,41,82,71,759,362,53,11,91,10,24,4

Iiaeaethaaiiy aca?iiae?? aeae?a /aa?aoeth c aeieia?/ieie
oaeoi?aie. A?aeiii, ui i?ae aeaeaie ? iaaeieeoi?i na?aaeiaeuai iino?eii
a?aeaoaa?oueny aca?iiae?y. Aaee/eia iciaee (oaiioeio) caeaaeeoue a?ae
aeaio oaeoi??a: niaaeeiaiai, coiiaeaiiai aaiioeiii ? ianiaaeeiaiai,
coiiaeaiiai na?aaeiaeuai.

Aeey eiiieaenii? ioe?iee oaiioeio c iaoith aeae?eaiiy
ia?niaeoeaieo aeae?a, iaea?eueo no?eeeo aei aieeao aeieia?/ieo oaeoi??a,
aoea ?ic?iaeaia iiaeaeue i?iaeoeoeaiino? /aa?aoeth (oaae. 3). Aeyaeaii,
ui ianiaa /anoea ao??ii? ie?? ia 79 % coiiaeaia aaiioeiii. I?ioyaii
aaaaoaoe?? a?ae oace aooii?caoe?? aei aeic??aaiiy ian?iiy niinoa??aaeiny
iinooiiaa cieaeaiiy neioaco ao??ii? ie??. Ia a?aei?io a?ae ianiai?
/anoee ao??ii? ie??, ia i?iaeoeoeai?noue /aa?aoeth aieeaathoue ye
aaiioei, oae ? aeieia?/i? oaeoi?e. Iieacaii, ui /anoea aieeao aaiioeio
neeaea 53 %, a aeieia?/ieo oaeoi??a — 33 %. Iaea?eueoa i?iaeoeoeai?noue
?ineei /aa?aoeth niinoa??aa?oueny a ia??iae oea?o?iiy. Aeenia?n?eiee
aiae?c i?aeoaa?aeea ia?niaeoeai?noue /ioe?ueio aeae?eaieo aeae?a: /.
eaioi?iiai, /. oaeaaaiii?ecaieoaaoiai, /. aeioeiiai ? /. cae/aeiiai
(53777). Aeae?eai? c?acee ?iciiiaeai? aeeaoethaaiiyi, aeeth/ai? o
naeaeoe?eiee i?ioean, noai?aiee iaoi/iee. Aiae?c ao??iie?eiino? a
iniiai? oace ?icaeoeo ca 1995-1997 ?iee iieacaa, ui iioeiaeuei? oiiae
aeey neioaco ao??ii? ie?? /aa?aoeth oa?aeoa?ecothoueny na?aaeiueith
oaiia?aoo?ith iia?o?y 17-19 N ? e?euee?noth iiaae?a, ui ia ia?aaeuothoue
5-10 ii ca i?aeoaciaee ia??iae.

Oaaeeoey 3

Iiaeaeue aeenia?n?eiiai aiae?co aeae?a ?iaeo /aa?aoeue

Aeaea?aei i?ieeaino?Nooi?iue naiaiaeeNa?aaei?e eaaae?aoI/?eoaaia
cia/aiiy na?aaeiueiai eaaae?aooIiaoi?iino? (R)r-1?iee (G)g-1I1
2e+r2vgf+rf2vg+rv2gf+rvf2gOace (F)f-1M22e+r2vgf+rg2vf+rv2gf+rvg2fAeaee
(V)v-1M32e+r2vgf+rf2vg+rg2vf+rfg2vG-F(g-1)(f-1)M42e+r2vgf+rv2gfV-G(v-1)(
g-1)M52e+r2vgf+rf2vgV-F(v-1)(f-1)M62e+r2vgf+rg2vfV-G-F(v-1)(g-1)(f-1)M72
e+r2vgfIiieeea (V-R)(v-1)(r-1)fgM82e

*AA?AOeUe EAIOI?IEE — IIAA IA?NIAEOEAIA AO??IIE?EIA ? E?EA?NUeEA ?INEEIA

C aea/aieo aeae?a iaea?eueoee ?ioa?an ye ao??iie?eia ? e?ea?nueea
?ineeia aeeeeea? /aa?aoeue eaioi?iee, ?io?iaeoeiaaiee c Ii?ooaae??.
Aea/aiiy aiaoii??, a?ieia?? ? a?io?i?? iiioeyoe?? /aa?aoeth eaioi?iiai
aeicaieeei aeyaeoe cia/i? a?aei?iiino? ia o?eueee ca na?aaei?ie
iieacieeaie ao??iie?eiino?, aea ? ca ?ioeie ooee?oa?ieie iciaeaie.
Aeae?eai? ia?niaeoeai? oi?ie c aenieei ai?noii ao??ii? ie??,
?aeiiaiaeiaai? aeey aaaaeaiiy a eoeueoo?o ia i?aaei? Oe?a?ie (oaae. 4).

Oaaeeoey 4

Oa?aeoa?enoeea iniiaieo ainiiaea?nueei-oe?iieo iciae aeae?eaieo
ni?oic?ace?a /aa?aoeth eaioi?iiai (1995-1997 ??.)

Iiia?O?iaeaeIaniaa/anoeaCa?? AIAi?no iniiaieoni?oi-ne?iae-yo??ii?ie??,(a
ia?a-eiiiiiaio?a,c?aceaie, oe/aa%a?ae?aooieo),%ne?i? ianeaaniethoii
nooi?
ianeea/aa1,8-oeeiaiee?iaeiie-oa?i?iaie5/41-5722,287,48163,974,12,16,05/5
6-4432,286,1698,375,02,35,12/5
-6392,256,2587,274,02,68,35/29-1482,285,18110,474,42,96,7

Aecia/ai? iniiai? iciaee i?iaeoeoeaiino? /aa?aoeth eaioi?iiai: /enei
caeicie, e?euee?noue ea?oie, iana ? ae?aiao? oeaio?aeueiiai nooea?ooy.
Oea aeicaiey? cae?enieoe iai?aaeaiee i?aea?? aaouee?anueeeo ia? aeey
no?auoaaiiy ? noai?thaaoe iia? oi?ie /aa?aoeth, ui iathoue oe?ieo
aaiaoe/io i?enoiniaai?noue ? ii?aeiothoue aenieo ao??iie?ei?noue c
iaeneiaeueiei ai?noii oe?iieo eiiiiiaio?a. Aea/aiiy o?i?/iiai neeaaeo
ao??ii? ie?? /aa?aoeth eaioi?iiai iieacaei eiai iiae?ai?noue c ianeii
aaeae?ioa. Ia a?aei?io a?ae inoaiiueiai, a ao??i?e ie?? /aa?aoeth
eaioi?iiai a?aenooi?e a-oaeaiae?ai, yeee aeeeeea? iia?/i? aea?a?/i?
?aaeoe??.

NOENEA AIOAI?*IA OA?AEOA?ENOEEA AeAAIAAeOessOE AEAe?A *AA?AOeTH

Ia iniia? eiiieaeniiai aeine?aeaeaiiy c?iaeaii noeneee iien
aiaoiii- ii?oieia?/ieo, a?ieia?/ieo ? oeeoiaaiaoe/ieo iniaeeainoae 12
aeae?a /aa?aoeth c ??cieo e?a?i na?oo a oiiaao I?aaeaiiiai aa?aaa E?eio.

Aecia/ai? i?aaiieaioe/i? ? o?ceei-o?i?/i? iieacieee ao??ii? ie??
?io?iaeooeaio?a (eie??, caiao, ?ic/eii?noue, u?euei?noue, eiao?oe??io
?ao?aeoe??, ianiaa /anoea). Aeaoaeueii aea/aiee ? iaaaaeaiee
eiiiiiaioiee neeaae ?o ie??. Aeey eiaeiiai aeaeo aecia/aii niiaoe/ia
/enei o?iiinii, aeey T. camphoratus Hoffm. et L?nk ? T. r?chard?? Pers.
Subsp. n?t?dus — aia?oa.

AENIIAEE

1. ?icoe?aia eieaeoe?y aeae?a ?iaeo Thymus L.: ?io?iaeoeiaai?
o?noue iiaeo aeae?a /aa?aoeth (T. argaeus Bo?ss et Bal., T. long?caul?s
C. Presl, T. cap?tellatus Hoffm. et L?nk, T. zyg?s Loefl ex L. ssp.
sylvestr?s, T. pannon?cus All., T. c?tr?odorus Pers.), noai?ai?
i?aeaeaeia? a?a?eaee oa iie?iei?aei? oi?ie. Aeyaeai? oe?ie? iiaeeeaino?
oi?iiooai?thaaeueiiai i?ioeano aiaoiii-ii?oieia?/ieo ? o?c?ieiai-
a?io?i?/ieo iciae ? aeanoeainoae. Aeiaaaeaia ia?niaeoeai?noue
aeei?enoaiiy iaea?aeaieo oi?i ye iiaiai aeo?aeiiai iaoa??aeo aeey
?io?iaeoeoe?? ? naeaeoe??.

2. Aeyaeaii aeaa e?eoe/ieo ia??iaee a iio?aa? /aa?aoeth aei
aieiae, a?aeiia?aeaeueieo ca oi?ioaaiiy o?iaeath ne?iaeie: e?oi?
a?ae?inoaiiy iaaii?a ae?oai? aaia?aoe?? (eeiaiue-na?iaiue) ? ia??iae
?ioaineaiiai ?inoo aaniith (o?aaaiue). I?e a?aenooiino? cia/ieo iiaae?a
(a?eueoa 10 ii) a oe? ia??iaee, aeey i?aeaeuaiiy o?iaeath ea?oeiai?
ne?iaeie /aa?aoeth, iaiao?aeii i?iaiaeeoe aaaaoaoe?ei? iieeae ii?iith,
ui caaacia/o? caieiaeaiiy a?oioo ia ieae/a 70-80% IA (iaeiaioi?
aieiai?iiino?).

3. Iiiiaeaiiy aaaaoaoe?? a on?o aeine?aeaeaieo aeae?a ianoa? i?e
i?aeaeuaii? na?aaeiueiaeiaiai? oaiia?aoo?e iia?o?y aei 4.5 N.
O?c?ieia?/iee i?i?ioi oaiia?aoo? aeey oace aooii?caoe?? T. vulgar?s
noaiiaeoue 10.5 N, T. camphoratus — 11.8 N; aeey ii/aoeo oea?o?iiy
a?aeiia?aeii noaiiaeoue 14.9 N ? 16.8 N. Ianiaa oea?o?iiy
?aiiueiea?oo/eo aeae?a ianoa? i?e iaeiie/aii? noi aeoeaieo oaiia?aoo?
iia?o?y aeua 5 N aeecueei 708 N, na?aaeiueiea?oo/eo — 871 N ,
i?ciueiea?oo/eo — 966 N ? /aa?aoeth eaioi?iiai — 1233 N. Oea iaiao?aeii
a?aoiaoaaoe i?e neeaaeaii? i?iaiico oea?o?iiy ? ?aeiioaaii? /aa?aoeth.

4. Aecia/ai? iioeiaeuei? cia/aiiy iaoaiaeaiaio?a, iaiao?aeieo aeey
?ioaineaiiai oea?o?iiy ? a?ineioaco ao??ii? ie?? /aa?aoeth. Iaeneiaeueia
e?euee?noue ?ice?eoeo ea?oie niinoa??aaeany i?e oaiia?aoo?? iia?o?y
19-20 N ? a?aeiini?e aieiaino? 55-65 %. Iaea?eueo? cia/aiiy ianiai?
/anoee ao??ii? ie?? a?aecia/ai? i?e na?aaei?e oaiia?aoo?? iia?o?y 17-19
N ? e?eueeino? iiaae?a, ui ia ia?aaeuo? 5-10 ii ca i?aeoaciaee ia??iae.

5. ?ic?iaeaia iiaeaeue i?iaeoeoeaiino? /aa?aoeth:

X = V + m + f + Vf + g + Vg +
fg + Vfg + e,

_______________ ____
_______________ __

aaiaoe/iee aoaeo aoaeo
aca?iiae?y aoaeo

aeiei- aeae — aeiaaeeiaeo

a?/ieo na?aaeiaeua i?e/ei

oaeoi??a

ui aeicaiey? aecia/eoe iaoai?ci iaeiie/aiiy ao??ii? ie?? ? aca?iica’ycie
nenoaie i?iaeoeoeaiino? oa ?? aeaiaio?a c a?ae?ioa?i?/ieie oaeoi?aie. Ia
i?iaeoeoeai?noue /aa?aoeth aieeaathoue ye aaiioei (52.9 %), oae ?
oaeoi?e na?aaeiaeua (32.7 %). Ia a?aei?io a?ae i?iaeoeoeaiino?, ianiaa
/anoea ao??ii? ie?? a cia/i?e i??? coiiaeaia aaiioeiii (79.2 %).

6. I?iaeoeoeai?noue /aa?aoeth eaioi?iiai ei?aeth? c /eneii
caeicie, e?euee?noth ea?oie, ianith ? ae?aiao?ii oeaio?aeueiiai
nooea?ooy, ui aeicaiey? cae?enithaaoe iai?aaeaiee i?aea?? aaouee?anueeeo
ia? aeey no?auoaaiiy ? noai?thaaoe iia? aenieii?iaeoeoeai? ni?oe
/aa?aoeth, ui iathoue oe?ieo i?enoiniaai?noue ? ii?aeiothoue aenieo
ao??iie?ei?noue c iaeneiaeueiei ai?noii oe?iieo eiiiiiaio?a.

7. Aeine?aeaeaiiy a?ineioaco iniiaieo eiiiiiaio?a iieacaei, ui
iaeiie/aiiy oeiieo ? e?iaeiieo a ao??i?e ie?? /aa?aoeth a cia/i?e i???
caeaaeeoue a?ae oace ?icaeoeo ? iiaiaeieo oiia. Iaeneiaeueiee neioac
oeiieo o oeiieuei?e a?oi? (T. vulgar?s, T. puleg?o?des, T. str?atus)
a?aecia/aany ia ii/aoeo oea?o?iiy. Aeeiai?ea ai?noo e?iaeiieo o
e?iaeiieuei?e a?oi? (T. vulgar?s (16275)) iaea aeaa iaeneioio — ia
ii/aoeo ? a e?ioe? oea?o?iiy. Iaeneiaeueia e?euee?noue 1.8-oe?iaieo o
oe?iaieuei?e a?oi? (T. camphoratus) a?aecia/aia o oac? ianiaiai
oea?o?iiy. Aieea oiia iiaiaee ia ai?no iniiaieo eiiiiiaio?a a ao??i?e
ie?? /aa?aoeth i?aeoaa?aeaeo?oueny ?o ei?aeyoe?eieie ca’yceaie.

8. Aea/aiiy iiioeyoe?? /aa?aoeth eaioi?iiai aeyaeei iayai?noue
aeaio a?ioei?a: aeainoaoaaiai ? c iaaei?icaeiooith /ieia?/ith noa?ith
(Oe*N). ?ineeie c Oe*N iathoue i?aeaeuaio ao??iie?ei?noue ?
i?iaeoeoeai?noue, ui i?aaenoaaey? aaeeeee iaoeiaee oa i?aeoe/iee
?ioa?an.

9. Ii??aiyeueia aea/aiiy eiiiiiaioiiai neeaaeo ao??ii? ie??
/aa?aoeth eaioi?iiai ? aaeae?ioa iieacaei noiae?noue iniiaieo oa?iaiiaeo
niieo/aiue aeine?aeaeoaaieo ianae. Ia a?aei?io a?ae ianea aaeae?ioa, a
ao??i?e ie?? /aa?aoeth eaioi?iiai a-oaeaiae?ai ia aeyaeaiee, ui
aeicaiey? ?aeiiaiaeoaaoe eiai aeey ae?iuoaaiiy ia i?aaei? Oe?a?ie ye
iiai? oe?iii? e?ea?nueei? ? ao??iie?eii? ?ineeie.

10. ?acoeueoaoe aeine?aeaeaiiy aieeao aeieia?/ieo oaeoi??a ia
?icaeoie /aa?aoeth iieacaee aeioe?euei?noue eoeueoeaoaaiiy /aa?aoeth c
iaoith iaea?aeaiiy a?iiane a ianooaaao, ui caaacia/othoue iio?aae
oa?iaoeaaoe/ii? i?iieneiaino? a ne?iaei?. Oea aeicaiey? noaaeoe ieoaiiy
i?i ai?iaaaeaeaiiy ia i?aaei? Oe?a?ie aeae?eaieo aeae?a ?ineei: /.
eaioi?iiai, /. aeioeiiai, /. oaeaaaiii?ecaieoaaoiai ? /. cae/aeiiai.

?AEIIAIAeAOe?? AE?IAIEOeOAO

1. Aeey n?aae aeei?enoiaoaaoe na?aeic?a?aia ian?iiy /aa?aoeth aai c
oa?i?iii caa??aaiiy ia a?eueoa aeaio-o?ueio ?ie?a. N?aao ian?iiy a
iieueiaeo oiiaao e?aua anueiai i?iaiaeeoe a ia?o?e aeaeaae? o?aaiy c
aeeaeiith caeeaaeaiiy 0.5 — 1.0 ni aai iiaa?oiaaei niiniaii. Aeey
a?aeai?o ? ioe?iee iin?aiiai iaoa??aeo ?iaeo Thymus L. aeei?enoaoe
??aiyiiy ?aa?an?? W=-49.1+169.5 Py, aea W — noiae?noue, ?y —
u?euei?noue.

2. I?e aecia/aii? no?ie?a i?iaaaeaiiy aa?ioaoi?/ieo caoiae?a
ae?iuoaaiiy ne?ae a?aoiaoaaoe, ui aaniyia iiiiaeaiiy aaaaoaoe??
/aa?aoeth ii/eia?oueny i?e no?eeiio ia?aoiae? na?aaeiueiaeiaiai?
oaiia?aoo?e iia?o?y iiiaae 5 N.

3. Ca?? e?ea?nueei? ne?iaeie iaiao?aeii i?iaiaeeoe i?e noo?e
iiaiae? a ia??iae ianiaiai oea?o?iiy. Cae?aiiy /aa?aoeth cae/aeiiai,
oaeaaaiii?ecaieoaaoiai ? aeioeiiai ne?ae ii/eiaoe i?e iaeiie/aii?
aeoeaieo oaiia?aoo? iia?o?y aeua 5 C aeecueei 900 C, /aa?aoeth
eaioi?iiai — 1250 N.

4. I?e ieaioaaii? naeaeoe?eii-aaiaoe/ieo ?ia?o iaiao?aeii
aeoiaeeoe c oiai, ui aeiaiaee ?eoi ?ice?eooy ea?oie /aa?aoe?a ia? iaeei
iaeneioi i aeaa’yo?e-aeanyo?e aiaeei? ?aieo. I?aea?? aaouee?anueeeo ia?
aeey no?auoaaiiy ne?ae cae?enithaaoe c o?aooaaiiyi oniaaeeoaaiiy
iniiaieo iciae i?iaeoeoeaiino? ? a?ieia?/ieo iniaeeainoae (?ineeie c
iaaei?icaeiooith /ieia?/ith noa?ith (Oe*N) iathoue i?aeaeuaio
ao??iie?ei?noue ? i?iaeoeoeai?noue).

NIENIE INIIAIEO ?IA?O, IIOAE?EIAAIEO CA OAIITH AeENA?OAOe??

1. Ei?naeiaa N.I. Aeeyiea yeieiae/aneeo oaeoi?ia ia aeieiaeth
oeaaoaiey /aa?aoea // Athee. Ain. Ieeeone. aio. naaea. — 1997. — Aui.
78. — N. 50-54.

2. Ei?naeiaa N.I. Iniaaiiinoe iioiaaiaca /aa?aoea eaioi?iiai a
oneiaeyo THaeiiai aa?aaa E?uia // Athee. Ain. Ieeeone. aio. naaea. —
1997. — Aui. 78. — N. 55-57.

3. ?aaioyaia A.Ae., Oeuiaiei E.A., Ei?naeiaa N.I. *aa?aoe eaioi?iue
— ia?niaeoeaiia yoe?iianee/iia e eaea?noaaiiia ?anoaiea //Iniaaiiinoe
aeeeeiaoecaoeee iiiaieaoieo eio?iaeooeaioia, iaeaieeaathueo aeieiae/anee
aeoeaiua aauanoaa. — E?aniiaea?: EAAO eao. iau. aeie. e yeie. — 1995. —
N. 197-199.

4. Ei?naeiaa N.I., ?aaioyaia A.Ae. Iiaua ia?niaeoeaiua
eaea?noaaiiua ?anoaiey: Eioi?i. eenoie. — Neioa?iiieue: E?ui OeIOYE,
1998. — N 2-98. — 4 n.

5. Iaoiaee/aneea ?aeiiaiaeaoeee ii aicaeaeuaaieth e ia?a?aaioea
/aa?aoea eaioi?iiai / ?aaioyaia A.Ae., Oo?na Ae.E., Ei?naeiaa N.I. —
sseoa, 1997. — 15 n.

6. Ei?naeiaa N.I., ?aaioyaia A.Ae., Oo?na Ae.E., Aeiia?aaeia A.A.
Eco/aiea eiiiiiaioiiai ninoaaa yoe?iiai ianea iaeioi?uo aeaeia
/aa?aoea, eio?iaeooee?iaaiiuo a AIAN // Iaoa?. ?Y Iaaeaeoia?.
eiioa?aioeee “I?iaeaiu aeaiae?ieiaee, oeaaoiaiaenoaa, ieiaeiaiaenoaa,
aeiia?aaea?noaa e aeiiaeaeey». — Oii 1. — sseoa, 1996. — N. 90-93.

7. Ei?naeiaa N.I., ?aaioyaia A.Ae., Oo?na Ae.E., Oeuiaiei E.A.
Eco/aiea aeieiaee i?i?anoaiey naiyi ?iaea *aa?aoe i?e eio?iaeoeoeee ia
THaeiii aa?aao E?uia // Iaoa?. ?Y Iaaeaeoia?. eiioa?aioeee «I?iaeaiu
aeaiae?ieiaee, oeaaoiaiaenoaa, ieiaeiaiaenoaa, aeiia?aaea?noaa e
aeiiaeaeey». — Oii 1. — sseoa, 1996. — N. 94-96.

8. Ei?naeiaa N.I. I aeeyiee iiaiaeiuo oneiaee ia iaeiieaiea
yoe?iiai ianea o /aa?aoea a ia?eiae aeoeaiie aaaaoaoeee // Iaoa?. Y
Iaaeaeoia?. eiioa?aioeee «I?iaeaiu aeaiae?ieiaee, oeaaoiaiaenoaa,
ieiaeiaiaenoaa». — *anoue ??, Oeaaoiaiaenoai. — sseoa, 1997. — N. 44-48.

9. Ei?naeiaa N.I., ?aaioyaia A.Ae., Oo?na Ae.E. Eio?iaeoeoeey
iiauo aeaeia ?iaea Thymus L. // Iaoa?. 6 Iaaeaeoia?. iao/.-i?aeoe/.
eiio. «Iao?aaeeoeeiiiia ?anoaieaaiaenoai, yeieiaey e caei?iauea”. —
Neioa?iiieue. — 1997, Ae. 6-8. — N. 558.

10. Ei?naeiaa N.I. *aa?aoe eaioi?iue — ia?niaeoeaiia
eaea?noaaiiia ?anoaiea // A?ycaea/aaieoea «Iieiaee»/ Oac. aeiee.
iao/ii-i?aeoe/. eiio. e 110-eaoeth a?ycaea/aaieoeu «Iieiaee».
-Aaiaoi?ey. — naioya?ue 1997. — N. 75-76.

11. Ei?naeiaa N.I., Aenaiia TH.A., ?aaioyaia A.Ae. Eio?iaeoeoeey
?iaeia Thymus L. e Nepeta L. a oneiaeyo THAE // Iaoa?.
iaaeae.neii. “Iaoiaeieiae/aneea iniiau oi?ie?iaaiey, aaaeaiey e
eniieueciaaiey eieeaeoeee aaiaoe/aneeo ?ano?nia ?anoaiee». — Oa?ueeia.
-1996. — N. 175.

12. ?aaioyaia A.Ae., Ieo?ioaiiaa I.A., Eaaiiaa I.I., Ei?naeiaa
N.I. Iieo/aiea iaaeaeaeiauo aea?eaeia /aa?aoea e eo onei?aiiia
?aciiiaeaiea a oneiaeyo ?n v?tro // Iaoa?. iaaeaeoia?. iao/.-i?aeoe/.
eiio. «Ia?a?aaioea eaea?noaaiiiai nu?uey e i?iecaiaenoai oeoii?aia?aoia
aeey iaaeeoeeiu e naeueneiai oicyenoaa». — Aeiaou: IEOe AANOAO. -1996.
— N. 76.

Ei?naeiaa N.I. Aeieiai-a?ieia?/i? iniaeeaino? ? ao??iie?ei?noue
aeae?a ?iaeo Thymus L. ia I?aaeaiiiio aa?ac? E?eio. — ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
a?ieia?/ieo iaoe ca niaoe?aeuei?noth 03.00.05 — aioai?ea. — Aea?aeaaiee
I?e?onueeee aioai?/iee naae OAAI, sseoa, 1998.

Aeena?oaoe?y i?enay/aia aea/aiith aeieiai-a?ieia?/ieo iniaeeainoae
12 aeae?a ?iaeo Thymus L. a oiiaao I?aaeaiiiai aa?aaa E?eio. Aea/aia
a?io?i?y ? aeeiai?ea iaeiie/aiiy ao??ii? ie?? /aa?aoeth a iioiaaiac?.
?ic?iaeaia iiaeaeue i?iaeoeoeaiino? /aa?aoeth. Aea/ai? aeiiae /aa?aoeth
aei iniiaieo oaeoi??a ciai?oiueiai na?aaeiaeua. Aecia/ai? ? iaoaiaoe/ii
iaa?oioiaai? e?euee?ni? iieacieee iioeiaeueieo oa e?eoe/ieo oiia
oi?ioaaiiy o?iaeath ea?oeiai? ne?iaeie ? a?ineioaco ao??ii? ie??.
Aeae?eaii /ioe?e aeaee /aa?aoeth, ia?niaeoeaieo aeey ai?iaaaeaeaiue o
ae?iaieoeoai: T. camphoratus Hoffm. et L?nk., T. puleg?o?des L., T.
str?atus Vahl.,T. vulgar?s L. ?ic?iaeai? caniae ia?aeiii? aa?ioaoi?ee
ae?iuoaaiiy /aa?aoeth.

Eeth/ia? neiaa: /aa?aoeue, a?iii?oieia?y, aeieia?/i? oaeoi?e,
a?ineioac, ao??ia ie?y, i?iaeoeoeai?noue, iiaeaeue.

Ei?naeiaa N.I. Yeieiai-aeieiae/aneea iniaaiiinoe e
yoe?iiianee/iinoue aeaeia ?iaea Thymus L. ia THaeiii aa?aao E?uia. —
?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.05 — aioaieea. — Ainoaea?noaaiiue
Ieeeoneee aioaie/aneee naae OAAI, sseoa, 1998.

Aeenna?oaoeey iinayuaia eco/aieth yeieiai-aeieiae/aneeo
iniaaiiinoae 12 aeaeia ?iaea Thymus L. a oneiaeyo THaeiiai aa?aaa
E?uia. Eco/aia aeioeiey e aeeiaieea iaeiieaiey yoe?iiai ianea /aa?aoea
a iioiaaiaca. ?ac?aaioaia iiaeaeue i?iaeoeoeaiinoe /aa?aoea. Eco/aiu
o?aaiaaiey /aa?aoea e iniiaiui oaeoi?ai aiaoiae n?aaeu. Ii?aaeaeaiu e
iaoaiaoe/anee iainiiaaiu eiee/anoaaiiua iieacaoaee iioeiaeueiuo e
e?eoe/aneeo oneiaee oi?ie?iaaiey o?iaeay oeaaoi/iiai nu?uey e aeineioaca
yoe?iiai ianea. Auaeaeaii /aou?a aeaea /aa?aoea, ia?niaeoeaiuo aeey
aiaae?aiey a i?iecaiaenoai: T.camphoratus Hoffm. et Link., T.pulegioides
L., T.striatus Vahl., T.vulgaris L. ?ac?aaioaiu i?eaiu ia?ae/iie
aa?ioaoieee au?aueaaiey /aa?aoea.

Eeth/aaua neiaa: /aa?aoe, aeiii?oieiaey, yeieiae/aneea oaeoi?u,
aeineioac, yoe?iia ianei, i?iaeoeoeaiinoue, iiaeaeue.

Korsakova S.P. Ecology-b?olog?cal pecul?ar?t?es and
essent?al o?l of spec?es of the genus Thymus L. on the Southern coast
of the Cr?mea. — Manuscr?pt.

D?ssertat?on for the degree of cand?date of b?olog?cal
sc?ences on the spec?al?ty 03.00.05 — botany. — The State
N?k?tsky Botan?cal Garden, Yalta, 1998.

The d?ssertat?on ?s devoted to the study of the
ecology-b?olog?cal pecul?ar?t?es 12 spec?es of the genus Thymus L.
?n cond?t?ons of the Southern coast of the Cr?mea.
B?ochem?stry and dynam?c of accumulat?on essent?al o?l Thymus L. ?n
ontogenes?s ?s ?nvest?gated. A model of product?v?ty Thymus has been
worked out. The requ?rements Thymus to major factors of external
env?ronment are ?nvest?gated. Quant?tat?ve parameters of opt?mum and
cr?t?cal cond?t?ons ?n format?on of a crop of flower raw and
b?osynthes?s of essent?al o?l have been determ?ned and mathemat?cally
reasoned. Four perspect?ve spec?es Thymus for ?ntroduct?on ?n
manufacture are s?ngled out: T.camphoratus Hoffm. et L?nk.,
T.puleg?o?des L., T.str?atus Vahl., T.vulgar?s L. The technology of
pr?mary agr?cultural metods of grow?ng Thymus has been elaborated.

Key words: Thymus L., b?omorphology, ecolog?cal factors,
b?osynthes?s, essent?al o?l, product?v?ty, model.

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