IAOe?IIAEUeIA AEAAeAI?ss IAOE OE?A?IE

?INOEOOO A?Ae?IA?IEIA??

Ia i?aaao ?oeiieno

OAeE [574.58+(594:576.89)] (28)

TH?EOEIAOeUe AIEIAeEIE? ?AAIIAE*

AeAINOOEEIA? IIETHNEE OA ?O AIAeIA?IIOE ssE EIIIIIAIO A?Ae?IIA?ACEOA?IEO
NENOAI

03.00.17 — a?ae?ia?ieia?y

A a o i ? a o a ? a o

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

EE?A-1999

Aeena?oaoe?y ? ?oeiien.

?iaioa aeeiiaia a ?inoeooo? a?ae?ia?ieia?? IAI Oe?a?ie

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, n.i.n., Oa?/aiei Oa?an
Aiae??eiae/,

?inoeooo a?ae?ia?ieia?? IAI Oe?a?ie, caa. a?aeae?eii nai?oa?ii? oa
oaoi?/ii? a?ae?ia?ieia??.

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, n.i.n, Oaaoeiaa Ethaeieea
Aanee?aia, ?inoeooo a?ae?ia?ieia?? IAI Oe?a?ie, caa. a?aeae?eii
a?ae?ia?ieia?? ??/eiaeo nenoai.

eaiaeeaeao a?ieia?/ieo iaoe, E?ei/eoeueeee Iao?i sseiae/, Ee?anueeee
oi?aa?neoao ?i. O.Oaa/aiea, aeioeaio eaoaae?e ciieia??, a?ieia?/iiai
oaeoeueoaoo.

I?ia?aeia onoaiiaa: ?inoeooo ciieia?? ?i. Oiaeueaaocaia IAI Oe?a?ie,
i.Ee?a

Caoeno a?aeaoaeaoueny “2” ethoiai 1999 ?ieo i 15 aiae. ia can?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae.26.213.01 a ?inoeooo? a?ae?ia?ieia??
IAI Oe?a?ie ca aae?anith: 254210, i.Ee?a, i?. Aa?i?a Noae?ia?aaeo, 12.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? ?inoeoooo
a?ae?ia?ieia??.

Aaoi?aoa?ao ?ic?neaiee “22” a?oaeiy 1998 ?ieo

A/aiee nae?aoa? niaoe?ae?ciaaii? ?aaee,

eaiaeeaeao a?ieia?/ieo iaoe I.A.Nie?iiaa

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE.

Aeooaeuei?noue oaie. O ca’yceo ?c ?ioaineaiei i?e?iaeiei?enooaaiiyi,
ca?aaoethaaiiyi oa/?? ??/ie oa noai?aiiyi aiaeinoiaeu ci?ith?oueny
aeaeiaee neeaae oa oeaiioe/i? oa?aeoa?enoeee aeainooeeiaeo iiethne?a —
aaaeeeaiai eiiiiiaioo iae?iciiaaioino oa ia?eo?oiio i??nieo aiaeiei.

Aeainooeeia? iiethnee a?ae?a?athoue cia/io ?ieue a aiaeieo aeinenoaiao .
Oeae eean oaa?ei neeaaea? iaeio c a?oi, ui aeii?iothoue ii a?iian?.
Aeaoaeueia aea/aiiy aeainooeeiaeo iiethne?a iaiao?aeia aeey a??iiai
?icoi?iiy aaaaoueio a?ieia?/ieo i?ioean?a, ui a?aeaoaathoueny a
a?ae?iaeinenoaiao. Aoaeo/e o?eueo?aoi?aie-naaeeiaioaoi?aie, aeainooeeia?
iiethnee iaeii/anii ?c caaacia/aiiyi naaa ?aeath inaaeaeothoue o
aeaeyae? aaethoeiao?a cia/io e?euee?noue nanoiio, aeath/e ii/aoie
aeao?eoiei eaioethaai aeeaeaiiy a aaioin? i a?ae?a?ath/e aaaiio ?ieue a
i?ioeanao naiii/euaiiy aiaeiei. Cia/ia e?euee?noue aeainooeeiaeo
iiethne?a (aepaenaie, eoeueeiai oa aipioeieiai) ? ia’?eoii aeeaeaiiy ?ea
oa aeayeeo ?ioeo aiaeyieo oaa?ei. Aeainooeeia? iiethnee aenooiathoue a
?ie? oacy?a oe?eiai ?yaeo aiaeia?iio?a (ia?aceo?a oa eiiainae?a), ui
aeei?enoiaothoue iiethne?a ye noano?ao i?ee??ieaiiy, na?aaeiaeua
iaoeaiiy oa aeaea?aei aeaniiai aeeaeaiiy. Aiaeii/an, aeaye? aeaee
aeainooeeiaeo iiethne?a nai? aeei?enoiaothoue aia?a?th ?ioeo i?aai?ci?a,
ia?aceooth/e ia ee/eieiaeo noaae?yo. Ana oea na?ae/eoue i?i oa, ui
neia?ic o aeaeyae? ia?aceoecio, eiiainae?cio oa ?ioeo aeae?a
ni?a?nioaaiiy a?ae?a?a? cia/io ?ieue a aeeoo?aeo oeeeeao aeainooeeiaeo
iiethne?a i??nieo aiaeiei.

Aeey a?eueo iiaiiai ?icoi?iiy ?ie? aeainooeeiaeo iiethne?a oa ?o
aiaeia?iio?a a i??niiaiaeieo aeinenoaiao iaiao?aeia eiiieaenia aea/aiiy
?o ye neia?ioeaiioe/ii? ?aeiino?.

Aea/aiiy neia?ioeaiioe/ieo nenoai aeicaiey? ii?aeiaoe a?ae?ia?ieia?/i?
oa ia?aceoieia?/i? iaoiaee aeine?aeaeaiiy a?ae?ia?iio?a ia
i?aai?ciaiiio, iiioeyoe?eiiio oa a?ioeaiioe/iiio ??aiyo ?c
a?ae?ia?ieia?/ieo iiceoe?e, ?icaeyaeaoe ia?aceo?a oa eiiainae?a
iiethne?a ye ua iaeei o?io?/iee ??aaiue aiaeieo aeinenoai.

Ia nueiaiaei? iinoaea na?eicia i?iaeaia ai?ioueae ?c a?iia?aoeiaeaie o
aiaeiiinoa/aii?, ui iaoiiaeai? ia?inoaiiyi ae?aenaiaie a?ae?ioaoi?/ieo
nii?oae, aiaeicaai??a, eaiae?a, aiaeiaiae?a oa iaa?no?aeueieo
o?oaii?iaiae?a c?iooaaeueieo nenoai. Aea/aiiy neeaaeo oa aeeiai?ee
neia?ioeaiioe/ii? nenoaie “aeainooeeia? iiethnee — aiaeia?iioe”
aeooaeueia c iaeyaeo ia iiaeeeaa canoinoaaiiy io?eiaii? ?ioi?iaoe?? a
?ic?iaoe? a?ieia?/ieo iaoiae?a ai?ioueae c a?iia?aoeiaeaie,
noai?thaaieie ae?aenaiith.

Aeine?aeaeaiiy iaoai?ci?a ooieoe?iioaaiiy neia?ioeaiioe/ieo nenoai
iaiao?aeia oaeiae aeey ?icoi?iiy i?ioean?a iioe?aiiy aiaeia?iio?a a
iiioeyoe?yo oacy?a ? caiia?aaiith aeieeiaiiy ai?ciio?e a?ae?ia?iio?a, ui
iathoue ainiiaea?nueea cia/aiiy.

Eiiieaeniee i?aeo?ae aei aea/aiiy iiethne?a oa ?o aiaeia?iio?a c
neia?ioeaiioe/ieo iiceoe?e ? iiaeaeueoith ?ic?iaeith ?aeae oa iai?yie?a
ia?aceoioeaiieia??, caii/aoeiaaieo I.I. Ia?eaae/ai.

Ca’ycie ?iaioe c iaoeiaeie oaiaie. ?acoeueoaoe aeeiiaieo aeine?aeaeaiue
aoiaeyoue neeaaeiaith /anoeiith a eiiieaeni? ooiaeaiaioaeuei? ?iaioe
?inoeoooo a?ae?ia?ieia?? IAI Oe?a?ie, ui i?iaiaeeeenue a ia??iae c 1995
ii 1998 ??. a iaaeao aeaio ieaiiaeo oai: “?ic?iaeoe aeieia?/i? iniiae
ooieoe?iioaaiiy oa aeaoe e?euee?nio ioe?ieo i?ioeanai, ui
a?aeaoaathoueny o aiaeieiao aeioiiiiai oeio: “??/ea-ii?a” a oiiaao
?iioeuenii-noaa?e?ciaaiiai ?aaeeio” oa “Oai?aoe/i? iniiae
ooieoe?iioaaiiy oa no?eeino? aiaeieo oeaiic?a oa aeinenoai, ?o
a?ieia?/iiai ??ciiiai?ooy a eiioaeoieo ciiao ??ciiai ??aiy”.

Iaoa ?iaioe. Aeaoe caaaeueio oa?aeoa?enoeeo neia?ioeaiioe/i?e nenoai?:
“i??niiaiaei? aeainooeeia? iiethnee (oacy?) — aiaeia?iioe (ia?aceoe oa
eiiainaee)” ia iniia? ye?nieo oa e?euee?nieo iiioeyoe?eieo iieaciee?a oa
caaaeueieo a?ioeaiioe/ieo ca’yce?a i?ae ieie.

Caaaeaiiy.

1. I?iaanoe oeaiioe/iee oa iiioeyoe?eiee aiae?c aeine?aeaeaieo
aeainooeeiaeo iiethne?a.

2. Aeine?aeeoe aeaeiaee neeaae oa aeaoe oeaiioe/io oa?aeoa?enoeeo
iiioeyoe?yi aiaeia?iio?a.

3. Aeyaeoe ca’ycee i?ae oeaiioe/ieie oa iiioeyoe?eieie oa?aeoa?enoeeaie
aeainooeeiaeo iiethne?a oa ?o aiaeia?iio?a.

4. C SYMBOL 146 \f «Times New Roman Cyr» \s 14 ynoaaoe ye oeaiioe/i? oa
iiioeyoe?ei? oa?aeoa?enoeee aeainooeeiaeo iiethne?a aieeaathoue ia
aeaeiaee neeaae oa oeaiioe/i? oa?aeoa?enoeee iiioeyoe?e aiaeia?iio?a.

5. Aeine?aeeoe neeaae oa aeeiai?eo neia?ioeaiico “aeainooeeia? iiethnee
— aiaeia?iioe” o ia?eo?oii? oa aaioin?.

Aeine?aeeoe neeaae oa aeeiai?eo neia?ioeaiico a oiiaao ??/eiai?
aeinenoaie (ae?ei Aeanie) oa aiaeinoiaeua (aa?oiy ae?eyiea Eai?anueeiai
aiaeinoiaeua).

Ia iniia? aeeiiaiiai aiae?co oa aeaieo e?oa?aoo?e aeaoe caaaeueio
oa?aeoa?enoeeo neia?ioeaiioe/ii? nenoaie “aeainooeeia? iiethnee —
aiaeia?iioe”.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. A ?iaio? aia?oa i?e
i?iaaaeaii? a?ae?iia?aceoieia?/ieo aeine?aeaeaiue aoa aeei?enoaiee
eiiieaeniee a?ae?ia?ieia?/iee i?aeo?ae, ui aeaei ciiao ?icaeyaeaoe
aea/aieo ia?aceo?a oa eiiainae?a aeainooeeiaeo iiethne?a ye i?aai?cie,
ye? neeaaeathoue iaeei ?c aeueo o?io?/ieo ??ai?a aiaeiei. Ia iniia?
oaeiai i?aeoiaeo, ?c canoinoaaiiyi a?ae?ia?ieia?/ieo iieaciee?a, aea/ai?
neia?ioe/i? ca’ycee a?ae?iia?aceoa?ieo nenoai “aeainooeeia? iiethnee —
aiaeia?iioe”, aecia/aia ?ieue ? i?noea oeeo nenoai o
no?oeoo?ii-ooieoe?iiaeuei?e i?aai?caoe?? aiaeieo aeinenoai caaaeii.

C’yniaaii aieea oeaiioe/ieo oa iiioeyoe?eieo oa?aeoa?enoee aeainooeeiaeo
iiethne?a ia aeaeiaee neeaae oa oeaiioe/i? oa?aeoa?enoeee iiioeyoe?e
aiaeia?iio?a.

Aeine?aeaeaii neeaae oa naciiio aeeiai?eo neia?ioeaiico o ia?eo?oii? oa
aaioin? a oiiaao ??/eiai? nenoaie oa aiaeinoiaeua i?e ??cieo oeaiioe/ieo
oa?aeoa?enoeeao iiioeyoe?e aeainooeeiaeo iiethne?a.

Aeyaeaii a?aei?iiino? o ia??iaeao ?iciiiaeaiiy iiethne?a D. polymorpha
(Pall.) oa D. bugensis (Andr.).

Iaaaaeaii anaa?/io oa?aeoa?enoeeo aiaeia?iio?a ae?aenai — aaaeeeaiai
eiiiiiaioo ia?inoaiue.

Aeey aiaeiei Oe?a?ie aia?oa a?aei?/aii iayai?noue /a?a’yea Aspidogaster
limacoides (Deising), ye ia?aceoa aeainooeeiaeo iiethne?a.

Aia?oa o a?ae?ia?ieia?/ieo aeine?aeaeaiiyo ?ic?aoiaaii iio?e aia?a??, ui
i?ioiaeeoue /a?ac neia?ioe/io nenoaio “iiethnee Dreissena polymorpha —
?iooci??? Conchophthirus acuminatus (Clap.Lacm.)”.

Aeaii ioe?ieo ci?iai o ia?aceoieia?/i?e neooaoe??, ui a?aeaoeeny ia
aeine?aeaeaieo ae?eyieao e?oi?ae? c /ano noai?aiiy Eai?anueeiai
aiaeinoiaeua, ? iia’ycaieie ?c oeeie ci?iaie oeaiioe/ieie
oa?aeoa?enoeeaie aeainooeeiaeo iiethne?a.

Iaoiaeieia?/ia oe?ii?noue oa i?aeoe/ia cia/aiiy ?iaioe. ?acoeueoaoe
?iaioe iiaeooue aooe aeei?enoai? a aeieia?/ieo i?iaiicao, iia’ycaieo ?c
i?ioeanaie oi?ioaaiiy a?i??ciiiai?ooy aiaeieo aeinenoai ? yeino? aiaee a
ieo, ui a?aeaoaathoueny i?ae aieeaii ca?aaoethaaiiy oa/?? ??/ie oa ci?i
a?ae?ieia?/ieo iieaciee?a aiaeiei.

Io?eiai? aeai? i?i neeaae, aeeiai?eo, no?oeoo?i? iniaeeaino?
neia?ioeaiico “aeainooeeia? iiethnee — aiaeia?iioe” oe?oa ?ice?eaathoue
?ieue oe??? nenoaie a i?ioeanao ooieoe?iioaaiiy i??niiaiaeieo
a?ioeaiic?a.

Iaoa??aee ?iaioe a?aee?eaathoue iia? ia?niaeoeae o ?ic?iaoe? a?ieia?/ieo
iaoiae?a ai?ioueae c ia?inoaiiyie, c aeei?enoaiiyi o?aiaoiae Bucephalus
polymorphus Baer.

A iaoiaee/iiio ieai? caii/aoeiaaii ii?aeiaiiy oa noi?nia aeei?enoaiiy o
eiiieaenieo aeine?aeaeaiiyo ia?aceoieia?/ieo (aenoaineai?noue oa
?ioaineai?noue ?iaac??) oa a?ae?ia?ieia?/ieo (/enaeuei?noue, a?iiana)
iieaciee?a.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. ?acoeueoaoe aeena?oaoe?eii?
?iaioe aoee ii?eethaeiai? ia oaeeo i?aeia?iaeieo c’?caeao oa ia?aaeao:
Iaaeaeoia?iaeiia niaauaiea «I?iaeaiu aeae?iaeieiaee eiioeiaioaeueiuo
aiae e eo iaeaeioaoia» (Naieo-Iaoa?ao?a, ?inney, 18 — 21 iiya?y 1996
a.), ?? c’?cae A?ae?iaeieia?/iiai oiaa?enoaa Oe?a?ie, (Ee?a, 27 — 31
aeiaoiy 1997 ?.), VII European Multicolloquium of Parasitology (2 — 6
September 1996, Parma, Italy), IXth International Congress of
Parasitology (24 — 28 August 1998, Japan) .

Ioae?eaoe??. Ii iaoa??aeai aeena?oaoe?? iioae?eiaaii 8 ?ia?o: 3 noaoo? a
iaoeiaeo aeo?iaeao, 1 noaooy a ca??ieeo iaoeiaeo i?aoeue, 4 ?iaioe a
iaoa??aeao i?aeia?iaeieo iaoeiaeo c’?cae?a oa eiioa?aioe?e.

Ia’?i oa no?oeoo?a ?iaioe. Aeena?oaoe?y aeeeaaeaia ia 128 noi??ieao
oaenoo ? neeaaea?oueny c anooio, i’yoe ?icae?e?a oa aeniiae?a. Oaeno
?ethno?iaaii 40 ?enoieaie oa 19 oaaeeoeyie. Nienie oeeoiaaieo
e?oa?aoo?ieo aeaea?ae i?noeoue 124 iaca, c ieo 24 ia ?iicaii?e iia?.

INIIAIEE CI?NO ?IAIOE

ANOOI

O anooi? aecia/ai? aeooaeuei?noue oaie, iaoa oa caaaeaiiy, iaoeiaa
iiaecia, i?aeoe/ia oa iaoiaeieia?/ia oe?ii?noue ?iaioe.

IAOA??AEE OA IAOIAeE

Iaoa??aee aeey ?iaioe aoee c?a?ai? ia i?ioyc? 1996-1997 ?ie?a ia
e?oi?aeueieo ae?eyieao ae?ea Aeanie oa aa?oiuei? /anoeie Eai?anueeiai
aiaeinoiaeua. Ia aeacaieo aiaeieo ia’?eoao noi?ioaaeeny ?
ooieoe?iiothoue eioe/i? a?ioeaiice, ui iaeaea ia caciaee (ae?ei Aeanie),
aai /anoeiai caciaee (aa?o?a’y Eai?anueeiai aiaeinoiaeua) aieeao
ca?aaoethaaiiy oa/?? Aei?i?a, ye? oa?aeoa?ecothoueny aenieeie
iieacieeaie /enaeueiino? oa a?iiane aeainooeeiaeo iiethne?a (na?aaei?
iieacieee /enaeueiino?: ia?e?aieoeaa? aei 120 aec./i2, eoeueeia? — aei
300 aec./i2, ae?aenaie — aei 10 oen. aec./i2). ??ciiiai?oi?noue oiia
?nioaaiiy, e?euee?nia ? ye?nia i?aaenoaaeai?noue aeainooeeiaeo iiethne?a
ia aea?aieo ae?eyieao aanaeio Aei?i?a aeicaieeee aecia/eoe ?o a iaai?e
i??? iiaeaeueieie ? aeine?aeaeoaaoe ?nioth/? iiioeyoe?? aeii?ioth/eo o
aaioin? oa ia?eo?oii? aeae?a iiethne?a, ui iioe?ai? a aanaei? Aei?i?a, c
iiaeaeueoith ?ioa?i?aoaoe??th io?eiaieo aeaieo ia ?io? ae?eyiee
aei?i?ianueeeo aiaeinoiaeu oa ??/ie, ui aei ieo aiaaeathoue. Oea oae?
aeaee, ye Unio tumidus falcatulus Droet, U. pictorum ponderosus Spitzi
in Rossmaessler, Anodonta cygnea (L.), Colletopterum piscinale falcatum
(Droet), Rivicoliana rivicola (Lamark), Pisidium amnicum (Mull.),
Dreissena polymorpha (Pall.) , D. bugensis (Andr.).

E?oi?aeuei? a?ioeaiice aaioino ae?ea Aeanie oa aa?oiuei? ae?eyiee
Eai?anueeiai aiaeinoiaeua a?ae??ciyeeny ii iieacieeao aaaaonoaa oa
aeii?ioaaiiy iaaieo aeae?a aeainooeeiaeo iiethne?a. Aeoiaey/e c oeueiai,
i?e cai?? iaoa??aeo aoa canoiniaaiee iaoiae o?ainaeo. Caaaeii aoei
a?ae?a?aii aeecueei 500 aaioinieo oa 100 ia?eo?oiiieo i?ia.

I?e aeine?aeaeaii? iiethne?a aecia/aee ?o nenoaiaoe/ia iieiaeaiiy,
iieacieee /enaeueiino? oa a?iiane, ?ici??ii-a?eiao oa noaoaao no?oeoo?e
iiioeyoe?e. A i?ioean? iiaeaeueoiai aiae?co o noaoaa?e no?oeoo??
iiioeyoe?e aeae?eyee o?e a?oie: thaai?euei? iiethnee, noaoaaic??e? oa
noa??th/? iiethnee. O ?ici??ii-a?eia?e no?oeoo?? oe? a?oie a?aeiia?aeaee
iiethneai c iaaiith aeiaaeeiith /a?aiaoee. Aeai? uiaei ?ici???a
thaai?eueieo oa noaoaaic??eeo iniaei io?eiai? c aeanieo niinoa?aaeaiue
oa e?oa?aoo?ieo aeaea?ae. Noa??th/eie aaaaeaeenue iiethnee aeiaaeeia
/a?aiaoee yeeo iiiaaeaea o ?ici??iee ?ioa?aae:

[L + 2/3(Lmax-L)…Lmax],

aea L — aeiaaeeia noaoaaic??eeo iiethne?a;

Lmax — iaeneiaeueia aeiaaeeia /a?aiaoie aeine?aeaeaieo aeae?a
iiethne?a.

Aeey ?aeaioeo?eaoe?? oa e?euee?niiai iae?eo aiaeia?iio?a an? a?ae?a?ai?
iiethnee, aai /anoeia aea??ee i?aeaeaaaeenue iiaiiio ia?aceoieia?/iiio
?icoeio. Oaeeo ?icoei?a aoei aeeiiaii aeecueei 3500.

E?euee?noue aiaeia?iio?a aecia/aeanue oeyoii ?o i?ae?aooieo i?ae /an
ia?aceoieia?/iiai ?icoeio iiethne?a. Iano aiaeia?iio?a cia/ieo ?ici???a
aecia/aee c oi/i?noth aei 0,01 ia.

A?ioeaiioe/i? ca’ycee o neia?ioeaiioe/i?e nenoai? eeaneo?eoaaee,
ea?oth/enue nenoaiith, cai?iiiiiaaiith A.I. A?ee?i?oaaei (1951).

Noaoenoe/ia ia?iaea aeaieo aeeiiaia c aeei?enoaiiyi i?ia?ai STATGRAFIC
oa STATISTICA for WINDOWS.

?IEUe AeAINOOEEIAEO IIETHNE?A OA ?O AIAeIA?IIO?A A I??NIIAIAeIEO

AEINENOAIAO

O ?icae?e?, ea?oth/enue aeaieie e?oa?aoo?ieo aeaea?ae oa iaoa??aeaie
aeanieo aeine?aeaeaiue, aena?oeaii ?ieue aeainooeeiaeo iiethne?a oa ?o
aiaeia?iio?a a i??niiaiaeieo aeinenoaiao. Iieacaii, ye? ci?ie o noai?
iiioeyoe?e aeainooeeiaeo iiethne?a oa neeaaeiaeo ?o neia?ioeaiico
noaeeny o ca’yceo ?c c?inoaiiyi aio?iiiaaiiiai oeneo ia i?e?iaei?
a?ae?iaeinenoaie oa ca?aaoethaaiiyi oa/?? ??/ie. Aeacaii ia ia?niaeoeai?
iai?yiee aeey ?ic?iaee a?ieia?/ieo iaoiae?a ai?ioueae ?c ia?inoaiiyie
ae?aenai, ui noai?ththoue a?iia?aoeiaee o aiaeiiinoa/aii?. Aeaii
aeaoaeueiee iaeyae aiaeia?iio?a i??niiaiaeieo aeainooeeiaeo iiethne?a
Oe?a?ie, i?eaaaeai? aeai? ?o iaoiaaiiino? aeey oacy?a.

OeAIIOE*IEE OA IIIOEssOe?EIEE AIAE?C AeINE?AeAEAIEO AeAINOOEEIAEO

IIETHNE?A

Aeainooeeia? iiethnee o ae?e? Aeanie aoee i?aaenoaaeai? 9 aeaeaie.
Aeii?ioth/eie ii /enaeueiino? (N) oa a?iian? (B) aoee Unio tumidus (N-60
aec./i2, B-2700 a/i2), U. pictorum (N-60 aec./i2, B-2900 a/i2),
Rivicoliana rivicola (N-80 aec./i2, B-84 a/i2).

Ia e?oi?ae? aa?oiuei? ae?eyiee Eai?anueeiai aiaeinoiaeua aoei ciaeaeaii
8 aeae?a aeainooeeiaeo iiethne?a. O aaioin? ii /enaeueiino? oa a?iian?
aeii?ioaaee Colletopterum piscinale (N -15 aec./i2, B — 500 a/i2),
Anodonta cygnea (N -10 aec./i2, B — 430 a/i2), Unio pictorum (N — 10
aec./i2, B — 500 a/i2), oa Dreissena polymorpha c ia?inoaiue /a?aiaoie
ia?e?aieoeaaeo (N — 2-3 oen. aec./i2, B -1200-2000 a/i2). O ia?eo?oii?
ii /enaeueiino? oa a?iian? aeii?ioaaea D. bugensis (N — 8-10 oen.
aec./i2, B-6100 a/i2).

Aei neeaaeo aeine?aeaeoaaiiai neia?ioeaiico “aeainooeeia? iiethnee —
aiaeia?iioe” aoiaeyoue aeaeia? iiioeyoe?? iiethne?a c ??ciith
?ici??ii-a?eiaith no?oeoo?ith. Iiioeyoe?? Unio tumidus, Dreissena
polymorpha, D. bugensis, Rivicoliana rivicola o nai?io neeaae? i?noyoue
aanue niaeo? ?ici??ii-a?eiaeo a?oi, noaoaaa no?oeoo?a i?aaenoaaeaia
thaai?eueieie oa noaoaaic??eeie iniaeiaie ia anueiio i?ioyc?
aaaaoaoe?eiiai ia??iaeo.

A iiioeyoe?yo U. pictorum, Anodonta cygnea, Colletopterum piscinale,
Pisidium amnicum iieiaeo? ?ici??ii-a?eia? a?oie niinoa??aaeenue ia
e?oi?ae? eeoa a iaai? ia??iaee, a ?ici??ii-a?eiaa oa noaoaaa no?oeoo?e
aoee i?aaenoaaeai? ia?aaaaeii noa?oeie ?ici??ii-a?eiaeie a?oiaie oa
noaoaaic??eeie iniaeiaie.

Ia a?aei?io a?ae ae?ea Aeanie ia e?oi?ae? aa?oiuei? ae?eyiee
Eai?anueeiai aiaeinoiaeua ia aoea a?aei?/aia iiyaa oeueiai??/ii? iieiae?
ia?e?aieoeaaeo.

O noaoaa?e no?oeoo?? ?icae?eueiinoaoaaeo aeae?a (ia?e?aieoeaa? oa
ae?aenaie) ia?aaaaeaee naioe? (55-80% /enaeueiino? noaoaaic??eeo
iniaei).

CAAAEUeIA OA?AEOA?ENOEEA IIIOEssOe?E AIAeIA?IIO?A NEIA?IOeAIIOE*II?
NENOAIE.

An?o aeyaeaieo aiaeia?iio?a, aeoiaey/e c ?icaeoeo o aiaeieo aeinenoaiao,
iiaeia ?icae?eeoe ia aea? a?oie:

1. Aiaeia?iioe c i?yiei oeeeeii ?icaeoeo;

2. Aiaeia?iioe c iai?yiei oeeeeii ?icaeoeo.

Aei ia?oi? a?oie iaeaaeaoue i?aai?cie, an? aeeoo?a? noaae?? yeeo iia
SYMBOL 146 \f «Times New Roman Cyr» \s 14 ycai? c iiioeyoe??th iaeiiai
aeaeo oacy?ia, oiaoi, o iaoiio aeiaaeeo — eeoa c iiioeyoe?yie
aeainooeeiaeo iiethne?a. Na?aae cono??iooeo aeae?a aiaeia?iio?a aei
oe??? a?oie iaeaaeaee ?iooci??? oa ani?aeiaano?e.

Aei ae?oai? a?oie iaeaaeaoue i?aai?cie, ui aeei?enoiaothoue ia iaaieo
noaae?yo aeeoo?aiai oeeeeo iiioeyoe?? ??cieo oacy?a. Aeainooeeia?
iiethnee o oeueiio aeiaaeeo aenooiathoue o ?ie? i?ii?aeieo oacy?a. Aei
oe??? a?oie iaeaaeaoue ia?oai?oe oa ee/eiee o?aiaoiae, a oaeiae i?ioe
ee?u?a.

4.1. Oa?aeoa?enoeea aiaeia?iio?a c i?yiei oeeeeii ?icaeoeo.

?iooci???. Iiioeyoe?eiee aiae?c a?ioeaiioe/ieo ca SYMBOL 146 \f «Times
New Roman Cyr» \s 14 yce?a o nenoai? “aeainooeeia? iiethnee —
aiaeia?iioe” aeyaea 9 aeae?a ?iooci??e. An? aeaee iae?aaoii iia SYMBOL
146 \f «Times New Roman Cyr» \s 14 ycai? c aeainooeeiaeie iiethneaie ?
?ai?oa ?a?no?oaaeenue o aanaei? Aei?i?a oa ?ioeo ??/ie Oe?a?ie
(Eaaioeea, 1979; Noaaeie/aiei, 1984). A ia?e?aieoeaaeo aeyaeaii 8
aeae?a, a ae?aenai — 2 aeaee, a eoeueeiaeo — 1 aeae ?iooci??e.

Iaea?eueoa aenoaineai?noue ?iaac?? ?iooci??yie aeanoeaa Anodonta cygnea,
C.piscinale, D. polymorpha (55-100% /enaeueiino?). Cieaeaiiy
aenoaineaiino? ?iaac?? iiethne?a oeeo aeae?a (Anodonta cygnea, C.
piscinale — a eeii?, D. polymorpha — o na?ii?) iia’ycaia ?c iiyaith o
iiioeyoe?? oeueiai??/ii? iieiae? oa ca?eueoaiiyi aeie? thaai?eueieo
iniaei, ui a iaio?e i??? ?iaaciaai? ?iooci??yie (aenoaineai?noue ?iaac??
— 0-14%). Aenoaineai?noue ?iaac?? iiethne?a Unio tumidus, Dreissena
bugensis eieeaaeanue a iaaeao a?ae 0 aei 70% (20-30 % a na?aaeiueiio).
Iaeneioi aenoaineaiino? ?iaac?? oeeo iiethne?a ?iooci??yie i?eiaaea? ia
/a?aaiue i?nyoeue, eiee o ?ici??ii-a?eia?e no?oeoo?? iiioeyoe?e
ia?aaaaeathoue noa?o? a?oie, a iieiaeue noaiiaeoue eeoa 5-10% caaaeueii?
/enaeueiino?.

Iaea?eueoiai ?icaeoeo ?iooci??? aeinyaathoue a iiioeyoe?yo Unio pictorum
(aaioin ae?ea Aeanie) oa D. polymorpha (aaioin oa ia?eo?oii aa?oiuei?
ae?eyiee Eai?anueeiai aiaeinoiaeua), ui aeii?iothoue ii /enaeueiino? oa
a?iian?. Na?aaeiy /enaeuei?noue ?iooci??e a iiioeyoe?yo oeeo aeae?a
neeaaeaea 30-40 oen. aec./i2 a?ioiio iaoeaiiy oacy?a. Iaeneiaeueia
/enaeuei?noue ?iooci??e (U. pictorum — 83 oen. aec./i2, D. polymorpha —
61 oen. aec./i2) niinoa??aaeanue a eeii? i?nyoe?, eiee ?ici??ii-a?eiaa
no?oeoo?a iiioeyoe?? U. pictorum ia 100%, a D. polymorpha — ia 55% aoea
i?aaenoaaeaia noa?oeie a?eiaeie a?oiaie, a aenoaineai?noue ?iaac??
neeaaeaea 94% — aeey U. pictorum oa 92% — aeey D. polymorpha.
*enaeuei?noue ?iooci??e a iiioeyoe?yo ?ioeo aeae?a eieeaaeanue a iaaeao
a?ae 100 aei 22 000 aec./i2 noano?aoo iaoeaiiy oacy?ia. Aeey iiethne?a
Unio tumidus oa D. bugensis oa?aeoa?ia iecueea aenoaineai?noue ?iaac??
?iooci??yie, a iiethnee Anodonta cygnea oa C. piscinale iaee iecueeo
(ii??aiyii c iiethneaie ?iaeo Unio) u?euei?noue iiioeyoe?e ia
aeine?aeaeaieo ae?eyieao e?oi?ae? (10 -15 aec./i2).

Ani?aeiaano?e. ?ioith a?oiith aiaeia?iio?a c i?yiei oeeeeii ?icaeoeo ?
ani?aeiaano?e — i?aaenoaaieee eeano Aspidogastrea, oeio Plathelmintes —
ieane? /a?ae. Iaie aoei aeyaeaii aeaa aeaee oeeo i?aai?ci?a —
i?aaenoaaiee?a iaeiiai ?iaeo: Aspidogaster conchicola Baer — ia?aceo
ia?e?aieoeaaeo, A. limacoides — ia?aceo ae?aenai. *a?ae iaio aeae?a
eieae?cothoueny a ia?eea?ae?aeuei?e ii?iaeiei? oa ie?eao iiethne?a.
Iaea?eueoa aenoaineai?noue ?iaac?? oa?aeoa?ia aeey iiethne?a ?iaeo Unio
— 10-47%. Aeey Unio tumidus aeanoeaee aeuee iieaciee ?ioaineaiino?
?iaac?? oeei ia?aceoii ii??aiyii c Unio pictorum (na?aaeiy
?ioaineai?noue ?iaac?? U. tumidus — 1,8 aec./iniaeio, U. pictorum — 1,2
aec./iniaeio).

AEaao?ieoe? (A. cygnea oa C. piscinale) oa?aeoa?ecothoueny iecueeith
aenoaineai?noth (2-6%) oa aenieith ?ioaineai?noth ?iaac?? oeei ia?aceoii
(2,9-6,1 aec./iniaeio). Iaeneiaeueia ?ioaineai?noue ?iaac?? cao?eniaaia
aeey iiethnea A. cygnea ? neeaea 13 aec./iniaeio.

Ia e?oi?ae? ae?ea Aeanie oa aa?o?a’?a Eai?anueeiai aiaeinoiaeua ?c
iiethneaie ?iaeo Unio iia’ycaia iniiaia /anoeia iiioeyoe?? Aspidogaster
conchicola. Iieacieee a?iiane oeueiai ia?aceoa aoee iaeneiaeueieie a
/a?ai?, aeey iiioeyoe?? U. tumidus c ae?ea Aeanie aiie neeaaeaee 440
ia/i2 noano?aoo iaoeaiiy oacy?ia, a aeey iiioeyoe?? U. pictorum — 190
ia/i2. O iiethne?a c aa?oiuei? ae?eyiee Eai?anueeiai aiaeinoiaeua
a?iiana oeueiai ia?aceoa eieeaaeanue a iaaeao 5-10 ia/i2. A oeae /an
iiioeyoe?? iiethne?a oeeo aeae?a ia 95-100% aoee i?aaenoaaeai? noa?oeie
?ici??ii-a?eiaeie a?oiaie. Oae? aenie? cia/aiiy aenoaineaiino? ?iaac??
iiethne?a oa /enaeueiino? (a?iiane) /a?a?a A. conchicola aeey aiaeiei
Oe?a?ie cao?eniaai? aia?oa. Ca aeaieie ?ioeo aeine?aeiee?a
aenoaineai?noue ?iaac?? oeei ia?aceoii eieeaaeanue a iaaeao 0,5-5%
(Eaaioeea, 1979; Noaaeie/aiei, 1984), i?e oeueiio a?iiana ca iaoeie
i?ae?aooieaie ia ia?aaeuoaaea 50 ia/i2.

Ia?aceo A. limacoides aoa cono??iooee eeoa a na?ii? 1996?. a iiioeyoe?yo
iaio aeae?a ae?aenai o aaioin? oa ia?eo?oii? aa?oiuei? ae?eyiee
Eai?anueeiai aiaeinoiaeua (aenoaineai?noue ?iaac?? iiethne?a D.
polymorpha — 0,7%, D. bugensis — 0,8%, ?ioaineai?noue — 1 aec./iniaeio).
Aiane?aeie aeniei? u?eueiino? iiioeyoe?e ae?aenai a?iiana oeueiai
ia?aceoa nyaaea 80 ia/i2 noano?aoo iaoeaiiy oacy?a.

Ia?aceo A. limacoides ca iaoeie aeaieie aia?oa a?aei?/aiee a Oe?a?i? ye
ia?aceo aeainooeeiaeo iiethne?a. ?ai?oa oeae aeae iino?eii ?a?no?oaaany
o neeaae? ia?aceoioaoie ?ea Aei?i?a oa aiaeinoiaeu ?c cia/iith
aenoaineai?noth ?iaac?? (ycue, aonoa?a — 60%) (Oeoa?, 1989).
Aeainooeeia? iiethnee aaaaeathoueny ia?aeiieie oacyyie oeueiai ia?aceoa
(Oeiioaaaa, 1971).

4.2. Oa?aeoa?enoeea aiaeia?iio?a c iai?yiei oeeeeii ?icaeoeo.

Aeey i?aai?ci?a c iai?yiei oeeeeii ?icaeoeo aeainooeeia? iiethnee
aenooiathoue o ?ie? ia?oiai aai ae?oaiai i?ii?aeiiai oacy?ia. sse
ia?oiai i?ii?aeiiai oacy?ia, aeainooeeiaeo iiethne?a aeei?enoiaothoue
ia?oai?oe o?aiaoiae oa i?ioe ee?u?a, ye ae?oaiai — iaoaoea?ea???
o?aiaoiae.

Ia?oai?oe oa oea?ea??? o?aiaoiae. O aeine?aeaeai?e iaie
neia?ioeaiioe/i?e nenoai? aoei aeyaeaii 7 aeae?a o?aiaoiae ia noaae?yo
nii?ioeeno, ?aae?e, oea?ea??e. O ae?e? Aeanie aoei aeyaeaii 6 aeae?a
o?aiaoiae, o aa?o?a’yo Eai?anueeiai aiaeinoiaeua — 3 aeaee (1 aeae o
iiethne?a aaioino, 2 aeaee o iiethne?a ia?eo?oiio). A iaea?eueo?e i???
?iaaciaaieie aeyaeeenue iiethnee Pisidium amnicum (4 aeaee o?aiaoiae ?c
caaaeueiith aenoaineai?noth ?iaac?? — 80-90%). Aenoaineai?noue ?iaac??
iiioeyoe?e iiethne?a ?ioeo aeae?a ia ia?aaeuoaaea 10%. Iaeaeua
aenoaineai?noue ?iaac?? oa?aeoa?ia aeey o?aiaoiae Bunodera luciopercae
(Mull.), Cercaria pardoxa ecaudata Bidulina oa Gorgoderina vitelliloba
(Ssinitzin.), ui ia?aceoothoue o iiethne?a P. amnicum. O iaea?eueo?e
i??? ?iaaciaai?noue o?aiaoiaeith Bunodera luciopercae niinoa??aaeanue a
/a?ai? i?nyoe?, eiee o ?ici??ii-a?eia?e no?oeoo?? iiioeyoe?? iiethne?a
P. amnicum ia?aaaaeaee iieiaeo? a?oie, ?iaaciaai? ia 37%,
aenoaineai?noue ?iaac?? noa?oeo ?ici??ii-a?eiaeo a?oi oe??th
o?aiaoiaeith neeaaeaea 90%. Iaeneioi ?icaeoeo o?aiaoiaee C. pardoxa
ecaudata i?eiaaeaa ia na?iaiue i?nyoeue, eiee o ?ici??ii-a?eia?e
no?oeoo?? aoee i?aaenoaaeai? eeoa noa?o? a?oie, i?e oeueiio noa??th/?
iiethnee aoee ?iaaciaai? ia 59%, a ?io? noaoaaic??e? iiethnee — ia 37%.

Aenoaineai?noue ?iaac?? iiethne?a o?aiaoiaeaie ?ioeo aeae?a ia
ia?aaeuoaaea 10%. I?e oeueiio o?aiaoiaea Allocreadium isoporum (Loos)
aoea ciaeaeaia eeoa o iiethne?a Rivicoliana rivicola, e?i?ei? ?ici??e
yeeo aoee a?eueoa 15 ii, a o?aiaoiaea Bucephalus polymorphus — eeoa o
noaoaaic??eeo iiethne?a ?ici??ii a?eueoa i?ae 60 ii aeiaaeeie
(ia?e?aieoeaa?) oa 15 ii aeiaaeeie (ae?aenaie). Ia?aceooaaiiy o?aiaoiae
Bucephalus polymorphus a aiiaae? iiethne?a ni?e/eiy? iiaio aai /anoeiao
noa?ee?caoe?th oacy?ia. Iaaaoeaiee aieea aooeaoae?ae ia ?ai?iaeoeoeaio
caeaoi?noue iiethne?a iiaea aooe aeei?enoaiee aeey a?ieia?/ii? ai?ioueae
c ia?inoaiiyie ae?aenai (Caeoi, 1969; Oaaoeiaa, Oa?/aiei, 1986). Iaeiae,
canoinoaaiiy a?ieia?/ieo iaoiae?a, iaoe?eaieo ia cieaeaiiy a?iia?aoeiae
o aiaeiiinoa/aii?, ye? aacothoueny ia aeei?enoaii? ia?aceo?a ae?aenai,
iio?aao? aeineiiaeueii? iaoeiai-aeine?aeieoeueei? ia?aa??ee, ai
iia’ycaia ?c ci?iaie a a?ioeaiicao, ye? aaaeei ia?aaeaa/eoe ?
i?iaiicoaaoe.

Ee?u?. I?e aeine?aeaeai? neia?ioeaiico “aeainooeeia? iiethnee —
aiaeia?iioe” iaie aoei aeyaeaii aeaa aeaee ee?u?a, ui iaeaaeaoue ?iaeo
Unionicola. Ee?u Unionicola ypsilophora (Bonz) aoa ciaeaeaiee o
iiethne?a Anodonta cygnea oa Colletopterum piscinale ia e?oi?ae? ae?ea
Aeanie oa aa?oiuei? ae?eyiee Eai?anueeiai aiaeinoiaeua (aenoaineai?noue
?iaac?? — 0,9-2%, ?ioaineai?noue — 2-3 aec./iniaeio). Ee?u Unionicola
sp. aoa ciaeaeaiee o iiethne?a Dreissena bugensis (aenoaineai?noue
?iaac?? — 0,9-2%, ?ioaineai?noue — 1 aec./iniaeio) o ia?eo?oii?
aiaeinoiaeua. I?io oa aei?ineeo ee?u?a ?a?no?oaaee ia anueiio i?ioyc?
aaaaoaoe?eiiai ia??iaeo.

Iaoaoea?ea??? o?aiaoiae. Ia noaae?? iaoaoea?ea??? aoei aeyaeaii o?e
aeaee o?aiaoiae. Aenoaineai?noue oa ?ioaineai?noue ?iaac?? iiethne?a
oeenoaie o?aiaoiae ?iaeeie Echinostomatidae aoee iaeaeueie
(aenoaineai?noue ?iaac?? aei 70%, ?ioaineai?noue — aei 60 aec./iniaeio)
a ii??aiyii? c ?ioeie aeaeaie (aenoaineai?noue ?iaac?? 3-5%,
?ioaineai?noue — 1-4 aec./iniaeio). Na?aae oacy?a ?iaeeie
Echinostomatidae a?aei?/ai? Dreissena polymorpha, D. bugensis,
Rivicoliana rivicola, Amesoda solida.

A?eueo aeaoaeueii aca?iiae?th i?ae iiioeyoe?yie aeainooeeiaeo iiethne?a
oa oeenoaie o?aiaoiae ?iaeeie Echinostomatidae ii?aoeueiaaii ia
i?eeeaae? iiioeyoe?e D. polymorpha oa D. bugensis. Iaeneiaeueiee
?icaeoie oeeo aiaeia?iio?a a iiioeyoe?yo iaio aeae?a niinoa??aaany o
e?oi? i?nyoe? (na?aaeiy aenoaineai?noue ?iaac?? D. bugensis — 58%, D.
polymorpha — 45%; ?ioaineai?noue ?iaac?? 5,2 oa 3,1 aec./iniaeio,
a?aeiia?aeii). Iaeaeua aenoaineai?noue ?iaac?? iiethne?a oeeie
o?aiaoiaeaie niinoa??aaeanue a eeii? i?nyoe?, eiee o ?ici??ii-a?eia?e
no?oeoo?? iiioeyoe?e ia?aaaaeaee iiethnee noa?oeo a?oi (70-95%
/enaeueiino?). C eeiiy ii aa?anaiue /enaeuei?noue oeeno Echinostomatidae
a iiioeyoe?? D. bugensis ciaoiaeeeanue o iaaeao 3900-5900 aec./i2
noano?aoo iaoeaiiy oacy?a. O iiioeyoe?? D. polymorpha /enaeuei?noue oeeo
aiaeia?iio?a o aeacaiee ia??iae ciaoiaeeeanue o iaaeao 100-1900 aec./i2
noano?aoo.

A?eueo aenie? iieacieee /enaeueiino? oeeno o?aiaoiae Echinostomatidae o
iiioeyoe?? D.bugensis o e?oi?e ia??iae iiaeia iiynieoe a?eueoith
(ii??aiyii c D. polymorpha) /enaeuei?noth oeeo iiethne?a o ia?eo?oii?,
oa a?aenooi?noth o oeae ia??iae o ?ici??iiio neeaae? iiioeyoe??
iaei?iaaciaaii? iieiae?.

CAAAEUeIA OA?AEOA?ENOEEA NEIA?IOeAIIOE*II? NENOAIE “AeAINOOEEIA?
IIETHNEE — AIAeIA?IIOE”

5.1. Neio?c?ieia?/i? ca’ycee o nenoai? “aeainooeeia? iiethnee —
aiaeia?iioe”.

Iiioeyoe?? a?eueoino? aeyaeaieo iaie aeae?a aiaeia?iio?a ooai?ththoue
aeiaaio?eaae? a?ioeaiioe/i? ca’ycee ?c iiioeyoe?yie aeainooeeiaeo
iiethne?a. Aeey an?o aiaeia?iio?a iiethnee ? eiiaeeoe?iioth/eie
na?aaeiaeua i?aai?ciaie oa iin?yie aeaea?aea aeeaeaiiy. An?o
aiaeia?iio?a ?c iiethneaie iia’ycothoue i?yi? oii?/i?, o?io?/i? oa
oi?e/i? ca’ycee (?en.1). Oa?aeoa? o?io?/ieo ca’yce?a a?aei?iiee aeey
aiaeia?iio?a, ui aaaaeathoueny ia?aceoaie aai eiiainaeaie.

Ie??i aaciina?aaeiueiai o?io?/iiai ca’yceo ?c iiethneaie, eiiainaeuei?
i?aai?cie (?iooci???) iathoue i?yiee o?io?/iee ca’ycie ?c aaeoa??yie, ui
eiioeaio?othoueny o iaio?ei?e ii?iaeiei? aiane?aeie o?eueo?aoe??
iiethne?a, ?ioaineaii ?icaeaath/enue ia aaethoeiaoao. Naia aaeoa??? ?
iniiaiei aeaea?aeii aeeaeaiiy eiiainaeueieo ?iooci??e. ?iooci??? oaeiae
iia’ycai? ?c aeainooeeiaeie iiethneaie iiina?aaeeiaaiei o?io?/iei
ca’yceii, ai oaeaee?noue o?eueo?aoe?? oa ?ioaineai?noue aeeaeaiiy
aeainooeeiaeo iiethne?a aieeaathoue ia /enaeuei?noue aaeoa??e o
iaio?ei?e ii?iaeiei?.

Iiioeyoe?? aiaeia?iio?a, ye neeaaeia? neia?ioeaiico, anooiathoue i?ae
niaith a aioaaii?noe/i? oa iaeo?aeuei? a?aeiineie. E?oa?aoo?i? aeai?
na?ae/aoue i?i ?nioaaiiy iiaeeeaiai aioaaii?cio i?ae o?aiaoiaeaie
Bucephalus polymorphus oa ani?aeiaano?aie Aspidogaster conchicola, ui
i?aeoaa?aeaeo?oueny ? iaoeie aeaieie. O aeine?aeaeai?e neia?ioeaiioe/i?e
nenoai? niinoa??aaeenue oaeoe noi?nii? ?iaac?? iaeiiai aeaeo oacy?ia
(Pisidium amnicum) aeae?eueeiia aeaeaie o?aiaoiae i?e iaaenie?e
?ioaineaiino? ?iaac?? (1-2 ?aae??/inia.). Iiae?aia yaeua iienaia aeey
o?aiaoiae, ui ia?aceoothoue o /a?aaiiiaeo (Aeia?iaieueneee, Aaeaeoeiiia,
1983). Aca?iia?aeiineie i?ae o?aiaoiaeaie caeeoathoueny iaeo?aeueieie
aei oiai /ano, iiee i?ae ieie ia aeieea? eiieo?aioe?? ca o?io?/i?
?ano?ne.

sse iaeo?aeuei? iiaeia oa?aeoa?ecoaaoe aca?iia?aeiineie i?ae ??cieie
aeaeaie ?iooci??e c iaio?eii? ii?iaeieie aeainooeeiaeo iiethne?a. A 90%
aeiaaee?a ?iooci??? cono??/aeenue o aeaeyae? ??ciiaeaeiaeo oa?oiiaaiue.
O e?oa?aoo?? a?aei?/ai? oaeoe ii?aeaiiy ?iooci??e Conchophthirus sp.
?iooci???th Trichodina unionis Hampl. ?iooci??? Trichodina unionis
aeyaeyeenue iaie eeoa o noi?nieo oa?oiiaaiiyo c ?iooci??yie ?iaeo
Conchophthirus.

5.2. ?ieue ??cieo ?ici??ii-a?eiaeo oa noaoaaeo a?oi iiioeyoe?e iiethne?a
o oi?ioaaii? a?ioeaiioe/ieo ca’yce?a o neia?ioeaiioe/i?e nenoai?.

Iiooaei?noue a?ioeaiioe/ieo ca’yce?a o neia?ioeaiioe/i?e nenoai? iiaea
oa?aeoa?ecoaaoe iieaciee aenoaineaiino? ?iaac?? aeainooeeiaeo iiethne?a
(oiaoi /anoea iiioeyoe??, ui aca?iiae?? c aiaeia?iioaie).

Aeey on?o aeine?aeaeaieo aeae?a aeainooeeiaeo iiethne?a niinoa??aaeinue
c?inoaiiy aenoaineaiino? ?iaac?? o noa?oeo ?ici??ii-a?eiaeo a?oiao.
Ci?ie o ?ici??ii-a?eia?e no?oeoo?? iiioeyoe?e iaieio/a aaaeooue aei ci?i
o caaaeuei?e aenoaineaiino? ?iaac?? iiioeyoe??. A?eueo aeaoaeueii oe?
ci?ie aeine?aeaeaii ia iiioeyoe?yo ae?aenai iaio aeae?a.

I?enooi?noue a neeaae? oeaiic?a naia noa?oeo ?ici??ii-a?eiaeo a?oi
aecia/a? aenieo aenoaineai?noue ?iaac?? iiioeyoe??. Iiyaa iieiae? o
iiioeyoe?? oacy?ia aaaea aei ciaioaiiy oeueiai iieacieea.

Iaie ia aeyaeaii noaoenoe/ii-aeinoia??ii? a?aei?iiino? i?ae
aenoaineai?noth oa ?ioaineai?noth ?iaac?? iiethne?a ??cii? noao? (o
ia?e?aieoeaaeo oa ae?aenai). Iaeiae, aeey an?o aea/aieo aeae?a iiethne?a
aeyaeaii noaoenoe/ii-aeinoia??i? a?aei?iiino? a ?iaaciaaiino?
thaai?eueieo iniaei ii??aiyii ?c noaoaaic??eeie.

5.3. Aieea aeaeiaiai neeaaeo oa /enaeueiino? oa?oiiaaiue aeainooeeiaeo
iiethne?a ia oi?ioaaiiy a?ioeaiioe/ieo ca’yce?a o nenoai? «aeainooeeia?
iiethnee — aiaeia?iioe».

Ia oi?ioaaiiy a?ioeaiioe/ieo ca’yce?a o nenoai? «aeainooeeia? iiethnee —
aiaeia?iioe» aieeaathoue ia eeoa iieacieee no?oeoo?e iiioeyoe?e
aeainooeeiaeo iiethne?a, a e aeaeiaee oa /enaeueiee neeaae
aaaaoiaeaeiaeo oa?oiiaaiue oacy?a. ?iooci??y Conchoscutum inversum Raabe
aoea aeyaeaia o iiethne?a Unio pictorum eeoa a noi?nieo oa?oiiaaiiyo c
aeaao?ieoeyie. O noi?nieo oa?oiiaaiiyo iiethne?a U. tumidus oa U.
pictorum niinoa??aaeinue c?inoaiiy ?ioaineaiino? ?iaac?? ia?aceoii
Aspidogaster conchicola iiethne?a U. pictorum oa iiyaa ?iooci???
Conchophthirus anodontae (Ehrbg.) a iiethneao U. tumidus. ?iooci??y C.
acuminatus aoea ciaeaeaia o ia?e?aieoeaaeo eeoa i?e iayaiino? ia?inoaiue
ae?aenai ia /a?aiaoeao.

Oaeei /eiii, aeey a?eueoino? aeyaeaieo aeae?a aiaeia?iio?a ?niothoue
iaai? aeaee iiethne?a-oacy?a, a yeeo ?icaeaa?oueny a?eueoa /anoeia
iiioeyoe?? ia?aceoa aai eiiainaea. I?enooi?noue oeeo aeae?a oacy?a o
neeaae? neia?ioeaiico aaaea aei c?inoaiiy aenoaineaiino? ?iaac?? ?ioeo
aeae?a iiethne?a oa caaaa/aiiy aeaeiaiai neeaaeo ?o aiaeia?iio?a.

5.4. Iio?e aia?a?? /a?ac nenoaio “aeainooeeia? iiethnee — aiaeia?iioe”.

Aoei aeine?aeaeaii iio?e aia?a?? /a?ac neia?ioe/io nenoaio: “iiethnee
Dreissena polymorpha — ?iooci??? Conchophthirus acuminatus”. Ia iniia?
i?iaaaeaiiai aiae?co aecia/aii, ui ia i?ioyc? aaaaoaoe?eiiai ia??iaeo (c
o?aaiy ii aa?anaiue) /a?ac neia?ioe/io nenoaio “ae?aenaie-?iooci???”
i?ioiaeeoue aia?aaoe/iee iio?e aea?aaeaioiee 117-200 eAeae/aeiao.i2
noano?aoo iaoeaiiy iiethne?a. Naia oaeo aaee/eio neeaaeaa aeieia?/iee
?aoe?ii ae?aenai, oiaoi e?euee?noue a?aeo?eueo?iaaieo /anoie nanoiio. A
na?aaeiueiio 35% aia?a?? a?ae anueiai a?aeo?eueo?iaaiiai ae?aenaiaie
nanoiio niiaeeaaoueny aeine?aeaeoaaiith nenoaiith, neeaaeath/e ?aoe?ii
iiethne?a oa ?iooci??e. I?e oeueiio aeecei 21% aia?a?? anei?eth?oueny,
aeo?aoe ia iai?i oa i?iaeoeoe?y neeaaeathoue 12,7 oa 8,5%, a?aeiia?aeii.
Ae?aenaie inaaeaeoaaee o aeaeyae? aaethoeiao?a aeecueei 65% an???
a?aeo?eueo?iaaii? iane nanoiio.

Iaei??th c eaiie eaioethaa aeeaeaiiy, aeaea?aeii yeiai ? aia?a?y, ui
i?noeoueny a aaethoeiaoao, ie ?icaeyaea?ii eiiainaeueieo ?iooci??e c
iaio?eii? ii?iaeieie iiethne?a. ?iooci??? Conchophthirus acuminatus
caeaoi? aeei?enoiaoaaoe aeecueei 0,03-0,26% oe??? aia?a??. Aeiey
eiiainaeueieo ?iooci??e o aeano?oeoe?? i?aai?/ieo ?a/iaei caeiiii??ii
c?inoa? c? ca?eueoaiiyi aenoaineaiino? oa ?ioaineaiino? ?iaac??.
Ei?aeyoe?eiee aiae?c aeyaea aeinoia??iee ca’ycie i?ae /anoeith ?iooci??e
o niiaeeaaii? aaethoeiao?a oa iieacieeaie ?iaac??. Eiao?oe??ioe
ei?aeyoe?? neeaaeaee 0,89 oa 0,90 aeey aenoaineaiino? oa ?ioaineaiino?
?iaac??, a?aeiia?aeii.

5.5. A?ioeaiioe/i? ca’ycee aiaeia?iio?a c ?ioeie aeaeaie oacy?a.

Eeoa aiaeia?iioe c i?yiei oeeeeii ?icaeoeo iia’ycai? o?eueee c
iiioeyoe?yie aeainooeeiaeo iiethne?a. ?io? aeaee aiaeia?iio?a
(ia?aaaaeii ??ci? aeaee o?aiaoiae) iia’ycai? oaeiae ?c iiioeyoe?yie
ioao?a, caiiiaiaeieo, ?ea, /a?aaiiiaeo iiethne?a, eiiao oa
?aeiiiae?aieo, oiaoi ioiieththoue o nai?io ?icaeoeo ??ciiiai?oi?
neeaaeia? a?ioeaiico ? iaa?oue ??ci? aeinenoaie.

Aeaye? aeaee aiaeia?iio?a ia?aceoothoue o aeainooeeiaeo iiethneao eeoa
ia ee/eieiaeo noaae?yo naiai aeeoo?aiai oeeeeo, oiae? ye aei?ine?
i?aai?cie ? a?eueiiaeeao/eie (ee/eiee a??/aea, i?ioe ee?u?a).

Aeeiai?ea aeine?aeaeaii? neia?ioeaiioe/ii? nenoaie iia’ycaia ia eeoa c?
ci?iaie o iiioeyoe?yo aeainooeeiaeo iiethne?a oa ?o aiaeia?iio?a, a
oaeiae ?c i?ioeanaie, ui i?ioiaeyoue a a?ioeaiicao a oe?eiio. Cieeiaiiy
i?ii?aeieo oacy?a aiaeia?iio?a c iai?yieie oeeeeaie ?icaeoeo aaaea aei
ae?i?iaoe?? /anoeie iiioeyoe?? inoaii?o, ui iia’ycaia ?c aeaiei aeaeii
oacy?a, aai ?c cieeiaiiyi aeaeo a oe?eiio c aeaii? aeinenoaie (o
aeiaaeeo aocueei? ainoaeueii? niaoeeo?/iino?). Ca iaoith aeoieith,
iayai?noue a ??cieo neeaaeiaeo a?ioeaiico ??ciiiai?oieo aiaeia?iio?a ia
??cieo noaae?yo nai?o aeeoo?aeo oeeee?a iiaea na?ae/eoe i?i oa, ui a
a?ioeaiic? ia ii?ooai? no?oeoo?i? ca(ycee i?ae eiai neeaaeiaeie
eiiiiiaioaie.

AENIIAEE.

1. Aeine?aeaeaii neia?ioeaiioe/io nenoaio «aeainooeeia? iiethnee —
aiaeia?iioe» o neeaae? yei? aoei aeyaeaii 10 aeae?a aeainooeeiaeo
iiethne?a oa 24 aeaee aiaeia?iio?a. O ae?e? Aeanie aei neeaaeo
neia?ioeaiico aoiaeeoue 9 aeae?a iiethne?a oa 23 aeaee aiaeia?iio?a, ia
aa?oi?e ae?eyioe? Eai?anueeiai aiaeinoiaeua — 8 aeae?a iiethne?a oa 17
aeae?a aiaeia?iio?a. O ia?eo?oii? neia?ioeaiic i?aaenoaaeaii 2 aeaeaie
iiethne?a oa 6 aeaeaie aiaeia?iio?a, o aaioin?- 10 aeaeaie iiethne?a oa
23 aeaeaie aiaeia?iio?a.

2. O ae?e? Aeanie ii /enaeueiino? oa a?iian? aeii?iothoue iiethnee Unio
tumidus, U.pictorum, Rivicoliana rivicola (o aaioin?), aeainooeeia?
iiethnee o neeaae? ia?eo?oiio aoee a?aenooi?. Ia aa?oi?e ae?eyioe?
Eai?anueeiai aiaeinoiaeua o aaioin? ii /enaeueiino? oa a?iian?
aeii?iothoue Colletopterum piscinale, Anodonta cygnea, U. pictorum,
Dreissena polymorpha; o ia?eo?oii? cono??/aeenue 2 aeaee iiethne?a —
Dreissena polymorpha oa D.bugensis, ii /enaeueiino? oa a?iian?
aeii?ioaaee iiethnee D. bugensis.

3. Iiioeyoe?? iiethne?a U. tumidus, D. polymorpha, D. bugensis,
Rivicoliana rivicola ia i?ioyc? aaaaoaoe?eiiai ia??iaeo i?aaenoaaeai?
on?i niaeo?ii ?ici??ii-a?eiaeo oa noaoaaeo a?oi. O iiioeyoe?yo
Colletopterum piscinale, Anodonta cygnea, U. pictorum iiyaa
oeueiai??/ii? iieiae? oa thaai?eueieo iniaei niinoa??aaeanue a
eeii?-aa?ani?, iiethne?a Pisidium amnicum — o o?aai?-/a?ai?. A ?io?
i?nyoe? iiioeyoe?? iiethne?a oeeo aeae?a ia e?oi?ae? i?aaenoaaeai?
ia?aaaaeii noa?oeie ?ici??ii-a?eiaeie a?oiaie (noaoaaic??eeie
iniaeiaie). O noaoaa?e no?oeoo?? iiioeyoe?e ia?aaaaeaee naioe? (55-75
%).

4. O iiethne?a D. polymorpha oa D. bugensis niinoa??aaeenue a?aei?iiino?
o ia??iaeao ?iciiiaeaiiy. Iaeneioi iiyae oeueiai??/ii? iieiae? o
iiethne?a D. polymorpha i?eiaaeaa ia na?iaiue i?nyoeue, a D. bugensis —
ia aeiaoaiue.

5. Aeey 12 aeyaeaieo aeae?a aiaeia?iio?a aeainooeeia? iiethnee ?
aeao?i?oeaieie oacyyie, aeey 9 aeae?a — ia?oeie i?ii?aeieie, aeey 3
aeae?a — ae?oaeie i?ii?aeieie oacyyie. Na?aae aiaeia?iio?a c i?yiei
oeeeeii ?icaeoeo ii e?eueeino? aeae?a ia?aaaaeathoue ?iooci??? (10
aeae?a), c iai?yiei oeeeeii ?icaeoeo — o?aiaoiaee (10 aeae?a).

6. Aiaeia?iioe c i?yieie aeeoo?aeie oeeeeaie (?iooci??? oa
ani?aeiaano?e) iaea?eueoiai ?icaeoeo aeinyaathoue o aaioin? ae?ea Aeanie
a iiioeyoe?yo iiethne?a ?iaeo Unio oa a ia?eo?oii? aa?oiuei? ae?eyiee
Eai?anueeiai aiaeinoiaeua a iiioeyoe?yo ae?aenai.

7. Iaea?eueoiai ?icaeoeo ia?aceoe c iai?yieie oeeeeaie (o?aiaoiaee)
aeinyaathoue a aaioin? ae?ea Aeanie o iiioeyoe?? iiethnea Pisidium
amnicum (ia?oai?oe oaiaoiae) oa a ia?eo?oii? aa?oiuei? ae?eyiee
Eai?anueeiai aiaeinoiaeua (iaoaoea?ea??? o?aiaoiae).

8. O iiethne?a D.polymorpha oa D.bugensis aeyaeaii o?aiaoiaeo
Bucephalus polymorphus, ui ia?aceoo? o aiiaae?, aeeeeeath/e iiaio aai
/anoeiao noaoaao noa?ee?caoe?th oacy?ia. Oeae oaiiiai iiaea aooe
aeei?enoaiee aeey ?ic?iaee a?ieia?/ieo iaoiae?a ai?ioueae c
a?iia?aoeiaeaie o aiaeiiinoa/aii?, noai?thaaieie ae?aenaiaie.

9. Aia?oa ia Oe?a?i? a?aei?/aii iayai?noue ieaneiai /a?a’yea
Aspidogaster limacoides, ye ia?aceoa aeainooeeiaeo iiethne?a. A.
limacoides aoa aeyaeaiee o iiethne?a ?iaeo Dreissena o ia?eo?oii?
aa?oiuei? ae?eyiee Eai?anueeiai aiaeinoiaeua.

10. Ocaaaeueithth/a aeie-noaia a?ioeaiioe/ieo ca’yce?a na?ae/eoue, ui
on?o aiaeia?iio?a ?c aeainooeeiaeie iiethneaie iia’ycothoue
aaciina?aaei? oii?/i?, o?io?/i? oa oi?e/i? ca’ycee. Ie??i i?yiiai
o?io?/iiai ca’yceo eiiainaeuei? ?iooci??? c iaio?eii? ii?iaeieie
iia’ycai? iiina?aaeeiaaiei o?io?/iei ca’yceii ?c iiethneaie — oacyyie.

11. Aeey on?o aeine?aeaeaieo aeae?a iiethne?a niinoa??aaeinue c?inoaiiy
aenoaineaiino? ?iaac?? o noa?oeo ?ici??ii-a?eiaeo a?oiao oa o
noaoaaic??eeo iiethne?a, ii??aiyii c iieiaeoeie a?oiaie oa thaai?eueieie
iniaeiaie. I?enooi?noue o neeaae? iiioeyoe?e naia oeeo a?oi aecia/a?
aenieo aenoaineai?noue ?iaac?? iiioeyoe?? a oe?eiio. Iiyaa oeueiai??/ii?
iieiae? oa thaai?eueieo iniaei o iiioeyoe?? oacy?ia aaaea aei ciaioaiiy
aaee/eie oeueiai iieacieea.

12. Ia i?ioyc? aaaaoaoe?eiiai ia??iaeo (c o?aaiy ii aa?anaiue) /a?ac
neia?ioe/io nenoaio “iiethnee D. polymorpha — ?iooci??? Conchophthirus
acuminatus” i?ioiaeeoue aia?aaoe/iee iio?e aea?aaeaioiee 117-200
eAeae/aeiao.i2 noano?aoo iaoeaiiy iiethne?a. Ae?aenaie inaaeaeoaaee o
aeaeyae? aaethoeiao?a aeecueei 65% an??? a?aeo?eueo?iaaii? iane nanoiio.
?iooci??? Conchophthirus acuminatus caeaoi? aeei?enoiaoaaoe aeecueei
0,03-0,26% aia?a?? ooai?aieo aaethoeiao?a.

13. I?enooi?noue o ??ciiaeaeiaeo oa?oiiaaiiyo iiethne?a aeae?a, ui ?
iniiaieie oacyyie iaaieo a?oi aiaeia?iio?a aaaea aei c?inoaiiy
aenoaineaiino? ?iaac?? ?ioeo aeae?a iiethne?a oa caaaa/aiiy aeaeiaiai
neeaaeo ?o aiaeia?iio?a.

14. Cieeiaiiy i?ii?aeieo oacy?a aiaeia?iio?a aaaea aei ae?i?iaoe??
/anoeie iiioeyoe?? inoaii?o, ui iia’ycaia ?c aeaiei aeaeii oacy?a, aai
?c cieeiaiiyi aeaeo a oe?eiio c aeaii? aeinenoaie (o aeiaaeeo aocueei?
ainoaeueii? niaoeeo?/iino?). Iayai?noue a ??cieo neeaaeiaeo a?ioeaiico
??ciiiai?oieo aiaeia?iio?a ia ??cieo noaae?yo nai?o aeeoo?aeo oeeee?a
iiaea na?ae/eoe i?i oa, ui a a?ioeaiic? ia ii?ooai? no?oeoo?i? ca’ycee
i?ae eiai neeaaeiaeie eiiiiiaioaie.

Nienie ?ia?o, iioae?eiaaieo ca iaoa??aeaie aeena?oaoe??:

TH?eoeiaoe A.E. Ia?eiaeoea Unio tumidus ec onouey Aeaniu e a? ia?aceou
// I?iaeaiu aeae?iaeieiaee eiioeiaioaeueiuo aiae e eo iaeaeioaoia. Oac.
aeiee. (N.-Iaoa?ao?a, ?inney, 18-21 iiya?y 1996a). — N.-Iaoa?ao?a. —
1996. — N.65-66.

TH?eoeiaoeue A.?. ?iooci??? ?iaeo Conchophthirus (Thigmotrichida,
Conchophthiridae) o aeainooeeiaeo iiethneao ae?ea Aeanie // Ca??iee
iaoeiaeo i?aoeue aeeeaaea/?a Eai’yiaoeue-Iiae?euenueeiai iaaeaaia?/iiai
oi?aa?neoaoo. Aeione ??. — Oiaeueieoeueeee: A-ai “Iiae?eey”, 1997. — N.
172-177.

TH?eoeiaoe A.E. I?eiaiaiea iaoiaeia eeanoa?iiai e ?aa?anneiiiiai
aiaeecia aeey eco/aiey iiioeyoeee iieethneia Rivicoliana rivicola
(Bivalvia, Sphaeriidae) e Pisidium amnicum (Bivalvia, Pisidiidae) //
Aanoi. ciie. — 1997. — 31, ?3. — N. 89-91.

TH?eoeiaoe A.E. Iaeioi?ua aniaeou acaeiiaeaenoaey iiioeyoeee Unio
tumidus e Unio pictorum n eo ia?aceoaie e eiiiainaeaie // Aeae?iaeie.
aeo?i. — 1997. — 33, ?5. — N.107-109.

TH?eoeiaoe A.E. O?aiaoiaeu iaeioi?uo aeaeia oa?iaie e ai?ioei ec onouey
Aeaniu // Aeae?iaeie. aeo?i. — 1997. — 33, ?6. — N. 67-71.

TH?eoeiaoeue A.?. Ia?aceoe oa eiiainaee ia?e?aieoeaaeo oa i?ee??ieaieo
aei ieo ae?aenai // ?? c`?cae a?ae?iaeieia?/iiai oiaa?enoaa Oe?a?ie
(Ee?a, 27-31 aeiaoiy 1997?.). Oace. aeii. O. 1. — Ee?a: A?ae?iaeieia?/ia
oiaa?enoai Oe?a?ie, 1997. — N. 164-165.

Yurishinets V. Bivalve molluscs from the Mouth of the Desna River and
their parasites // VII European multicolloquium of Parasitology (2 — 6
Sept. 1996, Parma, Italy). Abstracts. — Roma: “La Sapienza”, 1996. — P.
25-26.

Yurishinets V. Some suppositions concerning life cycle of Aspidogaster
conchicola (Aspidogastrea, Aspidogastrida) // IXth International
Congress of Parasitology (ICOPA IX)(24-28 Aug. 1998, Chiba, Japan).
Abstracts. Poster Session / Parasitology International 47 (Suppl.). —
Tokyo, 1998. — P. 303-304.

Aiioaoe??:

TH?eoeiaoeue A.?. Aeainooeeia? iiethnee oa ?o aiaeia?iioe ye eiiiiiaio
a?ae?iia?aceoa?ieo nenoai.- ?oeiien.

Aeena?oaoe?y ia caeiaoooy a/aiiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
ca niaoe?aeuei?noth 03.00.17 — a?ae?ia?ieia?y. — ?inoeooo a?ae?ia?ieia??
IAI Oe?a?ie, Ee?a, 1999.

Aeena?oaoe?y i?enay/aia aea/aiith i??niiaiaeieo aeainooeeiaeo iiethne?a
oa ?o aiaeia?iio?a, ye ?aeeii? neia?ioeaiioe/ii? nenoaie. O ?iaio?
i?iaaaeaii oeaiioe/iee oa iiioeyoe?eiee aiae?c aeainooeeiaeo iiethne?a,
aeine?aeaeaii aeaeiaee neeaae oa aeaii oeaiioe/io oa?aeoa?enoeeo
iiioeyoe?e ?o aiaeia?iio?a. C’yniaaii, ye oeaiioe/i? oa iiioeyoe?ei?
oa?aeoa?enoeee aeainooeeiaeo iiethne?a aieeaathoue ia aeaeiaee neeaae oa
oeaiioe/i? oa?aeoa?enoeee iiioeyoe?e aiaeia?iio?a. Aeine?aeaeaii neeaae
oa aeeiai?eo neia?ioeaiico “aeainooeeia? iiethnee — aiaeia?iioe” o
ia?eo?oii? oa aaioin?, a oiiaao ??/eiai? aeinenoaie (ae?ei ?. Aeania) oa
aiaeinoiaeua (aa?oiy ae?eyiea Eai?anueeiai aiaeinoiaeua ?. Aei?i?i). Ia
iniia? i?iaaaeaiiai aiae?co oa aeaieo e?oa?aoo?e aeaii caaaeueio
oa?aeoa?enoeeo aeine?aeaeai?e neia?ioeaiioe/i?e nenoai?. Aia?oa o
a?ae?ia?ieia?/ieo aeine?aeaeaiiyo ?ic?aoiaaii iio?e aia?a??, ui
i?ioiaeeoue /a?ac neia?ioe/io nenoaio “iiethnee Dreissena polymorpha —
?iooci??? Conchophthirus acuminatus ”.

Eeth/ia? neiaa: neia?ioeaiic, a?ae?iia?aceoa?ia nenoaia, aeainooeeia?
iiethnee, aiaeia?iioe, no?oeoo?a iiioeyoe??, ia?eo?oii, aaioin.

TH?eoeiaoe A.E. Aeaonoai?/aoua iieethnee e eo yiaeiaeiiou eae eiiiiiaio
aeae?iia?aceoa?iuo nenoai. — ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/?iie noaiaie eaiaeeaeaoa aeieiae/aneeo iaoe
ii niaoeeaeueiinoe 03.00.17 — aeae?iaeieiaey. — Einoeooo aeae?iaeieiaee
IAI Oe?aeiu, Eeaa, 1999.

Aeenna?oaoeey iinayuaia eco/aieth i?aniiaiaeiuo aeaonoai?/aouo
iieethneia e eo yiaeiaeiioia, eae aaeeiie neiaeioeaiioe/aneie nenoaiu. A
?aaioa i?iaaaeai oeaiioe/aneee e iiioeyoeeiiiue aiaeec aeaonoai?/aouo
iieethneia, enneaaeiaai aeaeiaie ninoaa e aeaia oeaiioe/aneay
oa?aeoa?enoeea iiioeyoeee eo yiaeiaeiioia. Auyniaii, eae oeaiioe/aneea e
iiioeyoeeiiiua oa?aeoa?enoeee aeaonoai?/aouo iieethneia aeeytho ia
aeaeiaie ninoaa e oeaiioe/aneea oa?aeoa?enoeee iiioeyoeee yiaeiaeiioia.
Enneaaeiaaiu ninoaa e aeeiaieea neiaeioeaiioe/aneie nenoaiu
“aeaonoai?/aoua iieethnee — yiaeiaeiiou ” a ia?eoeoiia e aaioina, a
oneiaeyo ?a/iie yeinenoaiu (onouea ?. Aeania) e aiaeio?aieeeua (aa?oiee
o/anoie Eaiaaneiai aiaeio?aieeeua ?. Aeiai?). Ia iniiaaiee i?iaaaeaiiiai
aiaeeca e aeaiiuo eeoa?aoo?u aeaia iauay oa?aeoa?enoeea enneaaeiaaiiie
neiaeioeaiioe/aneie nenoaia. Aia?aua a aeae?iaeieiae/aneeo
enneaaeiaaieyo ?ann/eoai iioie yia?aee, eioi?ue i?ioiaeeo /a?ac
neiaeioe/aneoth nenoaio “iieethnee Dreissena polymorpha — eiooci?ee
Conchophthirus acuminatus”.

Eeth/aaua neiaa: neiaeioeaiic, aeae?iia?aceoa?iay nenoaia,
aeaonoai?/aoua iieethnee, yiaeiaeiiou, no?oeoo?a iiioeyoeee, ia?eoeoii,
aaioin.

Yurishinets V.I. Bivalve molluscs and their endobionts as component of
hydroparasitical systems. — Manuscript.

Thesis for a scientific degree of the candidate of biological sciences
by speciality 03.00.17 — hydrobiology. — Institute of a hydrobiology
National Academy of Science of Ukraine, Kiev, 1999.

The dissertation is devoted to study fresh-water bivalve molluscs and
their endobionts, as united symbiocenosis. Cenotic and population
analysis of bivalves and their endobionts has been investigated.
Influences of cenotic and population characteristics of bivalves on
species diversity and cenotic characteristics of endobionts were
studied. The structure and dynamics symbiocenosis “bivalve molluscs —
endobionts” in periphyton and benthos in river ecosystem (mouth of Desna
River) and reservoir ecosystem (upper site of Kanev Reservoir of Dnipro
River) were investigated. Complex characteristic of symbiocenosis is
proposed, based on own data and literature analysis. For the first time
in hydrobiological researches the stream of an energy which passes
through symbiotic system “molluscs Dreissena polymorpha — ciliates
Conchophthirus acuminatus” is calculated ”.

Key word: symbiocenosis, hydroparasitical system, bivalve molluscs,
endobionts, structure of a population, periphyton, benthos.

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?en. 1 Noaia neio?c?ieia?/ieo ca(yce?a o nenoai? “aeainooeeia? iiethnee
— aiaeia?iioe”.

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