— 8 —

OE?A?INUeEA AEAAeAI?ss AA?A?IEO IAOE

?INOEOOO OAA?EIIEOeOAA OAAI

IAAEAIEI Ethaeieea Ieeiea?aia

OAeE 57.08:636.22/.28.453.5

AeIAAICAA?AAEIA NA?AAeIAEUA AeEss E??IEIINA?AAOe?? NIA?IE
AOAA?A-IE?AeIEE?A OA NIINIAE EIAI AEAIOIAEAIIss

3.00.20 — a?ioaoiieia?y

Aaoi?aoa?ao

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

Oa?e?a — 1998

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia a ?inoeooo? oaa?eiieoeoaa OAAI.

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, i?ioani?, aeaaeai?e OAAI
Inoaoei Oaae?? ?aaiiae/.

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani? Aoa?ia Ieaen?e
Aeieo?iae/, aieiaiee iaoeiaee ni?a?ia?oiee a?aeae?eo a?ioaoiieia?? oa
aaiii? ?iaeaia??? ?inoeoooo oaa?eiieoeoaa OAAI;

eaiaeeaeao a?ieia?/ieo iaoe E?aia? Ia??y Einei?aia, iaoeiaee
ni?a?ia?oiee ?inoeoooo i?iaeai e??ia?ieia?? ? e??iiaaeeoeeie IAI
Oe?a?ie.

I?ia?aeia onoaiiaa: Aea?aeaaiee iaoeiai-aeine?aeiee ?inoeooo
aaoa?eia?ieo i?aia?ao?a ? ei?iiaeo aeiaaaie. Euea?a

Caoeno aeena?oaoe?? a?aeaoaeaoueny “23” aa?aciy 1999 ?ieo i 10-00 aiae.
ia can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee Ae.65.356.02 i?e ?inoeooo?
oaa?eiieoeoaa OAAI.

C aeena?oaoe??th iiaeia iciaeiieoeny o a?ae?ioaoe? ?inoeoooo
oaa?eiieoeoaa OAAI. I?ineii i?eeiyoe o/anoue a can?aeaii? ?aaee, aai
aeneaoe a?aeaoe a aeaio i?ei??ieeao, caoaa?aeaeaieo ia/aoeith.

Aae?ana ?inoeoooo: 312120, i. Oa?e?a, i/a Eoeeie/?

Aaoi?aoa?ao ?ic?neaii “ 16 ” ethoiai 1999 ?.

A/aiee nae?aoa? niaoe?ae?ciaaii?

a/aii? ?aaee, eaiaeeaeao n.-a. iaoe O.
I. Niaao?

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. A ?aae?caoe?? e?oiiiianooaaii? naeaeoe?? oaa?ei
iaeia c i?ia?aeieo i?noeue caeia? ooo/ia in?iai?iiy. Iniiaith
aeiei??iii? ia?aaoaeiae i?aai?caoe?? ooo/iiai in?iai?iiy noaa iaoiae
aeeaieiai caii?iaeoaaiiy ? o?eaaeiai caa??aaiiy nia?ie ie?aeiee?a
n?euenueeiainiiaea?nueeeo oaa?ei, ia iniia? ooiaeaiaioaeueieo
aeine?aeaeaiue a?o/eciyieo oa ca?oa?aeieo a/aieo: (?.?.?aaiia, 1910;
A.E.I?eiaaiia, 1962; ?.A.Ni??iia, 1949; O.?.Inoaoei, 1968; A.I.Ieaoia,
1961; I.Ae.Aoa?ia, 1964; C.Polge, A.U.Smith, A.S.Parkes, 1948, 1949;
H.Nagase, T.Niwa, 1964; R.Cassou, 1964; L.Simmet, 1972). Caaaeyee
aeeeth/i?e aoaeoeaiino? a?i noaa iniiaiei aeaiaioii iaoe?iiaeueieo
i?ia?ai ii ?iciiiaeaiith ? iie?auaiith oaa?ei an?o oeea?e?ciaaieo e?a?i
na?oo. Iaeiae, aeaye? ae??oaeuei? aniaeoe oaoiieia?? e??ieiina?aaoe??
nia?ie aea/ai? iaaeinoaoiuei, ui ia aea? ciiae aeei?enoiaoaaoe eiai c
iaeneiaeueiith aoaeoeai?noth. ?aae?caoe?y iioaioe?eieo iiaeeeainoae
iaoiaeo e??ieiina?aaoe?? nia?ie an?eyei caeaaeeoue a?ae neeaaeo ooo/ieo
na?aaeiaeu oa ?o nai?oa?ieo aeanoeainoae.

Iaia’yceiaei aioeoieiaei eiiiiiaioii a?aeiieo na?aaeiaeu ? aeiaoie
eo?y/eo y?oeue, ui iaiaaeo? no?iee caa??aaiiy ? ia aea? iiaeeeaino?
oeaio?ae?ciaaiiiai aeaioiaeaiiy oa iinoa/aiiy ?o ieai?iiei
i?aei?e?inoaai. E??i oiai, aeiaoie aoaeo/e oi?ioei aeeaeeueiei
na?aaeiaeuai aeey i?e?ii?aai?ci?a a ?yae? aeiaaee?a ? iin??i iaoiaaiii?
? nai?io?oii? i?e?ioei?e, yea iaaaoeaii aieeaa? ia nia?io, a oaeiae ia
a?aeoai?thaaeueio ooieoe?th ? noai caei?ia’y naieoeue. O ca’yceo c oeei
aeieeea iano?eia iaiao?aei?noue noai?aiiy noa?eeueieo aeiaaicaa?aaeieo
aeieia?/ii /enoeo na?aaeiaeu aeey e??ieiina?aaoe?? nia?ie ie?aeiee?a.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. ?iaioa aeeiiaia
a ?aieao iaoeiai-oaoi?/ieo eiiieaenieo i?ia?ai AANAI?E ino. 42.0107.I3;
ino. 77.08.02/?? aea?ae?a?no?aoe?? 010001316?; 01.8600048.345 oa IO?
OAAI ia 1991-1995 ??. “Oaeineiiaeeoe iaoiaee e??ieiina?aaoe?? oa
aeiaaicaa?aaeaiiy nia?ie n?euenueeiainiiaea?nueeeo oaa?ei c iaoith
iaeneiaeueiiai aeei?enoaiiy aaiaoe/ieo ?aca?a?a ieai?iiiai
oaa?eiieoeoaa”, “?ic?iaeoe ia?niaeoeai? iiaeae? ainiiaea?noa ii
ae?iaieoeoao iieiea, caaacia/oth/eo aenieo i?iaeoeoeai?noue oaa?ei,
iie?auoth/eo ye?noue i?e cieaeaii? aeo?ao ei?i?a, i?aoe? oa o?iain?a”.

Iaoa aeine?aeaeaiue — ?ic?iaeoe ? ai?iaaaeeoe noa?eeueia
aeiaaicaa?aaeia na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie aoaa?a,
noaiaea?oeciaaia ca a?io?i?/ieie ? nai?oa?ieie iieacieeaie oa niiniae
eiai aeaioiaeaiiy ? oaeineiiaeeoe ia oe?e iniia? oaoiieia?th
e??ieiina?aaoe?? nia?ie ie?aeiee?a.

Caaaeaiiy aeine?aeaeaiue:

1. ?ic?iaeoe niin?a iaea?aeaiiy c iaoeaiiai aeiaoea eiiiiiaioa c
aioeoieiaeie aeanoeainoyie.

2. Aea/eoe caoeni? aeanoeaino? eiiiiiaioo i?e ioieiaeaeoaaii? ia nia?i?
aoaay, aa?aia oa eio?a.

3. ?ic?iaeoe neeaae oa?iinoaa?eueiiai aeiaaicaa?aaeaiiai na?aaeiaeua
aeey e??ieiina?aaoe?? nia?ie aoaa?a ? oa?i?/iee niin?a eiai
noa?ee?caoe??.

4. ?ic?iaeoe niin?a o?i?ei-oa?i?/ii? noa?ee?caoe?? na?aaeiaeua c
iaoeaiei aeiaoeii.

5. ?ic?iaeoe anaioe/iee niin?a oa i?eno??e aeey ioe?iee nia?ie a
aa?iaoe/ieo oiaeiaeao.

6. I?iaanoe iaoeiai-ae?iaie/ee oa oe?ieee ae?iaie/ee aeine?ae ?
ai?iaaaeeoe ?ic?iaeaia na?aaeiaeua a ae?iaieoeoai.

Iaoeiaa iiaecia oa oai?aoe/ia oe?ii?noue aeine?aeaeaiue. Aia?oa
onoaiiaeaii, ui aeno?aeo aeiaoea a neeaae? e??ii?ioaeoeaiiai na?aaeiaeua
i?ney oa?iiia?iaee caa??aa? nai? iiia?aaei? a?ieia?/i? aeanoeaino? ?
caaacia/o? caoeno noaoaaeo ee?oei i?e e??ieiina?aoaaii?, aiaeia?/ii
iaoeaiiio aeiaoeo. Ia oe?e i?aenoaa? noai?aii oa?iinoaa?eueia noa?eeueia
na?aaeiaeua, aea iaoeaiee aeiaoie cai?iaii oa?iinoaa?eueiei aioeoieiaei
eiiiiiaioii (a.n. N?N? ? 745045).

Aia?oa onoaiiaeaii aoaeo ca?eueoaiiy aeoiaeo aioeoieiaiai eiiiiiaioo
i?ney aeno?aaoaaiiy eiai ?c caii?iaeaii-a?aeoaeiai aeiaoea.

Aia?oa aea/aia ae?y aioenaioeea — ia?aeeno aiaeith a neeaae?
na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie ? aeyaeaia aoaeoeaia
n?ia?a?aeia aioei?e?iaia ae?y eiai ni?eueii c i?ia??aii 65-75°N, ui
caaacia/o? iiaio noa?eeuei?noue oa?iieaa?eueiiai na?aaeiaeua ? ia
cieaeo? eiai caoenieo aeanoeainoae i?e e??ieiina?aaoe?? nia?ie. Ia oe?e
iniia? noai?aii noa?eeueia, aeiaaicaa?aaeia aeiaoieoo?eiath/a
na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie ? niin?a eiai aeaioiaeaiiy
(iaoaio N?N? ? 1828582 ? iaoaio Oe?a?ie ? 13138).

Aia?oa aeiaeaeaii niin?a oa ?ic?iaeaii i?eno??e aeey anaioe/ii? ioe?iee
nia?ie a aa?iaoe/i?e oiaeiaoe? (a.n.C?N? ? 1543598).

I?aeoe/ia cia/aiiy ?iaioe. Io?eiai? ?acoeueoaoe aeaee iiaeeea?noue
aeae?eeoe c iaoeaiiai aeiaoea oa?iinoaa?eueiee aioeoieiaee eiiiiiaio ?
aeei?enoaoe eiai a iaoeiai-aeine?aei?e ?iaio? i?e ?ic?iaoe?
oa?iinoaa?eueiiai aeiaaicaa?aaeiiai na?aaeiaeua aeey e??ieiina?aaoe??
nia?ie aoaa?a.

Aeyaeaiiy aioei?e?iaii? ae?? ia?aeeno aiaeith a aeiaoieoo?eiath/eo
na?aaeiaeuao aeicaieeei noai?eoe aeiaaicaa?aaeaia na?aaeiaeua aeey
e??ieiina?aaoe?? nia?ie ? oaoiieia?th eiai ae?iaieoeoaa, yea noaaeaia ?
?aeiiaiaeiaaia aeey ai?iaaaeaeaiiy I?aea?aeii/ith eii?n??th I?iaa?ii?iio
Oe?a?ie (i?ioieie ? 7 a?ae 10 ea?oiy 1992 ?.). Ia oe?e i?aenoaa?
i?aai?ciaaii oeaio?ae?ciaaia ae?iaieoeoai na?aaeiaeu ? iinoa/aiiy ?o
ieai?iiei i?aei?e?inoaai Oe?a?ie ? ?in?enuee?e Oaaea?aoe?? (aeo
ai?iaaaeaeaiiy IIAA “Aia??ii” a?ae 3 n?/iy 1998 ?.), ui aeeeth/eei
iiaeeea?noue caa?oaeiaiiy nia?ie ? noaoaaeo oeyo?a naieoeue i?e ooo/iiio
in?iai?ii? i?e?ioei?ith, yea iiaea ia?aaeaaaoenue ianoa?eeueieie
?icaaaethaa/aie.

C iaoith i?aeaeuaiiy nai?oa?ii-a?a??i?/iiai ??aiy ?iaioe ioieo?a
ooo/iiai in?iai?iiy ? ?aoe?iiaeueiiai aeei?enoaiiy nia?ie, noai?aii ?
ai?iaaaeaeaii a ae?iaieoeoai anaioe/iee niin?a oa i?eno??e aeey ioe?iee
nia?ie a aa?iaoeciaaieo nia?iiaeicao.

Iniaenoee aianie aeena?oaioa a ?ic?iaeo iaoeiaeo ?acoeueoao?a, ui
aeiinyoueny ia caoeno. Aeena?oaoe?eia ?iaioa aeeiiaia iniaenoi aaoi?ii,
naiino?eii i?iaaaeai? an? aenia?eiaioaeuei? oa oai?aoe/i?
aeine?aeaeaiiy, iaoeiai-ae?iaie/? aeine?aee i?iaaaeai? c iniaenoith
o/anoth aaoi?a ia aac? aenia?eiaioaeueii-iaoiaee/ii? noaioe?? ooo/iiai
in?iai?iiy aeine?aeiiai ainiiaea?noaa “Oe?a?iea”, Oa?e?anueeiai
ieaiia’?aeiaiiy oa I?aeia?iaeii? iaoeiai-ae?iaie/i? anioe?aoe??
“Aia??ii”.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. ?acoeueoaoe aeine?aeaeaiue ?
iniiai? iieiaeaiiy aeena?oaoe?? aeeeaaeai? ? iaaiai?ai? a aeiiia?aeyo
Ananithcii? eiioa?aioe?? “E??ia?ieia?y ? e??iiaaeeoeeia”, 1984, Oa?e?a;
10 th International Congress on Animal Reproduction and Artificial
Insemination (10-14 June, 1984 — Illinois, U.S.A.); iaoeiai-ae?iaie/?e
eiioa?aioe?? “Iiaua iaoiaeu e aeioaoiieiaey a aeeaioiiaiaenoaa”, 1991,
Eeaa; I?aeia?iaei?e eiioa?aioe?? “Oniaoe a nia?aiaiiie e?eiaeieiaee”
(21-25 ai?aey 1992, Oa?ueeia). A iiaiiio ia’?i? ?acoeueoaoe
aeine?aeaeaiue iaaiai?thaaeenue ia ia’?aeiaiiio can?aeaii? a?aeae?e?a
a?ieia?? ?iciiiaeaiiy oa ooo/iiai in?iai?iiy n?euenueeiainiiaea?nueeeo
oaa?ei, a?ioaoiieia?? a?aeoai?aiiy oaa?ei oa o?c?ieia??
n?euenueeiainiiaea?nueeeo oaa?ei.

Iniiai? ?acoeueoaoe aeena?oaoe?? aeeeaaeai? o 3 iaoeiaeo i?aoeyo, 5
iaoa??aeao iaoeiaeo eiioa?aioe?e ? eiia?ano, 2 aaoi?nueeeo na?aeioeoaao,
2 iaoaioao Oe?a?ie oa ?in??.

No?oeoo?a oa ianya aeena?oaoe??. Aeena?oaoe?y neeaaea?oueny ?c anooio,
iniiaii? /anoeie, aeniiae?a, ?aeiiaiaeaoe?e ae?iaieoeoao, aeiiiaiaiiy ?
nieneo aeei?enoaieo aeaea?ae. ?iaioa aeeeaaeaia ia 132 noi??ieao
iaoeiiieniiai ae?oeo, i?noeoue 24 oaaeeoe?, 3 ?ethno?aoe??, 9
aeiaeaoe?a. Nienie e?oa?aoo?e aeeth/a? 212 aeaea?ae, a oiio /ene? 62
?iicaiieo aaoi??a.

Iaoa??aee ? iaoiaee aeine?aeaeaiue

Ia’?eoii aeine?aeaeaiue aoee: nia?ia aoaa?a, aa?ai?a ? eio??a, aeiaoie
eo?y/eo y?oeue ? eiai eiiiiiaioe, naieoe? aaeeei? ?iaaoi? ooaeiae.
Oa?iinoaa?eueiee aioeoieiaee eiiiiiaio aeeo/aee c aeiaoea aeno?aaoaaiiyi
ia neiino?oeiaaiiio iaie i?eno?i?. A?aeieaaiiy nia?i??a a?ae caoenieo
eiiiiiaio?a cae?enithaaee oeaio?eooaeiei iaoiaeii, i?iao ia
oaiia?aoo?iee oie i?iaiaeeee ca AeANO 20909.4-75. I?aea?? eiiiiiaio?a
na?aaeiaeua i?iaiaeeee iaoiaeii cono??/ieo ?yae?a ca A.E.I?eiaaiiaei,
1962. Iniioe/iee oene na?aaeiaeua aei??thaaee e??ineii?/iei iaoiaeii.
Eiioeaio?aoe?th aiaeiaaeo eii?a (?I) aecia/aee ia ?iiiiao?? AA-74 (ca
AeANO 20909.5-75). Caii?iaeoaaee nia?io ca Oa?e?anueeith oaoiieia??th.
?ooeea?noue, aeaeeaai?noue a aiaeeiao, iieaciee aaniethoii? aeaeeaaiino?
nia?i??a aecia/aee, ca?aeii c /eiieie AeANOaie ia iaoeaio ? caii?iaeaio
nia?io (AeANO 23745-79, AeANO 26030-83, AeANO 27777-88). Aeine?aeaeaiiy
aenia?eiaioaeueieo c?ace?a na?aaeiaeu ia aaeoa??ieia?/io caa?oaeiai?noue
i?iaiaeeee ca iaoiaeeeith, caoaa?aeaeaiith AOA INA N?N?, 1969. An?
aeine?aeaeaiiy aeeiioaaee ia ?icae?eaieo ayeoeyoao.

A on?o aeine?aeao i?e aea/aii? caie?aeithth/i? caeaoiino? ii??aiyeuei?
a?oie oaa?ei i?e in?iai?ii? i?aeae?aee ca i?eioeeiii ia?-aiaeia?a.
Naieoeue in?iaiyee aeaie?aoii a iaeio ioioo iaii- aai a?cioea?a?eaeueiei
niiniaii. Io?eiai? aeai? i?aeaeaaaee noaoenoe/i?e ia?iaoe?
caaaeueiii?eeiyoeie iaoiaeaie. Ioe?ieo ?acoeueoao?a i?iaiaeeee ca
e?eoa???i Nouethaeaioa. ??cieoey aaaaeaeanue a??ia?aeiith i?e ? < 0,05. ?ACOEUeOAOE AeINE?AeAEAIUe OA ?O IAAIAI?AIIss sse na?ae/aoue io?eiai? ?acoeueoaoe eiiiiiaioe eo?y/iai aeiaoea aeno?aaiaai? i?aai?/ieie ?ic/eiieeaie (nie?o, ao??) ? aaaaeai? a na?aaeiaeua aeey ?icaaaeaiiy nia?ie a eiioeaio?aoe?? 3% caiia?aathoue oaiia?aoo?iiio oieo noaoaaeo ee?oei aoaay, aa?aia oa eio?a ia iaeiiio ??ai? c 30%-iith eiioeaio?aoe??th iaoeaiiai aeiaoea, i?i ui na?ae/aoue aeai? oaaeeoe? 1. C oaaeeoe? 1 aeaeii, ui ?ooeea?noue nia?i??a aoaay a na?aaeiaeu? c iaoeaiei aeiaoeii aei oieo noaiiaeea 7,1 aae?a, a i?ney oieo 3,9 aae?a, ??cieoey i?ae iieacieeaie ?ooeeaino? noaoaaeo ee?oei aoea aenieia??ia?aeiith (? < 0,001), i?e oeueiio eiao?oe?aio e??i?acenoaioiino? (R) noaiiaea 0,55. A aeine?ae? ?ooeea?noue nia?ie, ?icaaaeaii? ?ic??aeaeoaa/ai c ai?noii aioeoieiaiai eiiiiiaioo noaiiaeea 6,8 aae?a, a i?ney oieo 3,95 aae?a, i?e oeueiio eiao?oe?aio e??i?acenoaioiino? noaiiaea 0,58, ui aoei ia ??ai? eiio?ieth. I?e aea/aii? ca’yceo aioeoieiaiai eiiiiiaioo c oeeoiieaciaoe/ieie iaia?aiaie noaoaaeo ee?oei i?iaiaeeee aaaaoi?aciaa oeaio?eooaeia a?aeieaaiiy eiai a?ae nia?i??a ?cioii?/iei oeoe?iai-nieueiaei aooa?ii c iine?aeoth/ei aecia/aiiyi no?eeino? ?o aei oaiia?aoo?iiai oieo. I?e oeueiio ia nia?i? aoaay eiao?oe?aio e??i?acenoaioiino? (R) noaiiaea i?ney ia?oiai a?aeieaaiiy 0,58, i?ney i’yoiai 0,58. A eiio?ie? oe? iieacieee aoee a?aeiia?aeii 0,55 ? 0,61. ??cieoey i?ae aeine?aeii ? eiio?ieai aoea noaoenoe/ii iaa??ia?aeia (? > 0,5). Aiaeia?/ia
caeaaei?noue aeyaeaia ? a aeine?aeao ia noaoaaeo ee?oeiao aa?aia ?
eio?a. Iieaciee ?ooeeaino? nia?i??a aoaay i?ney ?icaaaeaiiy ?
iine?aeoth/eo a?aeieaaiue a aeine?aeieo ? eiio?ieueieo c?aceao
a??ia?aeii cieaeoaaany (? < 0,01), a eiao?oe??io e??i?acenoaioiino? eeoaany ia iaeiiio ??ai? (? > 0,5). C aeaieo aene?aeaeaiue aeo?ea?, ui
aioeoieiaee eiiiiiaio (3%) ? iaoeaiee aeiaoie (30%) caoeuathoue nia?i??
aoaay, aa?aia ? eio?a a?ae oaiia?aoo?iiai oieo ia iaeiiio ??ai?.

Oaaeeoey 1

Aieea aioeoieiaiai eiiiiiaioo ? iaoeaiiai aeiaoea a

na?aaeiaeu? ia e??i?ac?noaioi?noue nia?i??a aoaay, aa?aia ? eio?a

Aoaie ia?iaee ?ooeea?noue nia?i??a, aaeeR-eiao?oe?aio

e??i?ac?noaioiino?nia?iena?aaeiaeua c aeiaoeiina?aaeiaeua c aioeoi-

eiaei eiiiiiaioiina?aaeiaeua c aeiaoeiina?aaeiaeua c aioeoieiaeiaei
oieoi?ney oieoaei oieoi?ney oieoeiiiiiaioiiNia?ia aoaa?a (12)?icaaa-

eaiiy

7,1±0,18

3,95±0,22

6,8±0,25

3,95±0,14

0,55±0,1*

0,58±0,24*A?aeie-aaiiy

5,9±0,18

3,4±0,29

5,8±0,20

3,1±0,28

0,57±0,04*

0,55±0,04*Nia?ia aa?ai?a (10)?icaaa-eaiiy

6,3±0,11

2,4±0,15

6,4±0,13

2,3±0,13

0,38±0,02*

0,35±0,02*A?aeie-aaiiy

6,2±0,08

2,3±0,15

6,2±0,08

2,2±0,11

0,37±0,03*

0,35±0,03*Nia?ia eio??a (15)?icaaa-eaiiy

6,5±0,34

2,6±0,3

6,6±0,34

2,7±0,3

0,40±0,05*

0,40±0,03*A?aeie-aaiiy

6,2±0,4

2,6±0,4

6,1±0,6

2,8±0,4

0,41±0,04*

0,45±0,03** A??ia?aei?noue ??cieoe? (? > 0,5).

Caoenia ae?y aeno?aeoo iiynith?oueny ooai?aiiyi ia iiaa?oi? iaia?aie
nia?i?y e?iii?ioa?iiaiai i?ioa?eo, caiia?aath/iai iioeiaeaeoth/?e ae??
oieiaeo. Caa?aaeaiiy oeueiai i?ioa?eo i?ney aaaaoi?aciaiai a?aeieaaiiy
na?ae/eoue i?i i?oeiee o?i?/iee ca’ycie aioeoieiaiai eiiiiiaioo c
iaia?aiaie ee?oeie, ui ocaiaeaeo?oueny c aeine?aeaeaiiyie O.?.Inoaoei,
I.I.Iaaeaiei, 1976; I.I.Iaaeaiei, 1981 ? aenia?eiaioaeueii
i?aeoaa?aeaeaii J.Fulkes, 1978, yeee oaeiae niinoa??aaa i?oeiee ca’ycie
e?i?ae?a aeiaoea, i?/aieo ?aae?iaeoeaiei aoaeaoeai N17 c ieaciaoe/ieie
iaia?aiaie nia?i??a, iacaaaoe oeae ca’ycie o?i?/iei.

Oaeei /eiii, iaoeie aeine?aeaeaiiyie aia?oa aecia/aii ??aaiue caoenoo
noaoaaeo ee?oei eiiiiiaioaie aeiaoea a?ae oaiia?aoo?iiai oieo ?
i?aeoaa?aeaeaia iiaeaeue (O.?.Inoaoei, 1963) caoenii? ae?? aeiaoea ia
aeea? ee?oeie. Io?eiai? ?acoeueoaoe a?aee?eaaee ia?niaeoeao noai?aiiy
ooo/ieo na?aaeiaeu aeey ??cieo aeae?a oaa?ei c e?aueie nai?oa?ieie,
a?ieia?/ieie, aeieia?/ieie ? oaoi?/ieie iieacieeaie.

Cia/iee ?ioa?an i?aaenoaaey? ieoaiiy aea/aiiy iiaeeeaino? cai?ie
iaoeaiiai aeiaoea a ooo/ieo na?aaeiaeuao eiai eiiiiiaioaie i?e aeeaieiio
caii?iaeoaaii? nia?ie aoaa?a.

Aeey i?iaaaeaiiy aeine?ae?a iaie aoee io?eiai? ? aeei?enoai? aeey
aeine?aeaeaiue nie?oiaee, ao??iee, nie?o-ao??iee ? aoeaoiiiaee aeno?aeoe
aeiaoea. A ?acoeueoao? i?iaaaeaieo aeine?aeaeaiue onoaiiaeaii, ui
iaea?eueo aenieo aioeoieiao ae?th i?e e??ieiina?aaoe?? nia?ie
i?iyaeythoue ao??iee ? nie?oiaee aeno?aeoe. ?ooeea?noue nia?ie a
ao??iiio aeno?aeo? i?ney ?icii?iaeoaaiiy noaiiaeee 4,3 aae?a ?
aeaeeaai?noue (Sa) 22,4 iaeeieoe?, a nie?oiaiio aeno?aeo? oe? iieacieee
noaiiaeee 4,2 ? 21,2, a?aeiia?aeii. Aenieia??ia?aeia cieaeaiiy
?ooeeaino? nia?i??a a ii??aiyii? c eiio?ieai (? < 0,001) i?ney e??ieiina?aaoe?? a?aei?/aii a aoeaoiiiaiio aeno?aeo?. I?e aecia/aii? iioeiaeueii? eiioeaio?aoe?? aeno?aeoo aeiaoea a ooo/iiio na?aaeiaeu? nia?io, ?icaaaeaio ?icaaaethaa/aie aei?iaoaaee ia no?ee?noue aei oaiia?aoo?iiai oieo ? ia aeaeeaai?noue ?? i?e oaiia?aoo?? 0...+5°N. Iaea?aeai? ?acoeueoaoe na?ae/aoue, ui a aeaieo oiiaao iioeiaeueia eiioeaio?aoe?y aeno?aeoo a na?aaeiaeu? noaiiaeea 3%. I?e oeueiio eiao?oe?aio ?acenoaioiino? noaoaaeo ee?oei aei oaiia?aoo?iiai oieo aoa ia iaeiiio ??ai? c eiio?ieai R = 0,76. C iaoith oaeineiiaeaiiy niiniao io?eiaiiy aioeoieiaiai eiiiiiaioo aea/aee aieea ?oei?aii? ae?? iecueei? oaiia?aoo?e ia e?euee?ni? oa ye?ni? iieacieee aeno?aeoo, aeeo/aiiiai c aeiaoea i?ney ia?iaee eiai ??aeeei acioii. Nooi?iue ?oei?aii? ae?? ia no?oeoo?o aeiaoeiaeo eoeueie iaoeaiiai aeiaoea anoaiiaethaaee i?e?ioioia?aooaaiiyi noniaic?? aeiaoea aei ? i?ney e??iia?iaee. Anoaiiaeaii, ui i?ney aeeaieiai caii?iaeoaaiiy ? iine?aeoth/iai a?aeoaiaiiy aeine?aeieo c?ace?a aeiaoea niinoa??aathoueny aeano?oeoeai? ci?ie aeiaoeiaeo eoeueie ? aeo?ae c ieo e?iii?ioa?i?a o aeaeyae? ie?aieo ino??ae?a. Io?eiaiee aoaeo aaco?oueny ia aioo?eee?oeii?e e?enoae?caoe?? aiaee ? iaoai?/iiio iioeiaeaeaii? e?enoaeaie ieaciaoe/ieo iaia?ai. ?acoeueoaoe aea/aiiy eiiiiiaioo iieacai? a oaaeeoe? 2, ye? na?ae/aoue i?i oa, ui aeo?ae aioeoieiaiai eiiiiiaioo a aeine?ae? noaiiaea 16,4%, i?ioe 11% a eiio?ie?. I?e ii??aiyii? e?euee?nieo iieaciee?a aeine?aeieo ? eiio?ieueieo i?ia, ??cieoey i?ae ieie ia ei?enoue aeine?aeo aoea aenieia??ia?aeiith (? < 0,001) ? aoea aeuith ca iannith a 1,5 ?ace, i?e iiaiiio caa?aaeaii? ye?nieo aeanoeainoae. Oaeei /eiii, anoaiiaeaii, ui iiia?aaeiy e??iia?iaea iaoeaiiai aeiaoea caii?iaeoaaiiyi aei -196°N i?ecaiaeeoue aei aeano?oeoe?? aeiaoeiaeo eoeueie ? aeoiaeo c ieo e?iii?ioa?iiaeo niieoe, ui aeaei ciiao ?ic?iaeoe niin?a ca?eueoaiiy aeoiaeo aioeoieiaiai eiiiiiaioo i?e aeno?aaoaaii? c aeiaoea. Oaaeeoey 2. Aieea oiia e??iia?iaee aeiaoea ia aeo?ae aioeoieiaiai eiiiiiaioo Aeaee ia?iaie aeiao- ea ia?aae aeno?aao- aaiiyiAeo?ae aeno?aeoo, % Iiia?e aeine?ae?a 1 2 3 4 5 6 7A na- ?aaei., %Iaoeaiee aeiaoie (eiio?ieue) 8,8 12 12,2 12,2 13,2 8,8 10,2 11,0Caii?iaeaiiee aeiaoie 13,2 18,2 15,0 19,8 19,8 14,6 15,0 16,4??cieoe?, ?++++++0,001 Aaaeeeaei aoaiii aeine?aeaeaiue aoa iiooe iioeiaeueieo ?aaeei?a noa?ee?caoe?? na?aaeiaeua, yea i?noeoue aioeoieiaee eiiiiiaio ? aea/aiiy eiai aoaeoeaiino? i?ney aeiaaino?ieiaiai caa??aaiiy. Aeine?aeaeoaaee iiaeeea?noue oa?i?/ii? noa?ee?caoe?? na?aaeiaeu oeyoii eei’yo?iiy oa aaoieeaaoaaiiy. A aeine?aei? c?acee aiineee iiaeaeueio i?e?ioei?o (Bac.Subtilis). I?inoa?ee?ciaai? na?aaeiaeua ia?aa??yee ia aaeoa??aeueio caa?oaeiai?noue oa ?? aieea ia a?ieia?/i? iieacieee nia?ie i?ney e??ieiina?aaoe?? - a?aeoaiaiiy. ?acoeueoaoe aeine?aeo i?aaenoaaeai? a oaaeeoe? 3. Anoaiiaeaii, ui noa?ee?caoe?y eei’yo?iiyi (10...30 oa.), aaoieeaaoaaiiyi i?e 0,5 aoi. 20 oa. caaacia/othoue iiaio noa?eeuei?noue na?aaeiaeua, a eiai e??icaoeni? aeanoeaino? i?ney oa?i?/ii? ia?iaee a ii??aiyii? c eiio?ieai ia cieceeenue a?aeiinii noaoaaeo ee?oei. Aia?oa aeiaaaeaii, ui i?e ae?? ia aioeoieiaee eiiiiiaio na?aaeiaeua oaiia?aoo?aie, ia?aaeuoth/eie +100°N, a?i i?iyaey? oa?iinoaa?euei?noue ? iiai?noth caa??aa? nai? iiia?aaei? e??icaoeni? aeanoeaino?, ui aeaei ciiao aeei?enoaoe aoaeoeai? oa?i?/i? niiniae noa?ee?caoe?? na?aaeiaeu aac iia??oaiiy ?o o?ceei-o?i?/ieo ? nai?oa?ii-a?ieia?/ieo oa?aeoa?enoee. Aeine?aeio ia?o?th i?iaaoieeaaiaaiiai na?aaeiaeua caeeoaee i?e e?iiaoi?e oaiia?aoo?? ? aei?iaoaaee aieea no?ie?a caa??aaiiy na?aaeiaeua ia eiai nai?oa?i? ? e??icaoeni? aeanoeaino? i?e caii?iaeoaaii? nia?ie aoaa?a. ?acoeueoaoe aeine?aeaeaiue na?ae/aoue i?i oa, ui caa??aaiiy i?inoa?ee?ciaaiiai na?aaeiaeua ia i?ioyc? 2 ?ie?a, caaacia/eei iaae?eio eiai noa?eeuei?noue ? caa?aaeaiiy e??ii?ioaeoeaieo aeanoeainoae - ?ooeea?noue nia?ie i?ney ?icii?iaeoaaiiy noaiiaeea 4,1 aaee, aeaeeaai?noue a aiaeeiao - 9,3; iieaciee aaniethoii? aeaeeaaiino? - 18,3, ui ia a?ae??ciyeinue a?ae aeei?enoaiiy aeiaoieoo?eiath/iai na?aaeiaeua, aeaioiaeaiiai ex tempore. Oaaeeoey 3 Aieea ?aaeei?a noa?ee?caoe?? na?aaeiaeua ia caii?iaeaii-a?aeoaiai? nia?i?? (n=14) ?aaeei noa?ee?caoe?? (oa.)Iieacieee yeino? nia?ieEiio?ieueEei’yo?iiyAaoieeaao-aaiiy 0,5 aoi.10203020?ooeea?noue i?ney io?eiaiiy (aaee), I±m 7,8±0,17 7,8±0,17 7,8±0,17 7,8±0,17 7,8±0,17?ooeea?noue i?e aeaeiina?aaoe??, (aaee), M±m 4,1±0,14 4,0±0,13 4,1±0,15 4,1±0,12 4,1±0,14Aeaeeaai?noue a aiaeeiao i?e +38°N, M±m 8,5±0,5 8,1±0,6 8,4±0,8 8,6±0,8 8,6±0,7Iieaciee aaniethoii? aeaeeaaiino? i?e +38°N, M±m 22,1±1,4 21,7±1,8 22,1±2,1 23,5±1,7 23,5±1,6Aeinoia??i?noue ??cieoe?, ? >0,5

>0,5

>0,5

>0,5Aaeoa??ieia?/i?

aeine?aeaeaiiynooe?eueiee

??no

noa?.

noa?.

noa?.

noa?.

Ianooiiei aoaiii aeine?aeaeaiue aoei aecia/aiiy caie?aeithth/i?
caeaoiino? nia?ie, caeiina?aiaaii? a aeiaaicaa?aaeiiio ooo/iiio
na?aaeiaeu?. Iaoeiaee aeine?ae i?iaaaeaii a eieaini? OOO Ia?oc’?caeo
Aaeeei-Ao?eoenueeiai ?aeiio ia 140 aieiaao aaeeei? ?iaaoi? ooaeiae. Ca
aeaieie ?aeoaeueieo aeine?aeaeaiue ? ca ?icoaeaiiyie anoaiiaeaii, ui
caie?aeiai?noue i?ney ia?oiai in?iai?iiy a aeine?ae? noaiiaeea 71,2%, a
a eiio?ie? 69,2, ui aoei aeua ia ei?enoue aeine?aeo ia 2%. Oe?
?acoeueoaoe na?ae/aoue i?i oa, ui aeei?enoaiiy caa?aaeaiiai i?e
e?iiaoi?e oaiia?aoo?? i?ioyaii 2 ?ie?a na?aaeiaeua aeey e??ieiina?aaoe??
nia?ie caaacia/eei caie?aeithth/o caeaoi?noue naieoeue ia iaeiiio ??ai?
c? nia?iith, caeiina?aiaaiith a noaiaea?oiiio na?aaeiaeu?, ae?iaeaiiio
ia?aae ?icaaaeaiiyi nia?ie. Iaoeiai-ae?iaie/ee aeine?ae i?iaaaeaii a
eieainiao “?in?y” Iiai-Ieeiea?anueeiai ?aeiio Caii??aenueei? iaeano?;
“Eai?inueea ?ne?a” Iooe?nueeiai ?aeiio Noinueei? iaeano? ? aeine?aeiiio
ainiiaea?noa? “Oe?a?iea” ?inoeoooo oaa?eiieoeoaa OAAI, aea aoei
in?iaiaii 3299 ei??a ? oaeeoeue nia?iith ?icae?eaieo ayeoeyo?a. C
in?iaiaieo 1816 naieoeue aeine?aeiith nia?iith i?ioeaio caie?aeiaiino?
i?ney ia?oiai in?iai?iiy neeaaeaa a na?aaeiueiio 62,2%, ui aoei aeua a
ii??aiyii? c eiio?ieai ia 2% ? ia a?ae??ciyeinue a?ae a?aeiia?aeieo
iieaciee?a, io?eiaieo a ii??aiyeueiiio iaoeiaiio aeine?ae?.

Iaea?aeai? iiceoeai? ?acoeueoaoe aoaeoeaiino? aeei?enoaiiy
oa?iinoaa?eueiiai aeiaaicaa?aaeiiai na?aaeiaeua aeathoue i?aenoaao aeey
i?aai?caoe?? oeaio?ae?ciaaiiai eiai ae?iaieoeoaa ia i?aei?e?inoaao
oa?iaeieia?/ii? ? a?ieia?/ii? i?iieneiaino?. Iaeiae, oea iia’ycaii c
aeiaeaoeiaeie eai?oaeiaeeaaeaiiyie. Oiio, ia ia??iae ae??oaiiy oeueiai
ieoaiiy aeieeea iaiao?aei?noue iiooeo ? i?aeoe/iiai canoinoaaiiy
i?inoeo, aoaeoeaieo ? aeaoaaeo niinia?a noa?ee?caoe??
aeiaoieoo?eiath/eo na?aaeiaeu. Ia iaoo aeoieo, ca nai?ie
o?ceei-o?i?/ieie aeanoeainoyie aeey noa?ee?caoe?? na?aaeiaeu, iaea?eueo
aoaeoeaiei aioenaioeeii iiaea aooe ia?aeen aiaeith.

Aeey aea/aiiy ae?? aioenaioeea i?iaaee aeine?aeaeaiiy ii aecia/aiith
nooiaiy oiene/iino? ??cieo eiioeaio?aoe?e 33%-iiai ia?aeeno aiaeith aei
nia?ie aoaa?a ? ??aiy eiai aaeoa?eoeeaeii? ae?? a neeaae? na?aaeiaeua c
aeiaoeii. Onoaiiaeaii, ui ia?aeen aiaeith a eiioeaio?aoe?? 0,02…0,03%
ia aieeaa? oiene/ii ia noaoaa? ee?oeie. I?e eiioeaio?aoe?? 0,02%
a?aei?/aii noeioethth/o ae?th i?iaeoeo?a ?iciaaeo I2I2 ia nia?i??. I?e
oeueiio na?aaeiy aeaeeaai?noue noaoaaeo ee?oei aoea aeua a ii??aiyii? c
eiio?ieai ia 2 aiaeeie, iieaciee aaniethoii? aeaeeaaiino? ia 4,6
iaeeieoeue (59,7 a aeine?ae? i?ioe 55,1 a eiio?ie?), a aaeoa?eoeeaeia
ae?y aeine?aeiiai aioenaioeea a na?aaeiaeu? i?iyaeyeanue aaea i?e 0,03%.

Onoaiiaeaii, ui i?e caii?iaeaii? nia?ie a na?aaeiaeu? c ai?noii
aioenaioeea a eiioeaio?aoe?? 0,02…0,03% ?ooeea?noue nia?i??a i?ney
aeaeiina?aaoe?? neeaea 4,8 i?ioe 4,4 aae?a, a eiio?ie?. Oaea ae
caeaaei?noue aoea a?aei?/aia ? ca iieacieeii ?? aaniethoii? aeaeeaaiino?
22,3 i?ioe 20,3%. Ui-aei aaeoa?eoeeaeiiai aieeao I2I2 ia i?e?ioei?o
i/aaeaeii, ui i?e ?? eiioeaio?aoe?? 0,02% a na?aaeiaeu? cao?eniaaia
iayai?noue (8-10 i?e?ii?aai?ci?a a 1 ie), a ii/eiath/e c 0,03%
eiioeaio?aoe?? ? aeua an? c?acee aoee noa?eeueieie, a a eiio?ie?
niinoa??aaany nooe?eueiee ??no i?e?ioei?e.

?acii c oeei a?aei?/aii, ui ia?aeen aiaeith a aeicao, ia?aaeuoth/eo
0,03% ii?yae c aaeoa?eoeeaeiith ae??th i?iyaey? oiene/io ae?th ia
nia?i??. Eiia??ii oea iiyith?oueny oei, ui i?e i?aeaeuaiieo
eiioeaio?aoe?yo ia?aeeno aiaeith ia an? aoiie eenith anoeaathoue
ia’?aeiaoenue a iieaeoee ? aiie ae?thoue ia noaoaa? ee?oeie a oae?e aea
i??? caoaii, ye ? ia i?e?ii?aai?cie ca ?aooiie aoiia?iiai eenith.

Aeey aea/aiiy n?ia?a?aeii? ae?? I2I2 ni?eueii c oa?iiia?iaeith, a
oaeiae aecia/aiiy i?i?iaeueii? eiioeaio?aoe?? aaeoa?eoeeaeii? o?i?/ii?
?a/iaeie i?iaaee niaoe?aeuei? aeine?aeaeaiiy,?acoeueoaoe yeeo iieacai? a
oaaeeoe? 4. Aeai? oaaeeoe? aeacothoue ia oa, ui aeei?enoaiee
n?ia?a?aeiee niin?a noa?ee?caoe?? caaacia/ea noa?eeuei?noue na?aaeiaeua
i?e aeicao aioenaioeea 0,1…0,01% ia 100 ie. Oaeee ?aaeei noa?ee?caoe??
aeicaieea cieceoe noa?ee?coth/o aeico I2I2 a 10 ?ac?a ? oieeiooe
oiene/iino? aioenaioeea.

Oaaeeoey 4

Aecia/aiiy i?i?iaeueii? noa?ee?coth/i? aeice I2I2

ni?eueii c iaa??aii i?e +65° N/40 oa.

IieacieeeEiio-

?ieueE?euee?noue I2I2 (ie) a na?aaeiaeu? 0,1
1,090,080,070,060,050,040,030,020,01??aaiue aae-

oa??aeueii?

caa?oaeia-iino?

106

106

106

106

106

106

106

106

106

106

106?acoeueoaoe

aaeoa??iei-

a?/ieo

aeine?aeaeaiue

++++

nooe.

??no

noa?

noa?

noa?

noa?

noa?

noa?

noa?

noa?

noa?

noa?

C iaoith io?eiaiiy iaeneiaeueiiai naioth/iai aoaeoo a?ae n?ia?a?aeiiai
niiniao noa?ee?caoe?? aecia/aee aeiionoeio aa?oith iaaeo iaa??ao
na?aaeiaeua c aeiaoeii.

A ?acoeueoao? aeine?aeaeaiue onoaiiaeaii, ui iioeiaeueiei ?aaeeiii
i?ia??ao na?aaeiaeua c aeiaoeii aoee oaiia?aoo?e +65…75°N i?e
aeniiceoe?? 40 — 20 oaeeei. Aneiiceoe?y aeine?aeieo c?ace?a i?e
oaiia?aoo?? + 60°N ia aa?aiooaaea naiaoe?? na?aaeiaeua, a i?aeaeuaiiy
oaiia?aoo?e aei + 80°N i?ecaiaeeei aei aeaiaoo?aoe?? aeiaoea.

Aiae?coth/e ?acoeueoaoe aeine?ae?a ii ?ic?iaoe? o?i?ei-oa?i?/iiai
niiniao noa?ee?caoe?? ia iniia? aeei?enoaiiy ia?aeeno aiaeith c
aeiciaaiei iaa??aii ae?eoee aeniiaeo, ui a?i caaacia/o? noa?ee?caoe?th ?
iiaea aooe canoiniaaiee aeey naiaoe?? a?ieia?/ieo na?aaeiaeu c ai?noii
oa?iieaa?eueieo eiiiiiaio?a. Aeae? aea/aee aieea aeiaaicaa?aaeiiai
na?aaeiaeua i?ney o?i?ei-oa?i?/ii? noa?ee?caoe?? ia ye?ni? iieacieee
nia?ie i?e e??ieiina?aoaaii?. Ca eiio?ieue a?aee na?aaeiaeua NAeN-1
(noa?eeueia aeiaaicaa?aaeia na?aaeiaeua) noa?ee?ciaaia ?aae?aoe?eiei
iaoiaeii. Ii??aiyeuei? ?acoeueoaoe aeine?aeaeaiue i?aaenoaaeai? a
oaaeeoe? 5

Aeai? oaaeeoe? na?ae/aoue, ui o?i?ei-oa?i?/iee niin?a noa?ee?caoe??
na?aaeiaeu ia aieeaa? iaaaoeaii ia noaoaa? ee?oeie a i?ioean? ?o
caii?iaeoaaiiy. Onoaiiaeaii c a??ia?aei?noth (? < 0,001), ui cai?iiiiiaaiee niin?a aeaioiaeaiiy na?aaeiaeua ni?ey? iie?auaiith a?ieia?/ii? aeoeaiino? nia?i??a i?e eiina?aoaaii? a ii??aiyii? c na?aaeiaeuai i?inoa?ee?ciaaiei i?enei?aieie aeaeo?iiaie, i?i Oaaeeoey 5 Aieea niiniao noa?ee?caoe?? ia caii?iaeaii- a?aeoayi? nia?i?? (n=10) Iieacieee nia?i??a?aae?aoe?eiee (eiio?ieue) O?i?ei-oa?- i?/ieeA??ia?aei?noue, ??ooeea?noue i?ney ?icaaaeaiiy, aaee 7,6±0,01 7,6±0,01 >0,5?ooeea?noue i?ney caii?i-

aeoaaiiy-a?aeoaiaiiy,aaee

4,0±0,01

4,2±0,02

>0,5Aeaeeaai?noue nia?i??a,

aiaeeie

7,0±0,05

9,0±0,05

<0,05Iieaciee aaniethoii? aeaeeaaiino?, Sa 15,5±0,02 23,9±0,15 <0,001 ui na?ae/eoue iieaciee aaniethoii? aeaeeaaiino? nia?ie - 23,9 iaeeieoeue a aeine?ae?, i?ioe 15,5 iaeeieoeue a eiio?ie?. Io?eiai? aeai? aeaee i?aenoaao aeey aea/aiiy aieeao no?ie?a caa??aaiiy na?aaeiaeua c aeiaoeii i?ney o?i?ei-oa?i?/ii? noa?ee?caoe?? ia eiai aaeoa??ieia?/i? ? a?ieia?/i? iieacieee. Ia?aa??eo na?aaeiaeu aeine?aeaeoaaee c?aco i?ney ?o aeaioiaeaiiy, a iio?i /a?ac 1, 6, 12 ? 36 i?nyoe?a caa??aaiiy a oiiaao e?iiaoe. Eiio?ieai aoei na?aaeiaeua, aeaioiaeaia ex tempore. ?acoeueoaoe aeine?aeaeaiue i?aaenoaaeai? a oaaeeoe? 6. C oaaeeoe? aeaeii, ui an? aeine?aei? c?acee na?aaeiaeua, ye? caa??aaeenue a oiiaao e?iiaoe ia i?ioyc? aenia?eiaioo aoee noa?eeuei?, ui i?aeoaa?aeaeaii Oa?e?anueeith iaoeiai-ae?iaie/ith eaai?aoi???th, aenia?oece ?? 2805, 1807. Cai?iiiiiaaiee neeaae na?aaeiaeua ? niin?a eiai aeaioiaeaiiy caaacia/o? e??icaoeno nia?i???a i?ioyaii an?o aei?iaoaaieo no?ie?a caa??aaiiy (36 i?nyoe?a). Io?eiaii a??ia?aeia iie?auaiiy yeino? nia?ie i?ney caii?iaeaiiy a ii??aiyii? c eiio?ieai ca an?ia iieacieeaie aeine?aeo. C iaoith aecia/aiiy inoaoi/ii? a?ieia?/ii? iiaiioe?iiino? noaoaaeo ee?oei, caeiina?aiaaieo a aeine?aeaeoaaieo aeiaaicaa?aaeieo na?aaeiaeuao aoea aea/aia ?o caie?aeithth/a caeaoi?noue. In?iai?iiy i?iaiaeeee a ?aaeainiao “Ia?o?anueeee” *oao?anueeiai ?aeiio ? “*aia?aa” Oa?e?anueeiai ?aeiio. ?acoeueoaoe aeine?aeo na?ae/aoue, ui aeei?enoaiiy nia?ie, caeiina?aiaaii? a na?aaeiaeu? i?ney o?i?ei-oa?i?/ii? noa?ee?caoe?? aeaei iiceoeaiee aoaeo. Caie?aeiai?noue i?ney ia?oiai in?iai?iiy a aeine?ae? neeaea 74,0%, i?ioe 70,0% a eiio?ie?. Oea na?ae/eoue i?i aenieee ??aaiue caie?aeithth/i? caeaoiino? nia?ie, caeiina?aiaaii? a aeiaaicaa?aaeiiio na?aaeiaeu?, yea i?noeoue aeiaoie. Oaaeeoey 6 Aieea no?ie?a caa??aaiiy na?aaeiaeua a oiiaao e?iiaoe ia nai?oa?i? oa a?ieia?/i? iieacieee nia?ie (n=17) I ? n y oe ?Iieacieee0 16 12 36nia?ieEiio- ?ieueAein- e?aeEiio- ?ieueAein- e?aeEiio- ?ieue Aein- e?aeEiio- ?ieueAein- e?aeEiio- ?ieueAein- e?ae?ooeea?noue i?e io?eia- ii?, aaee 7,9 ±0,2 7,9 ±0,2 7,9 ±0,2 7,9 ±0,2 7,9 ±0,2 7,9 ±0,2 7,7 ±0,2 7,7 ±0,2 7,8 ±0,2 7,8 ±0,2?ooeea?noue i?ney caii?iaeoaaiiy-a?aeoaiaiiy, aaee 4,4 ±0,1 4,8 ±0,1 4,0 ±0,1 4,2 ±0,1 4,0 ±0,1 4,4 ±0,1 4,1 ±0,1 4,8 ±0,1 4,2 ±0,1 4,3 ±0,1Iieaciee aaniethoii? aeaeeaaiin- o?, Sa 20,3 ±1,5 21,7 ±1,3 17,5 ±1,3 19,1 ±1,2 22,9 ±1,5 23,9 ±1,5 23,7 ±1,0 24,4 ±1,1 22,9 ±1,2 23,4 ±1,2Aeaeeaai?noue a aiaeeiao9,0 ±0,59,4 ±0,97,3 ±0,27,7 ±0,38,5 ±0,49,0 ±0,59,0 ±0,59,0 ±0,58,7 ±0,49,0 ±0,5?acoeueoaoe aaeoa??ieia. aeine?aeaeaiue - noa?. - noa? - noa?. - noa?. - noa?. - noa?. - noa?. - noa?. - noa?. - noa?. ?ic?iaea aeiaaicaa?aaeieo na?aaeiaeu aeey e??ieiina?aaoe?? nia?ie aeaea ciiao onoiooe na?eici?oa aeaea?aei caa?oaeiaiiy nia?ie i?e?ii?aai?ciaie, ui aeaei iiaeeea?noue caaa?oeoe eiiieaen anaioe/ieo cania?a i?e e??ieiina?aaoe?? nia?ie ca Oa?e?anueeith oaoiieia??th. A i?ioean? aea/aiiy aeiaaicaa?aaeieo na?aaeiaeu aoei aeyaeaii ua iaeio c ?o ia?aaaa - i?ici??noue. A ia?oiio aa??aio? oea aeinyaa?oueny ca ?aooiie iiaiiai aeaeaeaiiy aeiaoeiaeo eoeueie i?e aeno?aaoaaii? aioeoieiaiai eiiiiiaioo, a ae?oaiio - ca ?aooiie ?o niiioaiii? naaeeiaioaoe?? i?e caa??aaii? na?aaeiaeua a aiioeao. Caaaeyee i?ici?ino? c’yaeeanue iiaeeea?noue ?ic?iaee anaioe/iiai niiniao oa i?eno?ith aeey ioe?iee yeino? nia?ie, aaciina?aaeiuei a aa?iaoeciaai?e nia?iiaeic?, ui ia?aaeaa/a?oueny aeiiaaie Oa?e?anueei? oaoiieia?? ooo/iiai in?iai?iiy ei??a ? oaeeoeue. ?ic?iaeaiee niin?a i?aeaioiaee aeic aeey i?e?ineii?? ca ?aooiie ?iia?n?eii? nenoaie ? i?eno??e aeey i?e?ineii?? nia?ie a aeanoe/ieo oiaeiaeao caaacia/o? aeinyaiaiiy a??ia?aeieo ?acoeueoao?a eiio?ieth yeino? nia?ie a ii??aiyii? c caaaeueiii?eeiyoei niiniaii i?e?ineii?? ?icaeaaeaii? e?aie? nia?ie. Ia caaa?oaiiy i?iaaaeaii oe?iea ae?iaie/a aeine?aeaeaiiy ii canoinoaaiith aeiaaicaa?aaeiiai na?aaeiaeua, noa?ee?ciaaiiai o?i?ei-oa?i?/iei niiniaii ? aea/aia aoaeoeai?noue eiai aeei?enoaiiy i?e e??ieiina?aaoe?? nia?ie ia ieaii?aei?e?inoaao Oe?a?ie ? ?in??. Ia iniia? i?iaaaeaieo aeine?ae?a iaie ?ic?iaeai? oaoi?/i? oiiae ? ?aaeaiaio aeaioiaeaiiy aeiaaicaa?aaeiiai na?aaeiaeua, a oaeiae i?aai?ciaaii aenia?eiaioaeueiee a?ioeao ii aeaioiaeaiith na?aaeiaeu. I?ioyaii aeine?aeo aeaioiaeaii ? ?aae?ciaaii ieai?iiei i?aei?e?inoaai 1137 e?o??a noa?eeueiiai aeiaaicaa?aaeiiai na?aaeiaeua. Caeiina?aiaaii a aeine?aeiiio na?aaeiaeu? 14,4 iei. nia?iiaeic ? in?iaiaii 3,505 iei. ei??a ? oaeeoeue. Caie?aeiai?noue i?ney ia?oiai in?iai?iiy neeaea 61,8%. Ca ?acoeueoaoaie i?aea?aeii/eo aei?iaoaaiue na?aaeiaeua ? niin?a eiai aeaioiaeaiiy ?aeiiaiaeiaai? aeey i?iieneiaiai ae?iaieoeoaa. Canoinoaaiiy aeiaaicaa?aaeieo na?aaeiaeu a ii??aiyii? n aiaeiaaie ia? ne?aeoth/? ia?aaaae: iiaeeea?noue i?iieneiai aeaioiaeaiiy na?aaeiaeu ? oeaio?ae?ciaaiiai iinoa/aiiy ?o ieai?iiei i?aei?e?inoaai; aeeeth/a?oueny iiaeeea?noue caa?oaeiaiiy nia?ie ? noaoaaeo oeyo?a naieoeue i?e ?o in?iai?ii? i?e?ioei?ith na?aaeiaeu; caaacia/o?oueny ?aoe?iiaeueia aeei?enoaiiy o?i?aaeoea?a ? aeiaoea; aeinyaa?oueny i?ici??noue na?aaeiaeua, ui caaacia/o? ia’?eoeaio ioe?ieo ?ooeeaino? nia?i??a, ? aeaei ciiao ?ic?iaeoe anaioe/iee niin?a i?e?ineii?? nia?ie a aa?iaoeciaaieo nia?iiaeicao. AENIIAEE 1. Eiiiiiaioe aeiaoea eo?y/eo y?oeue aeno?aaiaai? i?aai?/ieie ?ic/eiieeaie ? aaaaeai? a nia?io a eiioeaio?aoe?? 3% caiia?aathoue oaiia?aoo?iiio oieo noaoaaeo ee?oei aoaay, aa?aia ? eio?a ia iaeiiio ??ai? c 30%-iith eiioeaio?aoe??th iaoeaiiai aeiaoea. 2. Anoaiiaeaii i?oeiee ca’ycie aioeoieiaiai eiiiiiaioo aeiaoea c oeeoiieaciaoe/ieie iaia?aiaie nia?i??a. Aaaaoi?aciaa oeaio?eooaeia eiai a?aeieaaiiy a?ae nia?i??a ?cioii?/iei aoaeaaiaeii-nieueiaei aooa?ii ia iicaaaey? ?acenoaioiino? noaoaaeo ee?oei aei oaiia?aoo?iiai oieo. 3. Anoaiiaeaii, ui iiia?aaeiy e??iia?iaea iaoeaiiai aeiaoea caii?iaeoaaiiyi aei -196°N i?ecaiaeeoue aei aeano?oeoe?? aeiaoeiaeo eoeueie ? aeoiaeo c ieo e?iii?ioa?iiaeo niieoe, ui aeaei ciiao ?ic?iaeoe niin?a ca?eueoaiiy aeoiaeo aioeoieiaiai eiiiiiaioo i?e aeno?aaoaaii? c aeiaoea a 1,5 ?ace. 4. I?ia??aaiiy ?ic/ei?a aioeoieiaiai eiiiiiaioo i?e +100...110°N ia aieeaa? iaaaoeaii ia eiai oa?iino?ee? aeanoeaino? ? ia ii?ooo? caoeni? yeino? a?aeiinii nia?ie, ui aea? ciiao canoinoaaoe aoaeoeai? niiniae eiai noa?ee?caoe??. 5. ?ic?iaeaii na?aaeiaeua aeey e??ieiina?aoaaiiy nia?ie aoaa?a (a.n. ? 745045), yea a?ae?yciy?oueny oei, ui c iaoith caa?aaeaiiy aeniei? nai?oa?ii? ? a?ieia?/ii? yeino? nia?ie, na?aaeiaeua i?noeoue cai?noue aeiaoea oa?iinoaa?eueiee aioeoieiaee eiiiiiaio a oaeiio ni?aa?aeiioaii? ?ia??ae??io?a: 11% ?ic/ei eaeoice 93,0, ae?oea??i ie 4,0, oa?iinoaa?eueiee aioeoieiaee eiiiiiaio 3.0. 6. ?ic?iaeaii niin?a aeaioiaeaiiy na?aaeiaeua c aioeoieiaei eiiiiiaioii, yeee a?ae??ciy?oueny oei, ui c iaoith aeiaaicaa?aaeaiiy ? oieeiaiiy i?e?iaiiai oaeoi?o, na?aaeiaeua aa?iaoecothoue a aiioee ? i?ia??aathoue i?e 100...110°N i?ioyaii 20 oaeeei, iio?i caa??aathoue a oiiaao e?iiaoe aei iiiaioo aeei?enoaiiy. 7. Anoaiiaeaii, ui 33%-iee ia?aeen aiaeith, aianaiee a oa?iieaa?eueia na?aaeiaeua a eiioeaio?aoe?? 0,02% noeioeth? ?ooeea?noue ? aeaeeaai?noue nia?i??a i?e +38°N, a oiene/ia eiai ae?y ianooia? c 0,07%-ii? eiioeaio?aoe??, ui aeicaieeei aia?oa canoinoaaoe oeae aioenaioee aeey naiaoe?? oa?iieaa?eueieo na?aaeiaeu. 8. Canoinoaaiiy iaoiene/ii? aeice I2I2 (ia?aeeno aiaeith) a na?aaeiaeu? ni?eueii c i?ia??aii eiai i?e +70°N ia i?ioyc? 20 oaeeei, caaacia/eei noa?eeuei?noue iiia?aaeiuei caa?oaeiaiiai i?e?ii?aai?ciaie na?aaeiaeua ? ia cieceei eiai caoenieo yeinoae a i?ioean? e??ieiina?aoaaiiy nia?i??a. ?ooeea?noue nia?i??a i?ney aeaeiina?aaoe?? neeaaeaea 4,8 aaea i?ioe 4,0 aae?a a eiio?ie?, a ii iieacieeo aaniethoii? aeaeeaaiino? 23,9 i?ioe 15,5 iaeeieoeue, a?aeiia?aeii. 9. Cai?iiiiiaaiee iaie neeaae ? niin?a aeaioiaeaiiy na?aaeiaeua, yea i?noeoue aeiaoie (iaoaio ?in?? ? 1828582 ? Iaoaio Oe?a?ie ? 13138) caaacia/o? caa?aaeaiiy ii/aoeiaeo e??ii?ioaeoeaieo ? nai?oa?ieo yeinoae eiai i?ioyaii 36 i?nyoe?a caa??aaiiy a oiiaao e?iiaoe, ui aeicaieeei i?aai?coaaoe ae?iaieoeoai ? oeaio?ae?ciaaia iinoa/aiiy e??ii?ioaeoeaiiai na?aaeiaeua ieai?iiei i?aei?e?inoaai. 10. Ia iniia? aeei?enoaiiy i?ici?eo aeiaaicaa?aaeieo na?aaeiaeu ?ic?iaeaii anaioe/iee niin?a ? i?eno??e (a.n. ? 1543598) aeey ioe?iee nia?ie ia?aae in?iai?iiyi naieoeue, aaciina?aaeiuei a aa?iaoeciaai?e nia?iiaeic?, ui caaacia/o? /?oeo aeaeei?noue nia?i??a i?ae i?e?ineiiii, a?eueo oi/io ioe?ieo ?o ?ooeeaino?. 11. I?iaaaeaii iaoeiaee ? oe?ieee ae?iaie/ee aeine?ae ai?iaaaeaeaiiy aeiaaicaa?aaeaiiai na?aaeiaeua ia 14 i?aei?e?inoaao Oe?a?ie ? ?in??, a yeiio aeaioiaeaii 1137 e?o??a na?aaeiaeua, caaioiaeaii 15,3 iei.nia?iiaeic ? in?iaiaii 3 iei.500 oen.ei??a ? oaeeoeue. Caie?aeiai?noue a?ae ia?oiai in?iai?iiy neeaaea? 61,8%. I?e oeueiio aeeo/a?oueny aeaea?aei iioe?aiiy caoaeiee?a oai?ia, ye? iiaeooue ia?aaeaaaoenue c aeiaoeii i?e ooo/iiio in?iai?ii? ? nei?i/othoueny aeo?aoe aei?iaieiooiaieo eiiiiiaio?a ? ?aaeoea?a. ?AEIIAIAeAOe?? AE?IAIEOeOAO 1. Aeey i?aeaeuaiiy nai?oa?ii-a?a??ie/iiai ??aiy ?iaioe ieai?iieo i?aei?e?inoa ? caiia?aaiiy caoaeiee?a caoai?thaaiue, ye? ?iciianthaeaeothoueny c aeiaoeii ? ?ioeie eiiiiiaioaie, a oaeiae aeey ?aoe?iiaeueiiai aeei?enoaiiy an?o eiiiiiaio?a na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie aoaa?a i?iiiio?oueny oa?icaaiaeai, a?ioaa?eeai ? ieaiia’?aeiaiiyi aeaa aa??aioe aeiaaicaa?aaeieo na?aaeiaeu: - na?aaeiaeua, yea i?noeoue oa?iinoaa?eueiee aioeoieiaee eiiiiiaio cai?noue iaoeaiiai aeiaoea ? eiai oa?i?/io noa?ee?caoe?th i?e 110°N i?ioyaii 20 oaeeei (a.n. N?N? ? 745045); - na?aaeiaeua, yea i?noeoue aeiaoie ? o?i?ei-oa?i?/iee niin?a noa?ee?caoe??, aeeth/ath/ee a na?aaeiaeua oa?iaeiiaeiee ia?aeen aiaeith (I2I2) a eiioeaio?aoe?? 0,02...0,03% ? i?ia??a eiai i?e +70°N i?ioyaii 20 oaeeei (iaoaio N?N? ? 1828582, iaoaio Oe?a?ie ? 13138). 2. C iaoith i?aeaeuaiiy nai?oa?ii-a?a??ie/iiai ??aiy oa ?aoe?iiaeueii? i?aoe? ioieo?a ooo/iiai in?iai?iiy oaa?ei i?iiiio?oueny iaooeiaeaeoth/ee anaioe/iee niin?a oa i?eno??e aeey ioe?iee ?ooeeaino? nia?i??a i?e i?e?ineii?? a aa?iaoe/ieo oiaeiaeao (a.n.N?N? ? 1543598). NIENIE IIOAE?EIAAIEO AAOI?II I?AOeUe CA OAIITH AeENA?OAOe?? 1. A.n. ?745045. N?aaea aeey e?eieiina?aaoeee nia?iu / O.E. Inoaoei, I.I.Iaaeaiei, E.I.Iaaeaiei (NNN?).- ? 2623106/30-15; Cayaeaii 1.06.78; Ca?aa. a A?E NNN? 7.03.80.a?eo:”Ia iiaeeaaeeo iioaeeeiaaieth a ioe?uoie ia/aoe”. 2. Ostashko F.I., Pavlenko M.P., Pavlenko L.N. Antishock effect of yolk and components in cooling spermatozoids of bull, ram and boar/10th International Congress on Animal Reproduction and A.I.- Illinois (USA).- 1984.- V.1.- P. 209-211. 3. Iaaeaiei E.I. E?eieiina?aaoeey nia?iu aueia a aeieaio?aiyuaeny oa?iinoaaeeueiie n?aaea //Ananithciay eiioa?aioeey e?eiaeieiaee e e?eiiaaeeoeeiu.- Oa?ueeia, 1984.- O.2.- N. 245. 4. Iaaeaiei I.I., Iaaeaiei E.I. Eco/aiea caueoiuo naienoa aioeoieiaiai eiiiiiaioa aeaeoea ia nia?ia aueia, aa?aiia e o?yeia i?e e?eiaicaeaenoaee //Ananithciay eiioa?aioeey e?eiaeieiaee e e?eiiaaeeoeeiu.- Oa?ueeia, 1984.- O.2.- N. 246. 5. A.n. 1543598 NNN?. Ono?ienoai aeey iee?ineiiee nia?iu e yia?eiiia a yeanoe/iuo eainoeao /O.E.Inoaoei, I.I.Iaaeaiei, E.I.Iaaeaiei, A.I.Eociaoeia.- ? 44528131/30-15; Caaeaii 14.04.88; Ca?aa. a A?E NNN? 15.10.89. 6. Inoaoei O.E., Iaaeaiei I.I., Iaaeaiei E.I. Aeieaio?aiyuayny n?aaea aeey nia?iu e aa aeeyiea ia auaeeaaaiinoue e iieiaeioai?ythuoth niiniaiinoue nia?ieaa aueia i?e e?eieiina?aaoeee //Iao/ii-i?iecaiaenoaaiiay eiioa?aioeey “Iiaua iaoiaeu e aeioaoiieiaey a aeeaioiiaiaenoaa”.- E.: Ainaa?ii?ii ONN?, 1991.- N.38-39. 7. Inoaoei O.E., Iaaeaiei I.I., Iaaeaiei E.I. AEeaeeea aeieaio?aiyueany n?aaeu aeey e?eieiina?aaoeee nia?iu aueia //Oacenu 11 iaaeaeoia?iaeiie eiioa?aioeee 21-25 ai?aey 1992 “Oniaoe a nia?aiaiiie aeieiaee” / Einoeooo e?eiaeieiaee e e?eiiaaeeoeeiu AI Oe?aeiu.- Oa?ueeia, 1992.- N.134-135. 8. Inoaoei O.?., Iaaeaiei I.I., Iaaeaiei E.I. Iaoiaeeea aecia/aiiy aioeoieiaeo aeanoeainoae caoenieo eiiiiiaio?a a ?icaaaethaa/ao i?e ae?? iecueeeo oaiia?aoo? ia nia?i?? // Iiaa a iaoiaeao ciioaoi?/ieo aeine?aeaeaiue /?inoeooo oaa?eiieoeoaa OAAI.- Oa?e?a, 1992.- N.138-142. 9. Iao. 1828582 NNN?. N?aaea aeey e?eieiina?aaoeee nia?iu e niinia aa i?eaioiaeaiey / O.E.Inoaoei, I.I.Iaaeaiei, E.I.Iaaeaiei (NNN?).- ? 4948632; Cayaeaii 8.04.91; Ca?aa. a A?E NNN? 13.10.92. a?eo. “Ia iiaeeaaeeo iioaeeeiaaieth a ioe?uoie ia/aoe”. 10. Inoaoei O.?., Iaaeaiei I.I., Iaaeaiei E.I. ??aeeee aeiaaicaa?aaeaiee ?icaaaethaa/ aeey e??ieiina?aaoe?? nia?ie ie?aeiee?a // Iniiai? iaoeia? ?ic?iaee ?inoeoooo aeey ai?iaaaeaeaiiy o ae?iaieoeoai / ?inoeooo oaa?eiieoeoaa OAAI.- Oa?e?a, 1992.- N.13-14. 11. Iaaeaiei I.I., Iaaeaiei E.I. Aeiaaicaa?aaeaia na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie aoaa?a-ie?aeiee?a // Iiei/ii-i’ynia neioa?noai: I?aea.oaiao.iaoe.ca??iee.- E.:O?iaeae, 1994.- Aei.85.- N.70-72. 12. Iao. 13138 Oe?a?ia, IEE A 61 D 19/02. Na?aaeiaeua aeey eiina?aoaaiiy nia?ie oa niin?a eiai i?eaiooaaiiy / O.?.Inoaoei, I.I.Iaaeaiei, E.I.Iaaeaiei (Oe?a?ia).- Iioae. 28.02.97.- Athe.? 1. Iaaeaiei E.I. Aeiaaicaa?aaeia na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie aoaa?a oa niiniae eiai aeaioiaeaiiy.- ?oeiien. Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe ca niaoe?aeuei?noth 03.00.20 - a?ioaoiieia?y.- ?inoeooo oaa?eiieoeoaa OAAI.- Oa?e?a, 1998 ?. C aeei?enoaiiyi no/anieo iaoiae?a aeine?aeaeaiue anoaiiaeaii caoeniee aoaeo oa?iinoaa?eueiiai aioeoieiaiai eiiiiiaioo aeiaoea i?e ae?? iecueeeo oaiia?aoo? ia nia?i??. A iiaeaeueieo aeine?aeaeaiiyo aeiaaaeaii ?aeaioe/i?noue eiai a?ieia?/ii? ae?? i?e e??ieiina?aaoe?? noaoaaeo ee?oei a ii??aiyii? c iaoeaiei aeiaoeii. Onoaiiaeaia iiaeeea?noue o?i?ei-oa?i?/ii? noa?ee?caoe?? na?aaeiaeua c aeiaoeii. Ia oe?e iniia? ?ic?iaeaii ? aei?iaoaaii a ae?iaie/eo oiiaao c iiceoeaieie ?acoeueoaoaie aeiaaicaa?aaeia na?aaeiaeua ? niiniae eiai aeaioiaeaiiy. Aiii i?aeaeuo? nai?oa?ii-a?a??ie/iee ??aaiue ?iaioe ieai?iieo i?aei?e?inoa, caiia?aa? iioe?aiith ca?acieo ? iaca?acieo caoai?thaaiue, ye? ?iciianthaeaeothoueny c aeiaoeii, a oaeiae caaacia/o? ?aoe?iiaeueia aeei?enoaiiy aeiaoea ? an?o eiiiiiaio?a i?e aeaioiaeaii? na?aaeiaeua aeey e??ieiina?aaoe?? nia?ie aoaa?a. ?ic?iaeaii anaioe/iee niin?a oa i?eno??e aeey ioe?iee ?ooeeaino? nia?i??a a aeanoe/ieo oiaeiaeao c iaoith i?aeaeuaiiy nai?oa?ii-a?a??ie/iiai ??aiy ?iaioe ioieo?a ooo/iiai in?iai?iiy. Eeth/ia? neiaa: nia?ia, aeiaoie, aioeoieiaee eiiiiiaio, e??i?ac?noaioi?noue, e??ieiina?aoaaiiy, oa?iino?ee?noue, noa?ee?caoe?y, ia?aeen aiaeith, ?ia?n?y, caie?aeiaiiy. Iaaeaiei E.I. Aeieaio?aiyuayny n?aaea aeey e?eieiina?aaoeee nia?iu aueia e niiniau aa ecaioiaeaiey. - ?oeiienue. Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe 03.00.20 - aeioaoiieiaey. - Einoeooo aeeaioiiaiaenoaa OAAI. - Oa?ueeia, 1998 a. Aeenna?oaoeey iinayuaia aii?inai eco/aiey aeeyiey aeaeoea e aai eiiiiiaioia ia nia?iee i?e eo aeiioa?iee e nicaeaiey ia yoie iniiaa noa?eeueiie aeieaio?aiyuaeny n?aaeu aeey e?eieiina?aaoeee nia?iu aueia-i?iecaiaeeoaeae. N eniieueciaaieai nia?aiaiiuo iaoiaeia enneaaeiaaiee onoaiiaeaii, /oi eiiiiiaiou aeaeoea eo?eiuo yeoe, yeno?aae?iaaiiua i?aaie/aneeie ?anoai?eoaeyie e aaaaeaiiua a n?aaeo a eiioeaio?aoeee 3%, i?aaeio?aiytho io oaiia?aoo?iiai oiea iieiaua eeaoee auea, aa?aia e o?yea ia iaeiii o?iaia n 30%-iie eiioeaio?aoeeae iaoeaiiai aeaeoea. Onoaiiaeaia i?i/iay naycue aioeoieiaiai eiiiiiaioa n oeeoiieaciaoe/aneeie iaia?aiaie nia?ieaa, i?iyaeythuayny nio?aiaieai e?ei?acenoaioiinoe iieiauo eeaoie iinea eo iiiaie?aoiiai oeaio?eooaeiiai ioiuaaiey ecioiie/aneeie ?anoai?aie. Auyaeaii, /oi i?e aicaeaenoaee ia aioeoieiaue eiiiiiaio a ?anoai?a oaiia?aoo?, i?aauoathueo + 100°N, ii i?iyaeyao oa?iinoieeinoue e nio?aiiinoue naieo ia?aiia/aeueiuo caueoiuo naienoa. A iiaeaeueiuo yenia?eiaioao aeieacaia eaeaioe/iinoue aai aeieiae/aneiai aicaeaenoaey a n?aaiaiee n iaoeaiui aeaeoeii i?e e?eieiina?aaoeee nia?iu aueia-i?iecaiaeeoaeae. Ia yoie iniiaa ?ac?aaioaia n?aaea, ioee/athuayny oai, /oi n oeaeueth nicaeaiey aunieiai naieoa?iiai e aeieiae/aneiai ea/anoaa nia?iu, iianiaea?aeeo oa?iinoaaeeueiue aioeoieiaue eiiiiiaio, aianoi aeaeoea, /oi iicaieeei i?eiaieoue yooaeoeaiue oa?ie/aneee niinia noa?eeecaoeee e i?iaeeeoue n?ie o?aiaiey n?aaeu a aiioea aei 2-o eao aianoi iaeieo nooie. I?aaeeiaeai niinia oaaee/aiey auoiaea aioeoieiaiai eiiiiiaioa ec aeaeoea a 1,5 ?aca ca n/ao i?aaeaa?eoaeueiie aai e?eiia?aaioee. Onoaiiaeaia aiciiaeiinoue oeieei-oa?ie/aneie noa?eeecaoeee aeaeoieniaea?aeauae n?aaeu. Aaaaeaiea iaoiene/iie aeicu ia?aeene aiaei?iaea a oa?iieaaeeueioth n?aaeo, a ni/aoaiee n aa i?ia?aaii i?e + 70°N a oa/aiea 20 ieioo, iaania/eaaao noa?eeueiinoue caaaaeiii ianaiaiaiiie iee?ii?aaieciaie n?aaeu ia i?ioyaeaiee 36 ianyoeaa o?aiaiey e ia nieaeaao aa caueoiuo naienoa a i?ioeanna e?eieiina?aaoeee nia?ieaa. Ia iniiaaiee eniieueciaaiey i?ic?a/iuo aeieaio?aiyueony n?aae ?ac?aaioai anaioe/aneee niinia e ono?ienoai aeey ioeaiee nia?iu iaiin?aaenoaaiii a aa?iaoece?iaaiiie nia?iiaeica, iicaieythuee oeo/oeoue ioeaieo nia?iu, nieceoue aa iioa?e, iiauneoue naieoa?ii-aeaeaie/aneee o?iaaiue eneonnoaaiiiai inaiaiaiey. Iniiaiua ?acoeueoaou ?aaiou iaoee oe?ieia i?eiaiaiea a i?aeoeea ieaiaiiuo i?aaei?eyoee i?e nicaeaiee aaieia aeoaieicaii?iaeaiiie nia?iu aueia-i?iecaiaeeoaeae. I?aaeeiaeaiiua aa?eaiou noa?eeueiuo aeieaio?aiyueony n?aae iaania/eaatho iiauoaiea naieoa?ii-aeaeaie/aneiai o?iaiy ?aaiou ieaiaiiuo i?aaei?eyoee, i?aaeoi?aaeaeaiea caaieaaaiee, ?ani?ino?aiyth-ueony n aeaeoeii e ae?oaeie eiiiiiaioaie, a oaeaea ?aoeeiiaeueiia eniieueciaaiea anao eiiiiiaioia ca n/ao oeaio?aeeciaaiiiai aa i?iecaiaenoaa. Eeth/aaua neiaa: nia?ia, aeaeoie, aioeoieiaue eiiiiiaio, e?ei?acenoaioiinoue, e?eieiina?aaoeey, oa?iionoie/eainoue, noa?eeecaoeey, ia?aeenue aiaei?iaea, eiia?ney, iieiaeioai?yaiinoue. Pavlenko L. N. Long-term storage medium for bull semen cryopreservation and methods of its production. - Manuscript. The dissertation for the scientific degree of the Candidate of Sciences (Agriculture) in speciality 03.00.20 - Biotechnology. - The Institute of Animal Science of the UAAS. - Kharkov, 1998. Using modern research techniques, the protective effect of the heat- stable antishock component of egg yolk at exposure of semen to low temperatures has been established. In simulation experiments the identity of its biological effect at cryopreservation of reproductive cells as compared with native egg yolk has been proved. Feasibility of chemical-thermal sterilization of yolk-containing medium has been established. On the above basis a long-term storage medium and methods of its production have been developed and tested in production conditions with positive results. It increases sanitary-hygienic level of breeding units, prevents spreading contagious and non-contagious diseases transmitted with egg yolk, as well as provides for efficient use of yolk and all the components at producing the medium for bull semen cryopreservation. An aseptic method and device for spermatozoa motility evaluation in elastic packages to increase sanitary-hygienic level and efficient operation of AI units has been developed. Key words: semen, yolk, antishock component, cryoresistance, cryopreservation, heat stability, sterilization, peroxide, immersion, pregnancy rate. A?aeiia?aeaeueiee ca aeione: aeieoi? a?ieia?/ieo iaoe Aoa?ia I.Ae. I?aeienaii aei ae?oeo 27.12.1998 ?. Oi?iao 60 o 84/16 Ianya 1.0 oi.-ae?oe.a?e., 1,0 iae.-aeae.a?e. Caiiaeaiiy ? 17 Oe?aae 100 i?ei. Ae?eyiea iia?aoeaiiai ae?oeo ?inoeoooo oaa?eiieoeoaa OAAI 312120, i.Oa?e?a, i/a Eoeeie/?

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