?INOEOOO AIOAI?EE ?i. I.A. OIEIAeIIAI

IAOe?IIAEUeII? AEAAeAI?? IAOE OE?A?IE

A A A E I

??eia Aiae???aia

OAeE 581.526.53: 581.9 (477.53+477.54)

AeEOA?AIOe?AOe?ss ?INEEIIIAI IIE?EAO NOAI?A I?AAeAIII? *ANOEIE
E?AIAA?AAEIIAI E?NINOAIO OE?A?IE

03.00.05 – aioai?ea

AAOI?AOA?AO

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Ee?a – 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia a ?inoeooo? aioai?ee ?i I.A. Oieiaeiiai IAI
Oe?a?ie.

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe

Ae?AeOO ssE?A IAO?IAE*

?inoeooo aioai?ee ?i. I.A. Oieiaeiiai IAI Oe?a?ie

caa. a?aeae?eii aeieia?? o?oinenoai

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani?

AO?AeA ?A?NA ?AAI?AIA

?inoeooo aa?iaeieia?? oa a?ioaoiieia?? OAAI

i?ia?aeiee iaoeiaee ni?a?ia?oiee

aeieoi? a?ieia?/ieo iaoe

NIEIIAOA AIEIAeEIE? AIAe??EIAE*

Ee?anueeee oi?aa?neoao ?iai? Oa?ana Oaa/aiea,

a?ieia?/iee oaeoeueoao, caa. eaoaae?e aioai?ee

I?ia?aeia onoaiiaa: Oeaio?aeueiee aioai?/iee naae ?i. I.I. A?eoea IAI
Oe?a?ie, a?aeae?e i?e?iaeii? oei?e i. Ee?a

Caoeno a?aeaoaeaoueny “29” aa?aciy 1999 ?. i 1000 aiaeei? ia
can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee Ae. 26.211.01?inoeoooo aioai?ee
?i. I.A. Oieiaeiiai IAI Oe?a?ie ca aae?anith: 252601 i. Ee?a INI-1,
aoe Oa?auaie?anueea, 2

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? ?inoeoooo aioai?ee
?i. I.A. Oieiaeiiai IAI Oe?a?ie ca aae?anith: 252025 i. Ee?a, aoe. A.
AEeoiie?nueea, 28

Aaoi?aoa?ao ?ic?neaiee “ 24 ” _ethoiai_____ 1999 ?.

O/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee
?eue?inueea A.I.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. Coaiia? aeinenoaie ia?aaoaathoue o noai? iano?eei?
??aiiaaae ? ? aeoaea aeeiai?/ieie. ?ai?oa noaie caeiaee cia/i? i?inoi?e
(aeecueei 40% oa?eoi??? Oe?a?ie) ? a?ae?a?aaaee i?ia?aeio ?ieue o
oi?ioaaii? oeaiioe/ii? niaoeeo?ee eaiaeoaoo?a oa ooai?aii? iiooaeieo
/i?iicai?a. Oiane?aeie iaaei??iiai ?ici?thaaiiy oa?eoi??e, yea a ie?aieo
i?aaeaiieo ?aa?iiao ia?aaeuo? 85%, noaie aoee cieuai? ye ciiaeueiee oei
?ineeiiino? ? ca?ac ?niothoue eeoa o aeaeyae? eieaeueieo aeioii?a, ui
iai?eaeaoi? aeey ??eueieoeoaa. Ia nueiaiaei? ?o ieiua noaiiaeoue
aeecueei 1%. ?acii c oei noaie iathoue aaee/acio iaoeiao oe?ii?noue. Oea
nai???aeiee a?ii c? niaoeeo?/iith aaeaioeaiith aei aeieia?/ieo oiia
a?ioith, ui a?aeia?aaea?oueny o no?oeoo?? a?iii?o, niiniaao ?iciiiaeaiiy
oa ?icnaeaiiy ?ineei, ?ioaineaiiio aeaeiooai?aii?. Aeecueei 10% aeae?a
oei?e ? aiaeai?eaie oa noaaiaeai?eaie, oiio noaie iio?aaothoue
iaa?aeeeaaeieo ioi?iiieo caoiae?a. O ca’yceo c oeei iaaecae/aeii
aaaeeeaei nueiaiaei? ? ioi?iia ?nioth/eo, aeyaeaiiy oa ?aca?aoaaiiy
iiaeo noaiiaeo ae?eyiie ? anoaiiaeaiiy a?aeiia?aeieo ?aaeei?a ?o
caa?aaeaiiy. Uia caa?aaoe oe?ii? oi?eaeuei? caeeoee, iaiao?aeia ?ic?iaea
niaoe?aeueii? i?ia?aie ioi?iie noai?a. Ia iniaeeao oaaao caneoaiaothoue
eo/i? noaie, iioe?ai? a e?ninoaiia?e cii?, ai aiie ? iaea?eueo
a?aceeaeie ? i?ae aio?iiiaaiiei i?anii oaeaeei ci?iththoue oei?enoe/iee
neeaae oa ao?a/athoue aoaeiii? ?ene.

Oiio aaeeeo iaoeiao oe?ii?noue ia? i?iaaaeaia aeena?oaioii aeyaeaiiy
noaiiaeo ae?eyiie, ioe?iea noaio, anaa?/ia aea/aiiy aeieiai-oeaiioe/ieo
oa oei?enoe/ieo iniaeeainoae ?ineeiiiai iie?eao noai?a i?aaeaiii?
/anoeie E?aiaa?aaeiiai E?ninoaio Oe?a?ie (IEEO).

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. ?iaioa iia’ycaia
?c iaoeiaith oaiaoeeith a?aeae?eo aeieia?? o?oinenoai ?inoeoooo aioai?ee
?i. I.A. Oieiaeiiai IAI Oe?a?ie, cie?aia c oaiith 279 “Aeieia?/ia
ioe?iea ??ciiiai?ooy oei?e oa ?ineeiiino? oa?oiiaaiue Oe?a?ie” oa
a?aioii Agreement ? II 58 ca I?ia?aiith i?aeo?eiee a?i??ciiiai?oiino?
(BSP) “Ii??aiyeueia ioe?iea o?oi??ciiiai?oiino? caiia?aeieo noaiiaeo
aeinenoai Oe?a?ie c iaoith iioei?caoe?? ?aaeei?a ioi?iie”.

Iaoa ? caaaeaiiy aeine?aeaeaiiy. Aeaoe ioe?ieo oei?enoe/ii?,
aeieiai-oeaiioe/ii? ??ciiiai?oiino? ?ineeiiiai iie?eao noai?a IEEO. Aeey
aeinyaiaiiy iaoe aoee iinoaaeai? oae? caaaeaiiy:

— aeyaeoe ae?eyiee ?c aeia?a caa?aaeaiith i?e?iaeiith noaiiaith
?ineeii?noth IEEO;

— i?iaanoe ?iaaioa?ecaoe?th ?o oei?e oa aeeiiaoe nenoaiaoe/iee,
a?iii?oieia?/iee, aeieiai-oeaiioe/iee ? aaia?ao?/iee aiae?ce;

— ?ic?iaeoe eeaneo?eaoe?eio noaio ?ineeiiino? noai?a ia oei?enoe/i?e
iniia? (ca iaoiaeii A?aoi-Aeaiea) oa aeaoe aeieiai-oeaiioe/io
oa?aeoa?enoeeo neioaenii?a;

— c’ynoaaoe iniaeeaino? ?ici?uaiiy ?ineeiiiai iie?eao noai?a a
caeaaeiino? a?ae aeieia?/ieo oiia;

— aeyaeoe i?noeac?inoaiiy ??aee?nieo noaiiaeo aeae?a, ? caianaieo aei
“*a?aiii? eieae Oe?a?ie” a oaeiae oa?oiiaaiue ?c “Caeaii? eieae
Oe?a?ie”, aeaoe ?o oa?aeoa?enoeeo oa neeanoe ea?oinoaie ?o iioe?aiiy.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. C’yniaaii i?noea
aeine?aeaeaieo noai?a ii a?aeiioaiith aei noaiiaeo caiia?aeiee?a
Oe?a?ie. Ciaeaeaii a?eueoa 100 ae?eyiie (caaaeueiith ieiuath aeecueei
2,8 oen. aa), aea aeia?a caa?aaeaia i?e?iaeia noaiiaa ?ineeii?noue.
Oia?oa io?eiai? iiai? aeai? i?i oaeniiii?/iee neeaae oei?e noai?a IEEO –
644 aeaee c 264 ?iae?a, 51 ?iaeeie oa 3 a?aeae?e?a. Aeeiiai? ??
nenoaiaoe/iee, a?iii?oieia?/iee, aeieiai-oeaiioe/iee oa aaia?ao?/iee
aiae?ce, ?acoeueoaoe yeeo i?aeoaa?aeeee ca’ycie aeine?aeaeaii? oei?e ?c
oei?ith noa?iaeaaiueiai Na?aaecai’y. Aia?oa ?ic?iaeaii eeaneo?eaoe?eio
noaio ?ineeiiino? noai?a IEEO ia oei?enoe/i?e iniia? (ca iaoiaeii
A?aoi-Aeaiea), yea aeeth/a? 3 eeane, 10 nithc?a, 23 anioe?aoe?? oa 3
noaanioe?aoe??, i?e oeueiio iiooea/ai aoei aia?oa iienaii 2 nithce, 9
anioe?aoe?e oa 3 noaanioe?aoe??. Anoaiiaeaii no/ania iioe?aiiy
??aee?nieo aeae?a (12 aeae?a, caianaieo aei ?a?iiaenueeiai /a?aiiiai
nieneo, 27 aeae?a, caianaieo aei “*a?aiii? eieae Oe?a?ie”, oa 34, ui
iio?aaothoue ?aa?iiaeueii? ioi?iie) oa 10 oa?oiiaaiue, caianaieo aei
“Caeaii? eieae Oe?a?ie”.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. ?ic?iaeaia eeaneo?eaoe?eia
noaia ?ineeiiino? aeei?enoiao?oueny aeey i?aeaioiaee aeaeaiiy
“Neioaeniiii?y ?ineeiiino? Oe?a?ie”. Aeai? uiaei a?iii?oieia?/ieo,
aeieiai-oeaiioe/ieo, aaia?ao?/ieo iniaeeainoae aeae?a aeei?enoiaothoueny
i?e i?aeaioiaoe? “Aeieia?? aeae?a oei?e Oe?a?ie”. ?acoeueoaoe
aeine?aeaeaiiy ? aianeii o ?ic?iaeo i?iaeaie ioi?iie noaiiaeo aeinenoai.
Ca o/anoth aeena?oaioa cai?iiiiiaaii noai?aiiy 56 i?e?iaeii-caiia?aeieo
ia’?eo?a caaaeueiith ieiuath iaeaea 2,7 oen. aa.

Iniaenoee aianie caeiaoaa/a. ?iaioa ? naiino?eiei aeine?aeaeaiiyi
caeiaoaa/a, yeei i?iaaaeaii 8 aeniaaeeoe?e, aeeiiaii 698 aaiaioai?/ieo
iien?a, caeeaaeaii 4 aeieiai-oeaiioe/i? i?io?e?, c?a?aii 350 aa?aa?ieo
c?ace?a, c?iaeaiee iiaeeaeaiee aiae?c e?oa?aoo?e, noai?aii eiiniaeo
oei?e noai?a ?aa?iio aeine?aeaeaiue, ea?oe iioe?aiiy ??aee?nieo aeae?a
oa oa?oiiaaiue, aeiiiaiaii eeaneo?eaoe?eio noaio ?ineeiiino? ia
oei?enoe/i?e iniia?.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. ?acoeueoaoe aeiiia?aeaeeny ia
can?aeaiiyo a?aeae?eo aeieia?? o?oinenoai (1997, 1998), A/aii? ?aaee
?inoeoooo aioai?ee ?i. I.A. Oieiaeiiai IAI Oe?a?ie (1998),
i?aeia?iaeiiio nai?ia?? “?acaeoea nenoaiu iaaeainoaea?noaaiiuo iniai
io?aiyaiuo i?e?iaeiuo oa??eoi?ee” (Ia?oiaa, 1996), “VI iieiae?aeiie
eiioa?aioeee aioaieeia a Naieo-Iaoa?ao?aa” (1997), eiioa?aioe?yo
iieiaeeo o/aieo ? niaoe?ae?no?a “Aeooaeuei? ieoaiiy aioai?ee ? aeieia??”
(sseoa, 1993; Aeiiaoeuee, 1995; Oa?e?a, 1996; Iaeana, 1997; Oa?nii,
1998), i?aeaoc?anueeeo iaoeiai-iaoiaee/ieo eiioa?aioe?yo c i?iaeai
i?e?iaeie/eo iaoe (Iieoaaa, 1993, 1995, 1997).

Ioae?eaoe??. Ca iaoa??aeaie aeena?oaoe?eii? ?iaioe iioae?eiaaii 25
i?aoeue (o oiio /ene? iaeia iiiia?ao?y – o ni?aaaoi?noa?, 5 noaoae a
“Oe?a?inueeiio aioai?/iiio aeo?iae?”, 2 noaoo? a “Oe?a?inueeiio
o?oioeaiieia?/iiio ca??ieeo”, 5 noaoae o ca??ieeao iaoeiaeo i?aoeue, 12
oac aeiiia?aeae.

No?oeoo?a oa ianya ?iaioe. Caaaeueiee ianya aeena?oaoe?eii? ?iaioe
noaiiaeoue 341 noi??ieo, c ieo iniiaiiai oaenoo 154. Aiia ?ethno?iaaia
27 ?enoieaie oa 17 oaaeeoeyie, neeaaea?oueny c? anooio, 6 ?icae?e?a,
aeniiae?a oa nieneo e?oa?aoo?e (377 iaeiaioaaiue, c ieo 34 ?iicaiieo), 4
aeiaeaoe?a: A) eiiniaeo oei?e aeueo noaeeiieo ?ineei noai?a IEEO, A)
neioaeniiii?y ?ineeiieo oa?oiiaaiue noai?a ?aa?iio aeine?aeaeaiue, A)
ea?oinoaie iioe?aiiy ??aee?nieo aeae?a oei?e, Ae) ea?oinoaie iioe?aiiy
??aee?nieo oa?oiiaaiue.

INIIAIEE CI?NO

?icae?e 1. Oa?aeoa?enoeea i?e?iaeieo oiia ?aa?iio aeine?aeaeaiue

I?ai?/ia iaaea ?aa?iio aeine?aeaeaiue i?ioiaeeoue ii e?i?? Cieioiiioa –
Ie?ai?iae – Iooe?ea ? ii ?. Ai?neea aei ei?aeiio c ?in??th, a i?aaeaiia
– Eiaaeyee – E?aniia?aae – Ci??a – Eoi’yinuee, cao?aeia – ii ?. Noea, a
no?aeia – ii ?. Ineie. Aea?oueny oa?aeoa?enoeea aaieia?/ii? aoaeiae,
a?oioo, ee?iaoo, ?ineeiiiai iie?eao IEEO.

O no/aniiio ?ineeiiiio iie?ea? IEEO noaie caeiathoue aeecueei 1% ieiu? ?
i?eo?i/ai? aei noee?a aaeie, y??a, o?aieythoueny ia no??ieeo i?aaeo
aa?aaao ??/ie N?aa?nueeee Aeiiaoeue, Ineie, Ai?neea, Aia/a oa ?o i?eoie,
aocueeeie no??/eaie — ia i?e??/eiaeo aiaeiae?eueieo ieaoi, a oaeiae ia
ai?aao. Aiie aeineoue aeia?a i?aaenoaaeythoue ?ineeii?noue aa?aenoeo
??ciio?aaii-oei/aeiai-eiaeeiaeo noai?a, a ni?aaaei? oei/aeiai-eiaeeia?
noaie i?aaenoaaeai? o?aaiaioa?ii. Iniaeeao a?oio neeaaeathoue noaiia?
ae?eyiee oce?nue oa e?niaeo aaeyaei, ye? oa?aeoa?ecothoueny aenieith
aeaeiaith ??ciiiai?oi?noth aiane?aeie ii?aeiaiiy ni?aaaei?o noaiiaeo oa
oce?nieo aeae?a. Oe? ae?eyiee na?ae/aoue i?i i?e?iaei?noue a E?ninoaio
aeaio ciiaeueieo oei?a ?ineeiiino?: e?niai? oa noaiiai?.

Ae?eyiee ?c inaiio?oiith noaiiaith ?ineeii?noth i?eo?i/ai? a iniiaiiio
aei ino??aii? nenoaie ae?ea Ai?neee oa ino?ia?a Aei?i?iaeca?aeeinueeiai,
E?aiai/ooeueeiai ? Ia/ai?cueeiai aiaeinoiaeu, a oaeiae e?aeo ai?iaeo
??/eiaeo oa?an. Iniaeeao a?oio ooai?ththoue ea?aiiaoi? noaie,
i?eo?i/ai? aei e?aeaeyieo a?aeneiiaiue a aeieeiao ??/ie N?aa?nueeee
Aeiiaoeue, Ineie, Aia/a oa ?o i?eoie.

Oi/a noaie ? ? ciiaeueiei oeiii ?ineeiiino? aeey IEEO, iaoiiaeaiei
o?ceei-aaia?ao?/ieie iniaeeainoyie ? ee?iaoii, iaeiae ae?eyiee ?c aeia?a
caa?aaeaiith noaiiaith ?ineeii?noth o?aieythoueny ??aeei, caeiath/e
a?aeiinii iacia/i? ieiu?.

?icae?e 2. ?noi??y aea/aiiy noai?a i?aaeaiii? /anoeie E?aiaa?aaeiiai
E?ninoaio Oe?a?ie

Aea/aiiy noai?a ii/eia?oueny c XVII no. ? aeoea?co?oueny a O?O no. o
ca’yceo c aeine?aeaeaiiyi i?e?iaee aoaa?i?e Ee?anueeiai o/aiaiai ie?oao
(O?aoooaooa?, 1853,1855; ?iaiae/, 1853, 1862, 1869; Iiio?aci?,
1886-1888, 1891). Cia/iee aianie c?iaeee o/anieee aeniaaeeoe??,
i?aai?ciaaii? E?niaei aeaia?oaiaioii aeey i?e?iaeie/i-?noi?e/iiai
aeine?aeaeaiiy Iieoaanueei? aoaa?i?? (Aeieo/a?a, 1953; E?aniia, 1890,
1891, 1894). Ocaaaeueithth/eie oei?enoe/ieie caaaeaiiyie e?ioey O?O no.
aoee i?aoe? ?.O. Oiaeueaaocaia “Oei?a THai-Caiaaeiie ?innee” (1886) oa
“Oei?a N?aaeiae e THaeiie ?innee, E?uia e Naaa?iiai Eaaeaca” (1895,
1897). Ieoaiiy aaiaceno ?ineeiiino? e?aeaeyieo a?aeneiiaiue ?ic?iaeyee
Ae.?. Eeoaeiia (1902) oa A.?. Oae??a (1902, 1905, 1907), aeeneon?? i?ae
yeeie a cia/i?e i??? aecia/eee iiaeaeueoee ?icaeoie aioai?/ieo
iai?yie?a.

Iniiai? oei?enoe/i? oa aaiaioai?/i? aeine?aeaeaiiy noai?a IEEO
i?eiaaeathoue ia e?iaoeue XIX – ii/aoie XX noie?oue (?ee?/aanueeee, 1926
– 1930, 1936; E?aniia, 1890, 1891, 1893, 1894; Eeaiiia, 1930, 1933,
1934, 1941; Eaa?aiei, 1918, 1924, 1925, 1928, 1931, 1936, 1939, 1940;
Ia/inueeee 1892, 1925; Eioia, 1927, 1928, 1931, 1939 oa ?io?). O
i?neyai?iiee ia??iae i?iaeiaaeoaaeiny iaeiie/aiiy oaeoe/iiai iaoa??aeo,
aea iniiaia oaaaa i?eae?eyeany iaienaiith ooiaeaiaioaeueieo
ocaaaeueithth/eo “?ineeii?noue O?N?. Noaie, eai’yieno? a?aeneiiaiiy,
i?nee”, 1973; “Oei?a O?N?” (1936-1965), “Aaiaioai?/ia ?aeiioaaiiy
Oe?a?inueei? ?N?” (1977), “Aaia?aoey ?anoeoaeueiiai iie?iaa Oe?aeiu”
(1982), “I?iae?iion ?anoeoaeueiinoe Oe?aeiu” (1991). I?iaeiaaeo?oueny
aea/aiiy inaiio?oii? ?ineeiiino? (Eeieia, 1981) oa e?aeaeyieo
a?aeneiiaiue ? ea?aiiaoieo noai?a (Eioia, 1953; Ii?icthe, 1971; Iaoeia,
1951), a oaeiae ocaaaeueiaiiy a?aeiiinoae uiaei oei?e, ?ineeiiino? ?
aaiaceno eo/ieo ? ??ciio?aaii-oei/aeiai-eiaeeiaeo noai?a (A?eee, 1955;
Naeaei, 1955, 1957). A inoaii? aeanyoee?ooy caa?oa?oueny oaaaa ia
iaiao?aei?noue ioi?iie noai?a a oe?eiio (Oea/aiei, 1993; Oea/aiei,
I?yaeei, 1990) ? oa?eoi??? IEEO cie?aia (I?ieoae?i oa ?i., 1979, 1980;
Ai?aeiaa, 1989; Ai?aeiaa oa ?i., 1990, 1993; Oea/aiei oa ?i., 1986;
Aoaianiaa oa ?i., 1992; Aiae???iei oa ?i., 1992, 1993, 1996; Aae?ae oa
?i., 1992, 1994-1996),

A?aenooi?noue aiae?co no/aniiai noaio ?ineeiiiai iie?eao noai?a IEEO
aeeeeeaea iio?aao aeaoaeueiiai aeine?aeaeaiiy ?o oei?enoe/ii? oa
aeieiai-oeaiioe/ii? aeeoa?aioe?aoe??.

?icae?e 3. Iaoa??aee oa iaoiaeeea aeine?aeaeaiue

A iniiao ?iaioe iieeaaeaii iaoa??aee iieueiaeo aeine?aeaeaiue,
i?iaaaeaieo aaoi?ii i?ioyaii 1993-1998 ??. ia oa?eoi??? IEEO. Iieueia?
aeine?aeaeaiiy i?iaiaeeeeny caaaeueiii?eeiyoeie iaoiaeaie (ia?o?ooiei oa
iai?anoaoe?iia?iei) (Iieaaay aaiaioaieea, 1959-1976). Cae?eniaii 698
aaiaioai?/ieo iien?a ia i?iaieo ae?eyieao (10o10 i). Aeey anoaiiaeaiiy
caeiiii??iinoae ?iciiae?eo ?ineeiieo oa?oiiaaiue o caeaaeiino? a?ae
a?oioiai-a?ae?ieia?/ieo oiia oa ?aeue?oo caeeaaeaii 4 aeieiai-oeaiioe/i?
i?io?e?. Aeey ia?iaee iaoa??ae?a aeei?enoiaoaaee eiii’thoa?iee aaie
aeaieo ii oei?? oa ?ineeiiino? a?aeae?eo aeieia?? o?oinenoai ?inoeoooo
aioai?ee ?i. I. A. Oieiaeiiai IAI Oe?a?ie oa eaoaae?e aioai?ee
Ee?anueeiai oi?aa?neoaoo. Aeei?enoaii i?ia?aie a?aae??ioiiai,
i?aeeiaoe?eiiai aiae?c?a, iaoiaeeeo o?oi?iaeeeaoe?? aeieia?/ieo oaeoi??a
(Ae?aeoo, Iethoa, 1994).

?icae?e 4. Oei?enoe/ia aeeoa?aioe?aoe?y noai?a i?aaeaiii? /anoeie

E?aiaa?aaeiiai E?ninoaio Oe?a?ie

Iaeiei c iaeaaaeeea?oeo caaaeaiue i?e aea/aii? ?ineeiiiai na?oo ?aa?iio
? ?iaaioa?ecaoe?y ? aiae?c eiai oei?e. Ie iiae?ey?ii iiaeyaee ss.I.
Ae?aeooa (1992) i?i oa, ui oei?a iiaea ia’?aeioaaoe aeaee ia eeoa ca
oa?eoi??aeueiei i?eioeeiii, a ? ca oeiieia?/iei (noeoii?noue aeae?a
iaaiiai oeio ?ineeiiino?), ine?eueee oaea oei?a oa?aeoa?eco?oueny
iaaieie aeanoeainoyie, ui caeiiii??ii ci?iththoueny. Naia c oeeo
iiceoe?e ?icaeyaea?oueny ? aiae?co?oueny oei?a noai?a IEEO.

4.1. Nenoaiaoe/iee aiae?c

Ia i?aenoaa? i?ea?iaeueieo, e?oa?aoo?ieo oa aa?aa?ieo aeaieo ie neeaee
eiiniaeo i?e?iaeii? oei?e noaeeiieo ?ineei noai?a, yeee ia?aoiao? 644
aeaee, ui iaeaaeaoue aei 264 ?iae?a, 51 ?iaeeie oa 3 a?aeae?e?a. Iai?aio
?ieue a?ae?a?athoue Magnoliophyta (99,6%), iacia/io /anoeio noaiiaeyoue
noaeeii? nii?ia? oa aieiian?ii? (0,4%). Na?aae Magnoliophyta ia
Liliopsida i?eiaaea? 16,6%, ia Magnoliopsida – 83%, ui noaiiaeoue
i?iii?oe?th 1:5. Oeae iieaciee aeecueeee aei a?aeiia?aeieo iieaciee?a
oei? Noa?iaeaaiueiai Na?aaecai’y (1:4-4,5) (Oieia/?a, 1970, 1974).
Iniiaia i?iii?oe?y (?iaeeie: ?iaee: aeaee) noaiiaeoue 1:5,2:12,6.

O aea/ai?e oei?? aeii?iothoue iaaaaaoi ?iaeei, ui i?eoaiaiii oei??
Oe?a?ie (Oaeya-Niniiei, Ae?aeoo, 1975) ? oei?? Aiea?eoeee a oe?eiio. O
?iaeeiiiio niaeo?? o?e ia?o? ?iaeeie i?noyoue 35,2% caaaeueii?
e?eueeino? aeae?a, ui aeaui a?eueoa, i?ae o oei?? Oe?a?ie (34,5%).
Aeanyoue i?ia?aeieo ?iaeei ioiieththoue 452 aeaee (70,2%) ? 184 ?iaee
(69,7%), a ?io? ?iaeeie i?aaenoaaeai? 192 aeaeaie ? 80 ?iaeaie, ui
noaiiaeoue, a?aeiia?aeii 29,8 oa 30,3%. Ianooii? ?iaeeie, ye? caeiathoue
a niaeo?? 11-20 i?noey, ia?aoiaothoue 7 -17 aeae?a ? caaaeii
aeeth/athoue 107 aeae?a (16,6%) oa 35 (13,2%), a ?aooa – 31 ?iaeeia
ioiieth? eeoa 85 aeae?a ? 45 ?iae?a, ui, a?aeiia?aeii, noaiiaeoue 13,2%
oa 17,1%. ?c ieo 10 ?iaeei (19,6%) i?noyoue ii 1 ?iaeo ? 1 aeaeo.
I?ia?aeia i?noea o niaeo?? oei?e noai?a IEEO, ye ? a a?eueoino?
aiea?eoe/ieo oei?, caeia? ?iaeeia Asteraceae (17,2%). ?iaeeia Poaceae
(9,5%) noi?oue ia ae?oaiio i?noe?, ai noaie a?aeiinyoueny aei oeaiic?a
o?aa’yienoiai oeio, ? iniiao ?ineeiiiai iie?eao oi?iothoue ceaee.
?iaeeia Fabaceae a aeine?aeaeai?e oei?? iin?aea? o?ao? i?noea (8,5%).
?iaeeia Caryophyllaceae ia?aaoaa? ia /aoaa?oiio i?noe? (6,8%), ui oaeiae
oa?aeoa?ii aeey oei?e Aaeeai, I?aaeaiii? ?a?iie ? Na?aaecaiiiii?’y a
oe?eiio. ?iaeeia Lamiaceae iin?aea? i’yoa i?noea (6,1%) ? ? iaei??th c
i?ia?aeieo o oei?ao Noa?iaeaaiueiai Na?aaecai’y (Caoe?ia, 1951), ui
i?aee?aneth? iaeaaei?noue noai?a aei oa?oiiaaiue ena?ioe/iiai oeio.
?iaeeia Brassicaceae caeia? oinoa i?noea (5,4%), ui iia’ycaii ?c
neiaio?ii?caoe??th oei?e. Ianooii? 7-10 i?noey a?aeiia?aeii caeiathoue
?iaeeie Scrophullariaceae (5,3%), Rosaceae (5%), Boraginaceae (3,3%),
Apiaceae (3,1%). sseui iieiaeaiiy Lamiaceae oa Caryophyllaceae ?
aeineoue aenieei ? iaoiiaeaia cia/iith o/anoth i?aaeaiieo aeae?a, oi
?iaeeia Cyperaceae ciaoiaeeoueny cia/ii ieae/a, i?ae o oei?? E?ninoaio a
oe?eiio (I?einueeee, 1969), ine?eueee noaiiaa oei?a a oe?eiio ? cia/ii
ena?ioe/i?oith ? iicaaaeaia ai?aaeueieo aeaiaio?a, oa?aeoa?ieo aeey
?aa?iio. A?aeiioaiiy ?iaeei Fabaceae+Caryophyllaceae aei Cyperaceae
noaiiaeoue 7,6, ui na?ae/eoue i?i aeineoue ena?io?oi? oiiae ?icaeoeo
(eoee – 3,3%, aieioa – 0,3%), a Asteraceae+Brassicaceae aei Rosaceae
noaiiaeoue 4,6, ui oa?aeoa?eco? oeiiao ceaeiao noaae?th ?icaeoeo
oeaiic?a, aeineoue aeaeaeeo a?ae ee?iaeniaiai noaio (Ae?aeoo, Iethoa,
1994)

?iaeiaee niaeo? a?aeia?aaea? iaea?eueo caaaeuei? ?ene oei?e. O oei??
noai?a IEEO anueiai ia?aoiao?oueny 264 ?iaee. Ca?aaeiy e?euee?noue
aeae?a o ?iae? noaiiaeoue 2,4, ui iaioa, i?ae aeey E?aiaa?aaeiiai
E?ninoaio a oe?eiio. A aeanyoe i?ia?aeieo ?iaeao i?noeoueny 16,3% a?ae
caaaeueii? e?eueeino? aeae?a, a o 15-oe – 22,3%. Aiae?c ?iaeiaiai
niaeo?a oei?e na?ae/eoue i?i iaioaaiiy oeiiaiai na?aaecaiiiii?nueeiai
?iaeo Centaurea – 15 aeae?a. Ii 13 aeae?a iathoue ?iaee Carex i
Veronica, a ii 11 aeae?a – oeiia? na?aaecaiiiii?nuee? ?iaee Astragalus
oa Dianthus. Ianooii? 6-7 i?noey o ?iaeiaiio niaeo?? caeiathoue ?iaee,
ye? iathoue ii 9 aeae?a – Potentilla i Silene, a 8-15 i?noey
iin?aeathoue ?iaee, ye? ia?aoiaothoue ii 8 aeae?a: Stipa, Salvia, Rosa,
Linum, Campanula, Vicia, Artemisia, Allium. I?ia?aei? ?iaee iathoue
ia?aaaaeii na?aaecaiiiii?nueea iioiaeaeaiiy.

4.2. A?iii?oieia?/iee aiae?c

Oa?aeoa?iith ?enith oei?e noai?a ? ia?aaaaeaiiy a ?? neeaae?
o?aa’yienoeo ?ineei — 593 aeaee (92,1%). Iiae?aiee a?iii?oieia?/iee
niaeo? i?eoaiaiiee oei?? noai?a oa aaiiyeiaeo a?aeneiiaiue
I?aaiaa?aaeiiai Ceaeiaiai Noaio (E?eoeueea, 1985). Ia ia?oiio i?noe? a
niaeo?? IEEO ciaoiaeyoueny o?aa’yieno? iie?ea?i?ee 63,4%. Ia ae?oaiio –
o?aa’yieno? iiiiea?i?ee – 28,7% (na?aae yeeo 14,3% iaeii??/iee?a), ui ?
niaoeeo?/iith ?enith na?aaecaiiiii?nueei? oei?e. Iaeiae oeae iieaciee
cia/ii ieae/ee, i?ae o oei?? noai?a I?aaiaa?aaeiiai Ceaeiaiai noaio
(36,5%). A?aeiinii aeniea o/anoue iiiiea?i?/ieo aeae?a na?ae/eoue, c
iaeiiai aieo, i?i na?aaecaiiiii?nuee? ?ene aeine?aeaeaii? oei?e
(?oaoeia, 1970), c ?ioiai – i?i iaaei??iee aieea ia oeth ?ineeii?noue
aio?iiiaaiieo oaeoi??a, ui i?aeoaa?aeaeo?oueny iayai?noth o neeaae?
iaeii??/iee?a aaeeei? e?eueeino? ?oaea?aeueieo aeae?a c oe?ieei a?aaeii.
*anoea eou?a oa eouee?a iaaeniea – a?aeiia?aeii, 3,7 oa 3,9%, iaeiae
aeaye? i?aaenoaaieee oe??? a?oie a?ae?a?athoue nooo?ao ?ieue o
oi?ioaaii? ?ineeiiiai iie?eao, ine?eueee aiie ? aeii?iaioaie a
eaeueoeeo?eueiinoaiiaeo, inaiio?oiinoaiiaeo oa noaiiaeo o?oioeaiicao.
Aea?aaa i?aaenoaaeai? o oei?? noai?a IEEO iaeiaioa – 0,3%, ui oaeiae ?
oa?aeoa?iith iciaeith.

Oei aoaeiae iaaecaiieo iaaii?a ei?aeth? c ee?iaoe/ieie oa aaeao?/ieie
oiiaaie. Aeecueei iieiaeie aeae?a (321, 49,8%) iathoue iai?a?icaoeia?
iaaiie, aeaui iaioa – 275 aeae?a (42,7%) – aac?icaoeia?. E?euee?noue
?icaoeiaeo aeae?a iacia/ia – 48 (7,5%), ui ? oa?aeoa?iei aeey oei?
iii??ii? ciie Aiea?eoeee.

A oiiaao noaio iai?aia iieiaeaiiy caeiathoue aaaaoaoeaii ia?ooii?
?ineeie c eaoaeaenaie – 34,2%, ye? a?ae?a?athoue cia/io ?ieue o
noai?aii? oeaiic?a, i?eo?i/aieo aei a?aeneiiaiue aaiiye?a oa eano.
Ei?ioei- oa aeiaaiei?aiaaeuia no?oeoo?a i?aecaiieo i?aai?a aeanoeaa
33,1% aeae?a noaiiai? oei?e IEEO. O?ao? i?noea iin?aeathoue aeaee aac
ei?aiaaeuii? no?oeoo?e – 176 aeae?a (27,3%). Aoeueaiei?aiaaeui? oa
oeeaoeeii? aeaee o neeaae? oei?e noaiiaeyoue a?aeiia?aeii, 0,8 ? 3,7%
a?ae caaaeueii? e?eueeino?, a?eueo?noue ?o ? aoaia?i?aeaie. Iacia/ia
e?euee?noue aeae?a (0,9%) iaeaaeeoue aei a?oie iaaecaiiinoieiiieo.

?acoeueoaoe a?iii?oieia?/iiai aiae?co na?ae/aoue i?i ii?aeiaiiy o
aeine?aeaeai?e oei?? aeaiaio?a oa?iio?eueieo oei? na?aaecaiiiii?nueeiai
oeio oa a?eaeieo aeaiaio?a ?a?iiaenueei-ac?aonueeiai oeio.

4.3. Aeieiai-oeaiioe/iee aiae?c

Aeieiai-oeaiioe/io no?oeoo?o oei?e a?aeia?aaea? e?euee?nia
ni?aa?aeiioaiiy aeae?a, i?eo?i/aieo aei iaaieo oeaiic?a, oa aieea ye
i?yiiae?th/eo (oaiea, aieiae iia?o?y, na?oea, o?i?/iiai neeaaeo
a?oio?a), oae ? iiina?aaeeiaaieo (ee?iaoo ? ?aeue?oo) aeieia?/ieo
oaeoi??a.

O oei?? iniiao (339 aeae?a) neeaaeathoue aai?e?eioio?oe. Cia/ia o/anoue
oa?io?o?a (14,3%) oa aaio?o?a (10,9%) aecia/a?oueny a?eaeieie ?enaie
ee?iaoo. I?aaenoaaeai?noue oaia?io?o?a ? oaiao?o?a o oei?? iacia/ia – 3
oa 5%, oi/a aeaye? i?aaenoaaieee a?ae?a?athoue nooo?ao ?ieue o
oi?ioaaii? ?ineeiiiai iie?eao. sse iieacaee ?ic?aooiee, i?iaaaeai? ss.I.
Ae?aeooii oa I.A. Iethoith (1994), aeineoue aeia?a ?iaeeeo?oueny noai
aeinenoaie ia iniia? a?aeiioaiiy noie oaia?io?o?a ? oaiao?o?a aei
oa?io?o?a, ui aeey noai?a IEEO noaiiaeoue 0,55 ? oa?aeoa?eco? ?o aaeeeo
ii?ooai?noue a?aeiinii ee?iaeniaiai noaio.

O oei?? noai?a IEEO aeae?eaii n?i aeieiai-oeaiioe/ieo a?oi. Aeecueei
iieiaeie aeae?a iaeaaeeoue aei a?oi noaiiaeo – 185 aeae?a (28,7%) oa
eo/ii-noaiiaeo – 132 aeaee (20,5%) ?ineei. Ia o?aoueiio i?noe?
ciaoiaeeoueny a?oia oce?nieo aeae?a – 18,2%. Aaeao?/i? aa??aioe noai?a
i?aaenoaaeai? inaiio?oiith (64 aeaee, 9,9%) oa ea?aiiaoiith (28 aeae?a,
2,8%) a?oiaie. Eo/ia a?oia i?aaenoaaeaia 40 aeaeaie (6,2%). Iayai?noue a
iaaeao ?aa?iio cia/ieo ieiu aio?iiiaai?ciaaieo oa?aeue iicia/eeany ia
no?oeoo?? oei?e, a neeaae? yei? c’yaeeanue cia/ia e?euee?noue
?oaea?aeueieo aeae?a (78 aeae?a, 12,1%).

Aeieiai-oeaiioe/iee aiae?c oei?e iieacaa cia/io ??ciiiai?oi?noue oei?a
aeaeiai? aaeaioaoe?? aei ie?aieo aeieia?/ieo oaeoi??a noaiiaeo oeaiic?a.

4.4. Aaia?ao?/iee aiae?c

A iniiao aaia?ao?/iiai aiae?co iieeaaeaii noaio aioai?ei-aaia?ao?/iiai
?aeiioaaiiy caiii? eoe? ? eeaneo?eaoe?th a?aae?a, ?ic?iaeaio A. Iiecaeai
oa ?i. (1965). Aeey oei?e noai?a IEEO oa?aeoa?iei ? ia?aaaaeaiiy
a?aeiinii aocueeeo oei?iciiaeueieo a?aaeia?/ieo oei?a. Aeecueei iieiaeie
aeae?a oei?e iathoue noana?aaecaiiiii?nueei-noaoaiia?aoi? (133 aeaee,
20,6%) oa noana?aaecaiiiii?nueei-oaiia?aoi? (263 aeaee, 25,3%) a?aaee.
?yae aeae?a i?eo?i/ai? aei iaei??? c oei?enoe/ieo cii:
noana?aaecaiiiii?nueei? (69 aeae?a, 10,7%), oaiia?aoii?, aai iii??ii?
(32 aeaee, 5%) oa noaoaiia?aoii? (5 aeae?a, 0,8%), a i?noeaciaoiaeaeaiiy
a IEEO ? eieae?oaoaie. Oaeei /eiii, yae?i oei?e noai?a IEEO oi?iothoue
aeaee, iioe?ai? a iaaeao noana?aaecaiiiii?nueei? ? oaiia?aoii? cii – 402
aeaee (62,4%). Aeecueei o?aouei? /anoeie caaaeueii? e?eueeino? aeae?a
(207, 32,1%) ia aeoiaeyoue ca iaae? noana?aaecaiiiii?nueei? oa
noaoaiia?aoii? cii, ui aeaco? ia oyae?iiy oei?e noai?a aeine?aeaeaiiai
?aa?iio naia aei noaiiai? ciie (Eaa?aiei,1942, 1961; Iiiia, 1963).

A?aaee aeine?aeaeaieo aeae?a oyae?thoue aei ?aa?ii?a c iii??iei nooiaiai
ieaai?/iino? ee?iaoo. Iaea?eueoith ? e?euee?noue aeae?a c
noaieaai?/ii-noaeiioeiaioaeueiei (173, 26,8%),
ieaai?/ii-noaeiioeiaioaeueiei (141, 21,9%) oa
noaieaai?/ii-eiioeiaioaeueiei (121, 18,8%) oeiaie a?aaeo. Aeaui iaioith
? o/anoue aeae?a c noaeiioeiaioaeueii-eiioeiaioaeueieie (88 aeae?a,
13,7%) oa noaieaai?/ii-noaeiioeiaioaeueieie (95, 14,8%) a?aaeaie.
E?euee?noue aeae?a, a?aaee yeeo iia’ycai? c i?aeaeuaiith ieaai?/i?noth
ee?iaoo iacia/ia, ui iiynith?oueny iineaaeaieie ca’yceaie aeine?aeaeaii?
oei?e ?c oeaio?aeueii-?a?iiaenueeeie iaii?aeueii-iiioaiieie oa
na?aaecaiiiii?nueeeie iiioaiieie oeaio?aie.

O oei?? noai?a IEEO ia?aaaaeathoue aeaee c ?a?iiaenueeei (268 aeae?a,
41,6%) oa ?a?iiaenueei-ac?aonueeei (242 aeaee, 37,6%), cie?aia
?a?iiaenueei-cao?aeiiac?aonueeei (154, 23,9%) oeiaie a?aaeo. A oe?e
a?oi? aeineoue cia/iith ? /anoea aeae?a c oe?ieeie ?a?ac?aonueeeie
a?aaeaie – 81 aeae (12,6%). Iacia/io aeieth neeaaeathoue aeaee c
?a?iiaenueei-no?aeiiac?aonueeei – 2 aeaee (0,3%) oa
?a?iiaenueei-ia?aaeiueiac?aonueeei – 5 aeae?a (0,8%) oeiaie iioe?aiiy.
O?aoueith ca /enaeuei?noth ? a?oia ?a?iiaenueei-nea??nueeiai a?aaoeio —
63 aeaee (9,7%). A?oia c ieth?e?aa?iiaeueiei oeiii a?aaeo i?aaenoaaeaia
39 aeaeaie (6,1%), na?aae yeeo cia/ia /anoeia ?oaea?aeueieo aeae?a.

O neeaae? oei?e ia?aoiao?oueny 77 (11,9%) aiaeai?/ieo aeae?a.
A?eueo?noue aiaeai?e?a a?aeiineoueny aei iiioe/iiai oeio – 40 aeae?a
(6,2%), cia/ii iaioa – aei na?iaonueeiai – 16 (2,5%),
iiioe/ii-iaiiiinueeiai – 10 (1,6%) oa iiioe/ii-na?iaonueeiai – 8 (1,2%)
? eeoa iacia/ia /anoeia – aei na?iaonueei-iaiiiinueeiai – 3 (0,5%).
Aenieee a?aenioie aiaeai?cio iiaeia iiynieoe nai???aei?noth
aeine?aeaeaii? oei?e ?, cie?aia, cia/iith o/anoth eaeueoeeo?oieo ?
inaiio?oieo aeae?a, ui coiiaeaii nai???aei?noth aeieia?/ieo oiia oa
i?ioeanaie aeaeiooai?aiiy (Eeieia, 1981).

?icae?e 5. Aeieiai-oeaiioe/ia aeeoa?aioe?aoe?y ?ineeiiiai iie?eao noai?a

Aeeoa?aioe?aoe?y ?ineeiiiai iie?eao ni?e/eiaia aieeaii aeaio neeaaeiaeo
— a?ieia?/ieie iniaeeainoyie iiioeyoe?e oa aeae?a, ye? aca?iiae?thoue
i?ae niaith, ? oaeoi?aie ciai?oiueiai na?aaeiaeua, ui iieeaaeaii a
iniiao eeaneo?eaoe?? ?ineeiiino?. A Oe?a?i? oaea eeaneo?eaoe?y
aacoaaeanue ia aeii?iaioieo i?eioeeiao.

5.1. Caaaeueia oa?aeoa?enoeea ?ineeiiino? noai?a i?aaeaiii? /anoeie
E?aiaa?aaeiiai E?ninoaio Oe?a?ie

A iniiao aeii?iaioii? eeaneo?eaoe?? noaiiai? ?ineeiiino? iieeaaeaii
i?eioeeie, ?ic?iaeai? ?.I. Eaa?aieii (1940, 1954, 1956). Ia oa?eoi???
IEEO eo/i? noaie i?aaenoaaeai? a?nueiiia oi?iaoe?yie: Stipeta
capillatae, S. pennatae, S. tirsae, Festuceta valesiace, Poeta
angustifoliae, Calamagrostideta epigeii, Bromopsideta inermis, Cariceta
humilis, ye? iioe?ai? ii anueiio ?aa?iio aeine?aeaeaiue. Iaeiae e?aua
aiie i?aaenoaaeai? a i?ai?/i?e /anoei? ? a??oa – ia iaae? c? noaiiaith
ciiith. Oeiia? ??ciio?aaii-oei/aeiai-eiaeeia? noaie, ui
oa?aeoa?ecothoueny aeii?ioaaiiyi iaciena?io?oieo oa ena?io?oieo
aea?ieiieo ceae?a oa aaeeeith o/anoth aiaiaeo ? ??ciio?aa’y,
i?aaenoaaeai? oi?iaoe?yie Stipeta lessingianae, S. zalesskii, S.
pulcherrime, S. capillatae, S. dasyphyllae, Festuceta valesiacae, Poeta
angustifoliae, Bromopsideta ripariae, B. inermis oa Koelerieta
cristatae.

Iniaeeao a?oio noaiiaeo oa?oiiaaiue neeaaeathoue ea?aiiaoi? oa
inaiio?oi? aaeao?/i? aa??aioe. Oa?oiiaaiiy ea?aiiaoieo noai?a ? aeineoue
aeeiai?/ieie. O i?noeyo ?icaeoeo e?aeaeyieo a?aeneiiaiue aiie
ci?iththoueny /aaa?ie/eiaeie oa?oiiaaiiyie oeio oii?ey??a. Inaiio?oi?
noaiia? oeaiice oi?iothoueny ia i?uaieo a?oioao ? ia oa?eoi??? IEEO
i?eo?i/ai? aei i?uaieo oa?an ??/eiaeo aeieei oa ino?ia?a
E?aiai/ooeueeiai, Aei?i?iaeca?aeeinueeiai oa Ia/ai?cueeiai aiaeinoiaeu.
A?eueo?noue aeae?a, ye? ooai?ththoue ?ineeiiee iie?ea ? inaiio?oaie.
Iai?aia ?ieue o neeaae? o?aainoith oeeo noai?a iaeaaeeoue
aaena?io?eueiei ? iaciena?io?eueiei aeaeai aea?ieiieo ceae?a (Festuca
beckeri, Koeleria glauca), ye? c?inoathoue eeoa ia i?neoaaoeo ? i?eiee
ia ia?aoiaeyoue ia noaeeieno? a?oioe.

5.2. Eeaneo?eaoe?y ?ineeiiino? ia oei?enoe/i?e iniia? (ca iaoiaeii
A?aoi-Aeaiea)

Ia iniia? aeeiiaieo 678 aaiaioai?/ieo iien?a iaoiaeii ia?aoai?aiiy
o?oioeaiioe/ieo oaaeeoeue (Einiai oa ?i., 1991) ?ic?iaeaia oei?enoe/ia
eeaneo?eaoe?y, ui aea? iiaeeea?noue ii??aiyoe aeai? c
cao?aeii?a?iiaenueeeie. Neioaeniiii?/ia noaia aeeth/a? 3 eeane, 4
ii?yaeee, 10 nithc?a, 23 anioe?aoe?? oa 3 noaanioe?aoe??. Oaea aeniea
??ciiiai?oi?noue coiiaeaia no?ieao?noth aeieia?/ieo (aaeao?/ieo)
/eiiee?a, ?ieue yeeo iaiaeiaeiaa.

Ceioaeniiii?/ia noaia ?ineeiiino? noai?a IEEO

Cl. Festuceta vaginatae Soo 1968 em Vicherek 1972

Ord. Festucetalia vaginatae Soo

All. Festucion beckeri Vicherek, 1972

Ass. Veronico dillenii-Secalietum sylvestri Shevchyk et V. Sl. 1996

Sass. Veronico dillenii-Secalietum sylvestri dianthosum platyodon Didukh
et Korotchenko 1996

Ass. Centaureo borysthenicae-Festucetum beckeri Vicherek 1972

Ord. Festuco-Astragaletalia arenarii Vicherek 1972

All. Koelerion glaucae Volk 1931

Ass. Thymo angustifolii-Festucetum beckeri Vicherek 1972

Cl. Festuco-Brometea Br.-Bl. et Tx. 1943

Ordn. Festucetalia valesiacae Br.-Bl. et R. Tx. 1943.

All. Cirsio-Brachypodion pinnati Hadac et Klika 1994 em Krausch 1961

Ass. Thymo marschalliani-Caricetum praecocis Korotchenko, Didukh, 1997

All. Fragario viridis-Orifolion montani Korotchenko, Didukh, 1997

Ass. Medicago romanicae-Poetum angustifoliae Tkachenko, Movchan et V.
Sl., 1987; Medicago-Festucetum valesiacae Wagner, 1940; Betonico
officinalis-Trifolietum montani Popova in Popova et al. 1986; Salvio
pratensis-Poetum angustifoliae Korotchenko, Didukh, 1997; Veronico
austriacae-Nhamaecytisetum austriaci Korotchenko, Didukh, 1997

All. Astragalo-Stipion Knapp 1944

Ass. Stipetum pennatae K. Jovanovic 1956; Astragalo austriaci-Salvietum
nutantis Korotchenko, Didukh, 1997; Thymo marschalliani-Crinitarietum
villosae Korotchenko, Didukh, 1997

Sass. Thymo marschalliani-Crinitarietum villosae Astragaletosum
onobrychis Korotchenko, Didukh, 1997

Ass. Stipetum lessingianae Soo 1948; Vinco herbaceae-Caraganetum
fruticis Korotchenko, Didukh, 1997

All. Festucion valesiacae Klika 1931

Ass. Festuco valesiacae-Stipetum capillatae Sillinger 1931; Salvio
nemorosae-Festucetum valesiacae Korotchenko, Didukh, 1997; Festucetum
valesiacae Solodkova et al., 1986; Tkachenko et al., 1987; Mirkin et
al., 1988

All. Artemisio-Kochion Soo 1959

Ass. Agropyro pectinato-Kochietum prostratae Zolyomi 1958 corr. Soo 1959

Sass. Agropyro pectinato-Kochietum prostratae Marrubietosum praecocis
Korotchenko, Didukh, 1997

All. Artemisio marschalliani-Elytrigion intermediae Korotchenko, Didukh,
1997

Ass. Astragalo dasyanthi- Elytrigietum intermediae Korotchenko, Didukh,
1997

All. Nentaureo carbonati-Koelerion talievii Romaschenko, Didukh et V.
Sl., 1996

Ass. Androsaco koso-poljanskii-Caricetum humilis Korotchenko, Didukh,
1997

Cl. Ielianthemo-Thymetea Romaschenko, Didukh et V. Sl. 1996

Ord. Ohymo cretacei-Hyssopetalia cretacei Didukh 1989

All. Artemisio hololeucae-Hyssopion cretacei Romaschenko, Didukh et V.
Sl. 1996

Ass. Onosmo tanaiticae-Androsacietum kozo-poljanskii Romaschenko, Didukh
et V. Sl. 1996; Artemisio hololeucae-Polygaletum cretaceae Didukh 1989;
Scrophulariae cretacei-Helianthemum cretacei Romaschenko, Didukh et V.
Sl. 1996

Eean Festuco-Brometea ? iaea?eueo ?ai?acaioaoeaiei ia aeai?e oa?eoi??? ?
aeeth/a? eo/i? oa oeiia? noaiia? oa?oiiaaiiy ia iiooaeieo /i?iicaiieo
a?oioao. Oeiia? (ni?aaaei?) noaie oa?aeoa?ecothoueny ia?aaaaeaiiyi
aaena?io?eueieo ? aa?eena?io?eueieo noaiiaeo ?ineei, aei yeeo o cia/i?e
e?eueeino? iino?eii aeii?oothoueny aaaaoi??/i? ena?iiacio?euei? ?
aaiacio?euei? ?ineeie: aieiaiei /eiii ??ciio?aa’y oa ei?aiaaeui? ceaee.
Eean Festuceta vaginatae aeeth/a? oa?oiiaaiiy inaiio?oieo noai?a,
i?eo?i/aieo aei ae?oai? i?uaii? ai?iai? oa?ane ??/ie. Eaeueoeeo?euei?
noaie eeano Helianthemo-Thymetea oi?iothoueny ia a?aeneiiaiiyo e?aeaee
i?e a?aenooiino? a?oioiaiai iie?eao, eoaeii? ?aaeoe?? (?I 7,5-8),
iecueeiai ai?noo aoiono (aei 1,5%) ? i?ia?aeueiiai acioo oa iayaiino?
ea?aiiaoiiai oeio canieaiiy (?iiauaiei oa ?i., 1996). Ianya neioaenii?a
oei?enoe/ii? oa aeii?iaioii? eeaneo?eaoe?? ??ciee ? ia ca?aa?oueny.

5.3. Aeieia?/ia oa?aeoa?enoeea ?ineeiiino? noai?a IEEO

Noaiia? aeinenoaie oi?ioaaeeny a oiiaao iaaeinoaoiuei? aieiaino? ee?iaoo
i?e ia??iaee/iiio aieea? aeianaiiy, iae?a, ui ia?aoeiaeaeaei iiya?
aea?aa. O noaiao iiiaae 80% a?iiane ui??/ii a?aeie?a? ? aiia oaeaeei
i?ia?ae?co?oueny, ui aecia/a? niaoeeo?eo e?oaiia?ao aeaiaio?a ?
oi?ioaaiiy iae?iaeth/?oeo /i?iicaiieo a?oio?a (Ae?aeoo oa ?i., 1998).

Aeey aecia/aiiy iniiaieo aeieia?/ieo oaeoi??a canoiniaaii iaoiae
o?oi?iaeeeaoe??, ?ic?iaeaiee o a?aeae?e? aeieia?? o?oinenoai ?inoeoooo
aioai?ee ?i. I.A. Oieiaeiiai IAI Oe?a?ie ss.I. Ae?aeooii oa I.A.
Iethoith (1994). Aiae?c i?iaiaeeee ca aeaa’youeia i?ia?aeieie
aeieia?/ieie oaeoi?aie: aieia?noth, eeneioi?noth a?oioo, caaaeueiei
nieueiaei ?aaeeiii, ai?noii i?ia?aeueiiai acioo, ea?aiiao?a, oa?i?/iei
?aaeeiii, aoi?aei?noth, ii?ici?noth, eiioeiaioaeuei?noth ee?iaoo.

Noaiia? aeinenoaie ? ?ano?niaeao?oeeoieie. Iniiaiei e?i?ooth/ei oaeoi?ii
noaiiaeo o?oioeaiic?a ii a?aeiioaiith aei e?niaeo ? eo/ieo ? aeao?oeeo
aieiae, ui iaoiiaeth? i?e?iaeia aace?nny ieaei?ieo i?noeac?inoaiue. A
iaaeao noaiiaeo aeioii?a iaea?eueoa aa??th? ai?no ea?aiiao?a eaeueoe?th
a a?oio?: a?aeiioaiiy ?aaeueii? aiie?ooaee aei iaeneiaeueii iiaeeeai? a
i?e?iae? noaiiaeoue 39,1%. Ioaea, naia ai?no ea?aiiao?a o a?oio?
a?ae?a?a? i?ia?aeio ?ieue o oi?ioaaii? ??cieo oei?a oa?oiiaaiue noaiiai?
?ineeiiino?, cie?aia aeeoa?aioe?aoe?? ia o?e eeane: inaiio?oieo,
ni?aaaei?o ? ea?aiiaoieo noai?a. Aeineoue cia/iith ? aiie?ooaea
eeneioiino? a?oioo – 17,8% oa ai?noo acioo – 17,6% ? aeaui iaioith
aa??aoe?y aieiaino? a?oioo – 13,3% oa caaaeueiiai nieueiaiai ?aaeeio
(10,2%). A?aeiinii iaaaeeea aiie?ooaea aieiaino? a?oioo iiynith?oueny
oei, ui noaie oi?iothoueny eeoa i?e oiia? iaaeinoaoiuei? caieiaeaiino?
a?oioo, i?e ca?eueoaii? ai?noo aieiae a a?oio? aiie cai?uothoueny
eo/ieie oa e?niaeie oa?oiiaaiiyie, a i?e ciaioaii? – a?aeaoaa?oueny
iionoaethaaiiy.

Oa?oiiaaiiy ia ??ai? eean?a oa ii?yaee?a aeineoue /?oei a?ae??ciythoueny
i?ae niaith ca a?eueo?noth aeieia?/ieo oaeoi??a, iniaeeai aaeao?/ieo.
Aiie?ooaea nithc?a oi/a e ia?ae?eaa?oueny ca ie?aieie oaeoi?aie, aea
caaaeaee iiaeia aeae?eeoe o? c ieo, ye? aecia/athoue aeieia?/io
niaoeeo?eo neioaenii?a. Oey aiie?ooaea cia/ii oe?oa (1-3 aaee) aeey
aaeao?/ieo oaeoi??a, i?ae ee?iaoe/ieo (1 aae). Oiio naia nithc aoei
ia?aii iniiaiith iaeeieoeath ii??aiyeueiiai aiae?co – ye aeieia?/io
nenoaio, ui ia? naith niaoeeo?eo. Ii??aiyiiy na?aaei?o iieaciee?a
aeieia?/ieo oaeoi??a aeey noai?a IEEO ?c aiaeia?/ieie iieacieeaie
noaiiaeo aeinenoai caiia?aeiee?a Oe?a?ie iieacaei ?o iiae?ai?noue
(Ae?aeoo oa ?i., 1998), ui na?ae/eoue i?i aaeao?/io ? ee?iaoe/io
oeiia?noue noaiiaeo aeinenoai. C ?ioiai aieo, i?iaaaeaiee aiae?c
na?ae/eoue i?i oa, ui oei?enoe/ia eeaneo?eaoe?y a?aeia?aaea? ia eeoa
aeaeiao niaoeeo?eo neioaenii?a, a e ?o aeieia?/io nooue oa ia? aenieo
i?iaiinoe/io oe?ii?noue.

?icae?e 6. Ioi?iia noaiiai? oei?e oa ?ineeiiino? a i?e?iaeii-caiia?aei?e

ia?aae? ?aa?iio aeine?aeaeaiue

Ine?eueee noaie ?noi?e/ii oi?ioaaeenue a oiiaao iino?eiiai aieeao
aeciaaiieo oaeoi??a. O ?ac? eiai onoiaiiy /e ci?ie aiie caciathoue
aeineoue nooo?aeo o?ainoi?iaoe?e (Ae?aeoo, Eenaiei, 1993). A oiiaao
aaniethoii caiia?aeiiai ?aaeeio ?o aeaiooaoe?y a?aeaoaa?oueny a iai?yieo
ieoaia?iiy (Oea/aiei, 1993). I?iaeaia ioi?iie noai?a iaaoaa? aaaeeeaiai
cia/aiiy o ca’yceo c aaeeeith ?ici?ai?noth ?aa?iio oa ?ioaineaiei eiai
inai?iiyi.

6.1. ??aee?ni? aeaee oei?e

Aaaeeeaei iieacieeii noaio caa?aaeaiino? i?e?iaeieo eiiieaen?a iaaii?
oa?eoi??? ? iayai?noue o neeaae? ?? oei?e ??aee?nieo aeae?a, ye? ?
aeoaea /ooeeaeie aei o?ainoi?iaoe?? aai caa?oaeiaiiy na?aaeiaeua ? oiio
cieeathoue a ia?oo /a?ao. Aeey nicieia?/ii? ioe?iee aeae?a aoea
aeei?enoaia cai?iiiiiaaia N.I. Noieeii (1982) ? aeiiiaiaia TH.?.
Oaeyaii-Niniieii oa ss.I. Ae?aeooii (1985) eiiieaenia oa?aeoa?enoeea
aeae?a, yea a?aoiaoaaea 10 iciae: aioai?ei-aaia?ao?/io cia/ou?noue
aeaeo, oa?aeoa? oi?eaeueiino? aeaeo aeey ?aa?iio, oaeniiii?/io
?ai?acaioaoeai?noue, e?euee?noue i?noeaciaoiaeaeaiue, ieiuo an?o
i?noeac?inoaiue, na?aaeith ?yni?noue aeaeo a oa?aeoa?ieo oeaiicao,
ni?yiiaai?noue ci?ie aeoeaiino? oeaiiiiioeyoe?e i?ae aieeaii
aio?iiiaaiieo oaeoi??a, oaeaee?noue caanaiiy aeoeaiino? oeaiiiiioeyoe?e
i?ae aieeaii aio?iiiaaiiiai oaeoi?a, oaeaee?noue a?aeiiaeaiiy
oeaiiiiioeyoe?e i?ney ?o ii?ooaiiy, i?aeoe/ia aeei?enoaiiy aeaeo
ethaeeiith /e aaeeaaiiy oaa?eiaie. C o?aooaaiiyi oeeo iieaciee?a aeey
noai?a IEEO aeae?eaii 67 aeae?a, ui iio?aaothoue ioi?iie.

Cie?aia, aei ?a?iiaenueeiai *a?aiiiai nieneo caianaii 11 aeae?a:
Dianthus lanceolatus, Scrophularia cretacea, Senecio borysthenicus,
Tragopogon ucrainicus, T. borystenicus, Silene cretacea, Androsace
koso-poljanskii, Genista tanaitica, Hyssopus cretaceus, Artemisia
hololeuca, Elytrigia stipifolia. Na?aae aeae?a, ui c?inoathoue ia
noaiiaeo ae?eyieao ?aa?iio aeine?aeaeaiue, 28 caianaii aei “*a?aiii?
eieae Oe?a?ie” (1996).

Iaeiei ?c iaeaoaeoeai?oeo caoiae?a ioi?iie i?noeac?inoaiue ??aee?nieo
aeae?a ia no/aniiio aoai? ? noai?aiiy i?e?iaeii-caiia?aeieo ia’?eo?a.
An? ??aee?ni? aeaee oei?e noai?a IEEO caaacia/ai? ioi?iiith ia noai?aieo
oa iioei?ciaaieo i?e?iaeii-caiia?aeieo oa?eoi??yo.

6.2. ??aee?ni? oa?oiiaaiiy noai?a IEEO

Ia noaiiaeo ae?eyieao aeyaeai? oa?oiiaaiiy, caianai? aei “Cae?iie eieae
Oe?aeineie NN?” (1987), a yeeo aeii?iothoue /e ni?aaeii?iothoue
??aee?ni? aeaee, aai oae?, ui ia?aaoaathoue ia iaae? a?aaeo ?
iio?aaothoue ioi?iie: (Amygdaleta nanae, Stipeta lessingianae, S.
capillatae, S. pulcherrimae, S. pennatae, S. dasyphyllae, S. tirsae, S.
zalesskii, Elytrigieta stipifoliae, Cariceta humilis). Aei “Caeaii?
eieae” ne?ae aeeth/eoe ??aee?ni? ?ae?eoia? oa?oiiaaiiy, a yeeo
aeii?iaioii /e ni?aaeii?iaioii aenooia? Androsace koso-poljanskii Ovcz.
Iacaai? oa?oiiaaiiy oa?aeoa?ecothoueny aenieith aeaeiaith iane/ai?noth
oa iayai?noth ??aee?nieo aeae?a.

6.3. I?e?iaeii-caiia?aei? oa?eoi???

Noai?aiiy ia?aae? i?e?iaeii-caiia?aeieo oa?eoi??e – iaea?eueo aoaeoeaiee
oeyo caa?aaeaiiy aaiioiiaeo ?ineeiiiai na?oo, oeiiaeo ? ??aee?nieo
eaiaeoaoo?a, i?aeo?eiaiiy ni?eyoeeaeo aeieia?/ieo oiia (Nioe, 1986).

I?e?iaeii-caiia?aeia ia?aaea Iieoaaueie aei 1992 ?. aoea ?icaeiaia
aeoaea neaai. A?aenioie caiia?aeiino? noaiiaea 0,5 i?e na?aaeiueiio a
Oe?a?i? 2,3. Oei?a oa ?ineeii?noue noai?a ioi?iiyeany o oanoe caeacieeao
i?noeaaiai cia/aiiy, ieiuath aeecueei 50 aa. Oa?eoi??th oe?eaaiai
caiia?aeieea “Aeaaeai?/iee noai” a i?neyai?ii? ?iee aoei ?ici?aii. Ca
?acoeueoaoaie i?iaaaeaieo aaiaioai?/ieo oa oei?enoe/ieo aeine?aeaeaiue
ca o/anoth aeena?oaioa ia?aaeo i?e?iaeii-caiia?aeieo oa?eoi??e iaeano?
aoei a cia/i?e i??? aeiiiaiaii ? ?icoe?aii. Ia?aaea
i?e?iaeii-caiia?aeieo oa?eoi??e a Oa?e?anuee?e iae. ia ii/aoeo
aeine?aeaeaiue aoea oaeiae iaaeinoaoiuei noi?iiaaia. A?aenioie
caiia?aeiino? ooo noaiiaeoue 0,9. Noaiia? ae?eyiee ioi?iiythoueny
o?aaiaioa?ii a aioiiieia?/ieo caeacieeao.

?aa?iiaeuei? eaiaeoaooi? ia?ee ? iiaith eaoaai???th a
i?e?iaeii-caiia?aeiiio oiiae? Oe?a?ie. Iacaaaeath/e ia oa, ui ieiua
Aeeeainueeiai ?aa?iiaeueiiai eaiaeoaooiiai ia?eo aeineoue cia/ia (11945
aa), ae?eyiee ?c noaiiaith ?ineeii?noth ia eiai oa?eoi??? iaei iioe?ai?,
i?eo?i/ai? a iniiaiiio aei cao?aeii?, iaio cae?niaii? /anoeie ?
caeiathoue noeee aaeie i?ai?/ii? oa i?aaeaiii? aeniiceoe??. I?iaeoiaaiee
?EI “Ia/ai?cueea iiea” (ieiua aeecueei 4,5 oen. aa), ?icoaoiaaiee ia
i?aaei? E?aiaa?aaeiiai E?ninoaio, ia iaae? c? noaiiaith ciiith, cia/ii
?icoe?eoue o/anoue noai?a o caiia?aei?e ia?aae?.

Ia ?nioth/eo ? cai?iiiiiaaieo i?e?iaeii-caiia?aeieo oa?eoi??yo IEEO a
aeinoaoi?e i??? i?aaenoaaeai? an? oeie noaiiai? ?ineeiiino?.
Iaeaieiai?oa aeey caa?aaeaiiy noaiiaeo ae?eyiie – oea caai?iia ?o
?ici?thaaiiy, oa?anoaaiiy oa caaoaeiae, aaeaea oae? caoiaee
ni?e/eiththoue aei ?o caaeaae?. E??i oiai, aeey a?aeiiaeaiiy
/enaeueiino? ??aee?nieo ? cieeath/eo noaiiaeo aeae?a o ia?o?e iieiaei?
e?oa caai?iiy?oueny aeeiooaaiiy, a oaeiae ?aaoeth?oueny nooi?iue
aeianaiiy.

AENIIAEE

1. Noaie, ui ? ciiaeueiei oeiii ?ineeiiino? i?aaeaiii? /anoeie
E?aiaa?aaeiiai E?ninoaio Oe?a?ie (IEEO) ? ca?ac caeiathoue ieiuo
aeecueei 1% aeine?aeaeaii? oa?eoi???, o?aieythoueny a ?aa?ii? aeineoue
/anoi aocueeeie no??/eaie eeoa ia i?eieaei?ieo ae?eyieao, ai?aao ?
noeeao aaeie ??/eiaeo aeieei ??cii? e?ooecie oa aeniiceoe??. Iaie
aeyaeaii aeecueei 100 ae?eyiie c aeia?a caa?aaeaiith i?e?iaeiith
?ineeii?noth eo/ii-noaiiaiai oeio.

2. Oei?a noai?a IEEO ia?aoiao? 644 aeaee, ui iaeaaeaoue aei 264 ?iae?a,
51 ?iaeeie oa 3 a?aeae?e?a. Nenoaiaoe/ia no?oeoo?a a?aeaeaa? ena?ioe/iee
oa?aeoa? ? oyae?? aei na?aaecaiiiii?nueeiai oeio.

3. Iniiao a?iii?oieia?/iiai niaeo?o oei?e noai?a IEEO neeaaeathoue
o?aa’yieno? iie?ea?i?ee c iai?a?icaoeiaith ? ?icaoeiaith aoaeiaith
iaaii?a oa eaoaeaenii. A aeieiai-oeaiioe/iiio niaeo?? ia?aaaaeathoue
noaiia? ? eo/ii-noaiia? c aaeeeith e?euee?noth oce?nieo aeae?a, ui
iiynith?oueny noaa?eaeiei, aai?ena?io?oiei oa?aeoa?ii oiia ?nioaaiiy,
ii?ooai?noth ia?aeiii? no?oeoo?e oeaiic?a, aeaeaeeo a?ae ee?iaeniaiai
noaio, ? oi?ioaaiiyi ?o ia i?noe? e?n?a.

4. Ca oeiaie a?aae?a o aeine?aeaeai?e oei?? ia?aaaaeathoue
noana?aaecaiiiii?nueei-noaoaiia?aoi?, noaieaai?/ii-noaeiioeiaioaeuei?
aeaee. Aenieee a?aenioie aiaeai?/ieo aeae?a (11,9%) na?ae/eoue i?i
naiiaooi?e oa?aeoa? noai?a ? i?i oa, ui a iaaeao noaiiaiai a?iio
?ioaineaii a?aaeoaathoueny i?ioeane aeaeiooai?aiiy.

5. O caeaaeiino? a?ae oei?a a?oio?a noaiiaa ?ineeii?noue i?aaenoaaeaia
o?ueiia eeanaie: Festuco-Brometea (eo/i? noaie, iioe?ai? ii an?e
oa?eoi??? oa ??ciio?aaii-oei/aeiai-eiaeeia? noaie, ui c??aeea
o?aieythoueny ia iaae? c? Noaiiaith ciiith) ia /i?iicaiieo a?oioao,
Helianthemo-Thymetea (eaeueoeeo?oiinoaiia? oa?oiiaaiiy, i?eo?i/ai? aei
i?aaeaiii-no?aeii? /anoeie aeine?aeaeaii? oa?eoi???) ia ea?aiiaoieo
?aiaeceiao, Festuceta vaginatae (inaiio?oiinoaiia? oa?oiiaaiiy,
o?aaiaioa?ii i?aaenoaaeai? ia ai?iaeo oa?anao ??/eiaeo aeieei) ia
i?uaieo a?aeeeaaeao.

6. Ca?aeii c ?ic?iaeaiith iaie eeaneo?eaoe?eiith noaiith ?ineeii?noue
noai?a i?aaeaiii? /anoeie E?aiaa?aaeiiai E?ninoaio i?aaenoaaeaia 3
eeanaie, 10 nithcaie, 23 anioe?aoe?yie oa 3 noaanioe?aoe?yie, c yeeo 2
nithce, 9 anioe?aoe?e oa 3 noaanioe?aoe?? iienai? aeena?oaioii.

7. Ia iniia? canoinoaaiiy iaoiaeeee o?oi?iaeeeaoe?? oa i?aeeiaoe?eiiai
aiae?co anoaiiaeaii, ui i?ia?aeieie /eiieeaie oi?ioaaiiy noai?a ?
a?ae?i?aaeei oa ai?no ea?aiiao?a o a?oio?, ye? coiiaeththoue iniaeeaino?
noaiiaeo a?oio?a (eeneioi?noue, o?ioi?noue, ai?no acioo oiui).
Anoaiiaeaii oa?aeoa? oa nooi?iue ei?aeyoe?? i?ae oaeoi?aie. ?aia nithco
aenooia? o??th neioaeniiii?/iith eaoaai???th, yea a?aeia?aaea?
niaoeeo?eo aeinenoaie ca eiiieaenii aeieia?/ieo oaeoi??a.

8. O oei?enoe/iiio neeaae? noai?a IEEO aeyaeaii 12 aeae?a, caianaieo aei
?a?iiaenueeiai *a?aiiiai nieneo, 27 aeae?a, caianaieo aei “*a?aiii?
eieae Oe?a?ie”, oa 34 aeaee, ye? iio?aaothoue ?aa?iiaeueii? ioi?iie.
Aeine?aeaeaii iioe?aiiy oa aeaia aooo?oinioe?ieia?/ia oa?aeoa?enoeea
oeeo aeae?a.

9. Anoaiiaeaii, ui ia noaiiaeo ae?eyieao o?aieythoueny oa?oiiaaiiy
aeanyoe oi?iaoe?e, caianaieo aei “Caeaii? eieae Oe?a?inueei? ?N?”
(Amygdaleta nanae, Stipeta. lessingianae, S. capillatae, S.
pulcherrimae, S. pennatae, S. dasyphyllae, S. tirsae, S. zalesskii,
Elytrigieta stipifoliae, Cariceta humilis) oa ??aee?ni? oa?oiiaaiiy
oi?iaoe?? Androsaceta koso-poljanskii, ui iio?aaothoue ioi?iie.

10. Cai?iiiiiaai? (ca o/anoth aaoi?a) aeey ioi?iie ae?eyiee noaiiai?
?ineeiiino? aeathoue ciiao ?icoe?eoe ?ai?acaioaoeai?noue
i?e?iaeii-caiia?aeii? ia?aae? ?aa?iio ? caaacia/eoe caa?aaeaiiy oeiiaeo
oa ??aee?nieo aeae?a ? oa?oiiaaiue.

NIENIE i?aoeue, IIOAE?EIAAIEO CA OAIITH AeENA?OAOe??

1. Ii??aiyeueia ioe?iea o?oi??ciiiai?oiino? caiia?aeieo noaiiaeo
aeinenoai Oe?a?ie c iaoith iioei?caoe?? ?aaeei?a ?? ioi?iie / ss.I.
Ae?aeoo, A.N. Oea/aiei, I.A. Iethoa, ?.A. Ei?io/aiei (Aaaei), O.A.
O?oeaeei / I?ae caa. ?aae. ss.I. Ae?aeooa. – ?inoeooo aioai?ee ?i. I.A.
Oieiaeiiai IAI Oe?a?ie. – Ee?a, 1998. – 75 n.

2. Ei?io/aiei (Aaaei) ?.A. ??aee?ni? oa?oiiaaiiy noai?a Iieoaaueie. //
Oe?. aioai. aeo?i. — 1996. — 53, ? 3. — N. 274-277.

3. Aae?ae-Niiey? I.I., Ei?io/aiei (Aaaei) ?.A. Oei?enoe/ia ? oeaiioe/ia
oa?aeoa?enoeea noai?a Iieoaaueie // Oai aea. — 1996.- 53, ? 5. — N.
554-561.

4. Ei?io/aiei (Aaaei) ?.A. Oeaiiaeieia?/i? iniaeeaino? eo/ii-noaiiaiai
caeacieea “Ia?aaeeoaaoa aea/a” // Oai aea. — 1998. – 55, ? 2. – N.
201-207.

5. Ei?io/aiei (Aaaei) ?.A. Neioaeniiii?/ia oa aeieia?/ia
aeeoa?aioe?aoe?y noaiiaeo oa?oiiaaiue i?aaeaiii? /anoeie E?aiaa?aaeiiai
E?ninoaio Oe?a?ie // Oai aea. — 1998. – 55, ? 4. – N. 403-410.

6. Ei?io/aiei-Aaaei ?.A., Iaaei?oa I.TH. Noaiiaa ?ineeii?noue
Aeeeainueeiai ?aa?iiaeueiiai eaiaeoaooiiai ia?eo // Oai aea. — 1998. –
55, ? 5. – N. 556-561.

7. Ae?aeoo ss.I., Ei?io/aiei (Aaaei) ?.A. Noaiiaa ?ineeii?noue
i?aaeaiii? /anoeie E?aiaa?aaeiiai E?ninoaio Oe?a?ie. ?. Eeane Festucetea
vaginatae oa Helianthemo-Thymetea // Oe?. o?oioeai. ca. — Ee?a, 1996, —
Na?. A. Aei. 2. — N. 56-63.

8. Ei?io/aiei (Aaaei) ?.A., Ae?aeoo ss.I. Noaiiaa ?ineeii?noue
i?aaeaiii? /anoeie E?aiaa?aaeiiai E?ninoaio Oe?a?ie. ??. Eean
Festuco-Brometea // Oai aea. — Ee?a, 1997. – Na?. A, aei. 1(6). – N.
20-39.

9. Aiae?eaiei O.E., I?yaeei A.E., Ei?io/aiei (Aaaei) E.A. ?anoeoaeueiue
ie? i?iaeoe?oaiiai ?aaeiiaeueiiai eaiaeoaooiiai ia?ea “Ia/aiaaeneia
iiea” e aai iao/iay cia/eiinoue // Iaoa?eaeu iaaeaeoia?iaeiiai naieia?a
“?acaeoea nenoaiu iaaeainoaea?noaaiiuo iniai io?aiyaiuo i?e?iaeiuo
oa??eoi?ee”. – Eeaa, 1996. — N. 14-20.

10. Aae?ae I.I., Naii?iaeia A.I., Noaoethe I.I., Ei?io/aiei (Aaaei) ?.A.
Ioi?iia noaiiai? oei?e oa ?ineeiiino? a i?e?iaeii-caiia?aei?e ia?aae?
Iieoaanueei? iaeano? // Nioe?ieia?/i? i?iaeaie naea: aeiiii?ea ?
iaoai?caoe?y n?euenueeiainiiaea?nueeiai ae?iaieoeoaa. Oii. 18. –
Iieoaaa, 1995. – N. 99-106.

11. Aae?ae I.I., Noaoethe I.I., Ei?io/aiei (Aaaei) ?.A. Oei?a ?
?ineeii?noue noai?a Iieoaaueie oa ?o ioi?iia // Iaoeia? caienee. Na??y
i?e?iaeie/a (Aei 80-??//y Iieoaanueeiai iaaeaaia?/iiai ?inoeoooo). –
Iieoaaa, — 1995. — N. 33-37.

12. Ae?aeoo ss.I., Ei?io/aiei (Aaaei) ?.A. Iaoeia? canaaee caa?aaeaiiy
eo/ieo noai?a i?aaeaiii? /anoeie E?aiaa?aaeiiai E?ninoaio Oe?a?ie //
Iaoa?eaeu iaaeaeoia?iaeiiai naieia?a “?acaeoea nenoaiu
iaaeainoaea?noaaiiuo iniai io?aiyaiuo i?e?iaeiuo oa??eoi?ee”. – Eeaa,
1996. — N. 39-42.

13. Andrienko T.L., Priadko O.I.,. Korotczenko (Babko) I.A Swait
roslinny projektowanego Regio?nalnego Parku Kraj?obrazowego
Pieczenizskie Pole na ukra?nskich stepach // Chronmy przyrode ojczysta.
– 1996. R. 52. — N 6. – P. 5-14.

14. Aiae???iei O.E., Aae?ae I.I., Eeanoia I.E., Ei?io/aiei (Aaaei) ?.A.
Iioei?caoe?y caiia?aeii? ia?aae? Iieoaaueie // Ae?oa? Ea?eoeinuee?
/eoaiiy. I?aeaoc?anueea iaoeiai-iaoiaee/ia eiioa?aioe?y c i?iaeai
i?e?iaeie/eo iaoe. – Iieoaaa, 1993. — N. 3.

15. Aae?ae I.I., Ei?io/aiei (Aaaei) ?.A. Iioe?aiiy caeeoe?a noai?a ia
Iieoaauei? // Aeooaeuei? i?iaeaie iaoiaeeee aeeeaaeaiiy aaia?ao??
??aeiiai e?ath. Iaoa??aee iaoeiai-i?aeoe/ii? eiioa?aioe??. – Iieoaaa,
1994. – N. 21-23.

16. Aae?ae I.I., Noaoethe I.I., Ei?io/aiei (Aaaei) ?.A., Iaaei?oa I.TH.
Aoaie iioei?caoe?? i?e?iaeii-caiia?aeii? ia?aae? Iieoaaueie // O?ao?
Ea?eoeinuee? /eoaiiy. I?aeaoc?anueea iaoeiai-iaoiaee/ia eiioa?aioe?y c
i?iaeai i?e?iaeie/eo iaoe. (Aeieia?/i? aniaeoe). – Iieoaaa, 1995. — N.
5-6.

17. Ei?io/aiei (Aaaei) ?.A. Oei?a oa ?ineeii?noue i?iaeoiaaiiai
aioai?/iiai caeacieea “O?i/eua Othothiieeiaa” (E?aiai/ooeueeee ?aeii).
// Ae?oa? Ea?eoeinuee? /eoaiiy. I?aeaoc?anueea iaoeiai-iaoiaee/ia
eiioa?aioe?y c i?iaeai i?e?iaeie/eo iaoe. – Iieoaaa, 1993. — N. 45-46.

18. Ei?io/aiei (Aaaei) ?.A. Noaiia? i?e?iaeii-caiia?aei? oa?eoi??? ye
iie?aiie aeey i?iaaaeaiiy aeieia?/iiai aeoiaaiiy // O?ao? Ea?eoeinuee?
/eoaiiy. I?aeaoc?anueea iaoeiai-iaoiaee/ia eiioa?aioe?y c i?iaeai
i?e?iaeie/eo iaoe. (Aeieia?/i? aniaeoe). – Iieoaaa, 1995. — N. 5-6.

19. Ei?io/aiei (Aaaei) ?.A. Caeiiii??iino? ?ici?uaiiy ae?eyiie noaiiai?
?ineeiiino? ia oa?eoi??? Iieoaanueei? iaeano? // Aeooaeueiua aii?inu
aioaieee e yeieiaee. Oacenu aeieeaaeia. Eiioa?aioeey iieiaeuo o/?iuo e
niaoeeaeenoia. – Oa?ueeia, 1996. — N. 64.

20. Ei?io/aiei (Aaaei) E.A. ?anoeoaeueiinoue noaiae thaeiie /anoe
Eaaiaa?aaeiie Eaninoaie Oe?aeiu. // Oacenu VI iieiae?aeiie eiioa?aioeee
aioaieeia a Naieo-Iaoa?ao?aa. – Naieo-Iaoa?ao?a – 1997, – N. 80-81.

21. Ei?io/aiei (Aaaei) ?.A. Ioi?iia noaiiai? ?ineeiiino? a ?aa?iiaeueieo
eaiaeoaooieo ia?eao E?aiaa?aaeiiai E?ninoaio // *aoaa?o? Ea?eoeinuee?
/eoaiiy. Anaoe?a?inueea i?aeaoc?anueea iaoeiai-iaoiaee/ia eiioa?aioe?y c
i?iaeai i?e?iaeie/eo iaoe, i?enay/aia iai’yo? A.I. Ea?eoeia. Ca??iee
noaoae. *. ?. – Iieoaaa, — 1997. — N. 110-112.

22. Ei?io/aiei (Aaaei) ?.A., Aae?ae I.I. Noai oa ia?niaeoeae ioi?iie
noaiiai? ?ineeiiino? i?aaeaiii? /anoeie E?aiaa?aaeiiai I?eaei?i?ia’y //
Eiioa?aioe?y iieiaeeo o/aieo ? niaoe?ae?no?a. Aeooaeuei? ieoaiiy
aioai?ee ? aeieia?? (19-21 aeiaoiy 1993, sseoa). Oace aeiiia?aeae. –
Ee?a, 1993. — N. 63.

23. Ei?io/aiei (Aaaei) ?.A., Iaaei?oa I.TH. Noaiia? ae?eyiee
Aeeeainueeiai ?aa?iiaeueiiai eaiaeoaooiiai ia?eo ia Iieoaauei? //
I?iaeaie aioai?ee ? i?eieia?? ia ii?ic? o?aoueiai oeny/ie?ooy. Iaoa??aee
O c’?caeo Oe?a?inueeiai aioai?/iiai oiaa?enoaa. – o Ee?a; Iieoaaa, 1997.
– N. 203.

24. Iaaei?oa I.TH., Ei?io/aiei (Aaaei) ?.A. ??ciiiai?ooy ?ineeiiiai
na?oo Aeeeainueeiai ?aa?iiaeueiiai eaiaeoaooiiai ia?eo oa eiai
nicieia?/ia cia/aiiy // Iaoa??aee Anaoe?a?inueei? iaoeiai-i?aeoe/ii?
eiioa?aioe?? niaoe?ae?no?a oa nooaeaionueei? iieiae? “No/ani? iiaeyaee c
iiioey?ecaoe?? i?e?iaeie/eo iaoe”, i?enay/aii? iai’yo? A.?.
Aa?iaaenueeiai. – Iieoaaa, 1998. – N. 39.

25. Ei?io/aiei (Aaaei) ?.A. Neioaeniiii?/ia oa aeieia?/ia
aeeoa?aioe?aoe?y noaiiaeo oa?oiiaaiue i?aaeaiii? /anoeie E?aiaa?aaeiiai
E?ninoaio // Aeooaeuei? ieoaiiy aioai?ee oa aeieia??. Iaoa??aee
eiioa?aioe?? iieiaeeo a/aieo aioai?e?a. – Oa?nii, 1998. – N. 61-62.

Aaaei ?.A. Aeeoa?aioe?aoe?y ?ineeiiiai iie?eao noai?a i?aaeaiii? /anoeie
E?aiaa?aaeiiai E?ninoaio Oe?a?ie. – ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
ca niaoe?aeuei?noth 03.00.05 – aioai?ea. – ?inoeooo aioai?ee ?i. I.A.
Oieiaeiiai IAI Oe?a?ie, Ee?a, 1999.

Aeena?oaoe?th i?enay/aii ieoaiiyi ioe?iee oei?enoe/ii? ?
aeieiai-oeaiioe/ii? ??ciiiai?oiino? ?ineeiiiai iie?eao noai?a i?aaeaiii?
/anoeie E?aiaa?aaeiiai E?ninoaio Oe?a?ie (IEEO). Aeyaeaii aeecueei 100
ae?eyiie (1% oa?eoi???) ?c aeia?a caa?aaeaiith i?e?iaeiith ?ineeii?noth
eo/ii-noaiiaiai oeio. Anoaiiaeaii, ui o neeaae? oei?e ia?aoiao?oueny 644
aeaee, ye? iaeaaeaoue aei 264 ?iae?a, 51 ?iaeeie oa 3 a?aeae?e?a.
Aeeiiaii ?? nenoaiaoe/iee, a?iii?oieia?/iee, aeieiai-oeaiioe/iee oa
aaia?ao?/iee aiae?ce, ?acoeueoaoe yeeo i?aeoaa?aeeee ca’ycie
aeine?aeaeaii? oei?e ?c oei?ith Noa?iaeaaiueiai Na?aaecai’y. ?ic?iaeaii
eeaneo?eaoe?eio noaio ?ineeiiino? noai?a IEEO ia oei?enoe/i?e iniia? (ca
iaoiaeii A?aoi-Aeaiea), yea aeeth/a? 3 eeane, 10 nithc?a, 23
anioe?aoe?? oa 3 noaanioe?aoe??, i?e oeueiio caeiaoaa/ai aia?oa iienaii
2 nithce, 9 anioe?aoe?e oa 3 noaanioe?aoe??. Anoaiiaeaii, ui
i?ia?aeieie /eiieeaie oi?ioaaiiy noai?a ? a?ae?i?aaeei oa ai?no
ea?aiiao?a o a?oio?. Aeiaaaeaii, ui ?aia nithco aenooia? o??th
neioaeniiii?/iith eaoaai???th, yea a?aeia?aaea? niaoeeo?eo aeinenoaie ca
eiiieaenii aeieia?/ieo oaeoi??a. Aeyaeaii 12 aeae?a, caianaieo aei
?a?iiaenueeiai *a?aiiiai nieneo, 27 aeae?a – c “*a?aiii? eieae Oe?a?ie”,
34 aeaee, ui iio?aaothoue ?aa?iiaeueii? ioi?iie oa oa?oiiaaiue 10
oi?iaoe?e, caianaieo aei “Cae?iie eieae Oe?aeineie NN?”. Ca o/anoth
aaoi?a cai?iiiiiaaii noai?aiiy 56 i?e?iaeii-caiia?aeieo ia’?eo?a
caaaeueiith ieiuath aeecueei 2,7 oen. aa.

Eeth/ia? neiaa: i?aaeaiia /anoeia E?aiaa?aaeiiai E?ninoaio Oe?a?ie,
noai, oei?a, aiae?c, ?ineeii?noue, oei?enoe/ia eeaneo?eaoe?y,
o?oi?iaeeeaoe?y aeieia?/ieo oaeoi??a, ??aee?ni? aeaee, ??aee?ni?
oa?oiiaaiiy.

Aaaei E.A. Aeeoa?aioeeaoeey ?anoeoaeueiiai iie?iaa noaiae thaeiie /anoe
Eaaiaa?aaeiie Eaninoaie Oe?aeiu. – ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/?iie noaiaie eaiaeeaeaoa aeieiae/aneeo iaoe
ii niaoeeaeueiinoe 03.00.05 – aioaieea. – Einoeooo aioaieee ei. I.A.
Oieiaeiiai IAI Oe?aeiu, Eeaa, 1999.

Aeenna?oaoeey iinayuaia eco/aieth ninoiyiey nio?aiiinoe, oei?enoe/aneie
e yeieiai-oeaiioe/aneie aeeooa?eioeeaoeee ?anoeoaeueiiai iie?iaa noaiae
thaeiie /anoe Eaaiaa?aaeiie Eaninoaie Oe?aeiu (THEEO). Iaia?oaeaii aieaa
100 o/anoeia, ia eioi?uo oi?ioi nio?aieeanue i?e?iaeiay ?anoeoaeueiinoue
eoaiai-noaiiiai oeia, eioi?ay caieiaao ieiei 1% ieiuaaee ?aaeiia
enneaaeiaaiee. Ii?aaeaeaii ianoi enneaaeiaaiiuo noaiae ii ioiioaieth e
noaiiui caiiaaaeieeai Oe?aeiu.

Iieo/aiu iieiua aeaiiua i oaeniiiie/aneii ninoaaa oei?u noaiae THEEO –
644 aeaea ec 264 ?iaeia, 51 naiaenoaa e 3 ioaeaeia. Onoaiiaeaiiay
nenoaiaoe/aneay no?oeoo?a (Asreraceae, Poaceae, Fabaceae,
Caryophyllaceae, Lamiaceae, Brassicaceae, Scrophullariaceae, Rosaceae,
Boraginaceae, Apiaceae) io?aaeaao ena?ioe/aneee oa?aeoa? e oyaioaao e
n?aaeecaiiiii?neiio oeio. Niaeo? aeiii?oieiae/aneie no?oeoo?u ii
iniiaiui iieacaoaeyi oa?aeoa?ecoaony i?aiaeaaeaieai o?aayienouo
iieeea?ieeia n iieo?icaoi/iui eee ?icaoi/iui no?iaieai iiaaaia e
eaoaeaenii. A yeieiai-oeaiioe/aneii niaeo?a i?aiaeaaeatho noaiiua e
eoaiai-noaiiua aeaeu oei?u, e eioi?ui a cia/eoaeueiie noaiaie
i?eniaaeeiythony iiooa/iua aeaeu, /oi io?aaeaao noaa?eaeiue,
aaieena?ioeoiue oa?aeoa? oneiaee nouanoaiaaiey, ia?ooaiiinoue ia?ae/iie
no?oeoo?u oeaiicia, ioaeae?iiuo io eeeiaeniaiai ninoiyiey, e
oi?ie?iaaiey eo ia ianoa eania. Yeieiai-oeaiioe/aneee aiaeec oei?u
iieacae cia/eoaeueiia ?aciiia?acea oeiia aeaeiaie aaeaioaoeee e
ioaeaeueiui yeieiae/aneei oaeoi?ai noaiiuo oeaiicia. Aaia?aoe/aneee
aiaeec THEEO naeaeaoaeuenoaoao i oii, /oi a niaeo?a oei?u i?aiaeaaeatho
noan?aaeecaiiiii?nei-noaoaiia?aoiua e noaieaaie/anei-noaeiioeiaioaeueiua
aeaeu. Iieacaii, /oi enneaaeiaaiiay oei?a oa?aeoa?ia aeey ainoi/ii –
aa?iiaeneeo noaiiuo e eaninoaiiuo oei?. Aunieee i?ioeaio yiaeaie/iuo
aeaeia – 11,9 – naeaeaoaeuenoaoao i naiiauoiii oa?aeoa?a oei?u, a oaeaea
i oii, /oi a i?aaeaeao noaiiiai aeiia eioaineaii eaeoo i?ioeannu
aeaeiia?aciaaiey.

Onoaiiaeaii, /oi a caaeneiinoe io oeia ii/au noaiiay ?anoeoaeueiinoue
i?aaenoaaeaia o?aiy eeannaie: ia /a?iic?iao – eeann Festuco-Brometea
(eoaiaua, ?ani?ino?ai?iiua ii anae oa??eoi?ee, e
?aciio?aaii-oei/aeiai-eiaueueiua noaie); ia ea?aiiaoiuo ?aiaeceiao –
eeann Helianthemo-Thymetea (nin?aaeioi/aiu a thai-ainoi/iie /anoe); ia
ian/aiuo ioeiaeaieyo – eeann Festuceta vaginatae (o?aaiaioa?ii ia
ai?iauo oa??anao ?a/iuo aeieei). Ia oei?enoe/aneie iniiaa (ii iaoiaeo
A?aoi-Aeaiea) ?ac?aaioaia eeanneoeeaoeeiiiay noaia ?anoeoaeueiinoe
noaiae THEEO, eioi?ay aeeth/aao 3 eeanna, 10 nithcia, 23 annioeeaoeee e
3 noaannioeeaoeee, ec eioi?uo 2 nithca, 9 annioeeaoeee e 3
noaannioeeaoeee aia?aua iienaiu nieneaoaeai. I?e iiiiue iaoiaeeee
oeoieiaeeeaoeee, a?aaeeaioiiai e i?aeeiaoeeiiiiai aiaeecia, onoaiiaeaii,
/oi aaaeoueie oaeoi?aie oi?ie?iaaiey noaiae yaeythony aeae?i?aaeei e
niaea?aeaiea ea?aiiaoia a ii/aa, eioi?ua iaoneaaeeaatho iniaaiiinoe
noaiiuo ii/a (eeneioiinoue, o?ioiinoue, niaea?aeaiea acioa e o.ae.),
auyaeaia ei??aeyoeey iaaeaeo ieie. Aeieacaii, /oi ?aia nithca aunooiaao
oie neioaeniiiie/aneie eaoaai?eae, eioi?ay io?aaeaao niaoeeoeeo
yeinenoaiu ii eiiieaeno yeieiae/aneeo oaeoi?ia.

Auyaeaiu ianoii?iec?inoaiey 12 aeaeia, aeeth/?iiuo a Aa?iiaeneee E?aniue
nienie, 27 – ec “E?aniie eieae Oe?aeiu” e 34 aeaeia, ioaeaeathueony a
?aaeiiaeueiie io?aia, enneaaeiaaii eo ?ani?ino?aiaiea e aeaia
aoooeoinioeeieiae/aneay oa?aeoa?enoeea. Ia noaiiuo o/anoeao auyaeaiu
niiauanoaa aeanyoe oi?iaoeee, caian?iiuo a “Cae?ioth eieao Oe?aeineie
NN?” (Amygdaleta nanae, Stipeta. lessingianae, S. capillatae, S.
pulcherrimae, S. pennatae, S. dasyphyllae, S. tirsae, S. zalesskii,
Elytrigieta stipifoliae, Cariceta humilis) e ?aaeeea niiauanoaa
oi?iaoeee Androsaceta koso-poljanskii, ioaeaeathueany a io?aia. N
o/anoeai aaoi?a i?aaeeiaeaii nicaeaiea 56 i?e?iaeii-caiiaaaeiuo iauaeoia
iauae ieiuaaeueth ieiei 2,7 oun. aa, eioi?ua iiauoatho
?ai?acaioaoeaiinoue noaiae, iaania/eaatho nio?aiiinoue oeie/iuo e
?aaeeeo aeaeia e niiauanoa.

Eeth/aaua neiaa: thaeiay /anoue Eaaiaa?aaeiie Eaninoaie Oe?aeiu, noaiue,
oei?a, aiaeec, ?anoeoaeueiinoue, oei?enoe/aneay eeanneoeeaoeey,
oeoieiaeeeaoeey yeieiae/aneeo oaeoi?ia, ?aaeeea aeaeu, ?aaeeea
niiauanoaa.

Babko I.A. Steppe plant cover differentiation of the Left-Bank
Forest-Steppe south part of Ukraine. – Manuscript.

Thesis for 1 candidate’s degree by speciallity 03.00.05. – botany. —
M.G. Kholodny Institute of Botany of the National Academy of Sciences of
Ukraine, Kyiv, 1999.

The thesis is dedicated to problem of an evaluation of floristic and
ecology-cenotic diversity of steppe plant cover of the Left-Bank
Forest-Steppe south part of Ukraine (LFSU). About 100 locations (1% of
area) with well conserved original vegetation of meadow-steppe type have
been found. It has been determined that flora composition numbers 644
species which have been assigned to the 264 genera, 51 families and 3
sections. There have been carried out its systematic, biomorphological,
ecology-cenotic and geographical analyses, results of which have
confirmed a relation of investigated flora with ancient Mediterranean
flora. Aclassified scheme of the LFSU steppe vegetation on a floristic
basis (by the Braun-Blanguet method) has been worked up. It includes 3
classes, 10 alliances, 23 associations and 3 subassociations. Besides, a
description was made of the 2 alliances, 9 associations and 3
subassociations by the author. It has been established that the key
factors of steppe shaping are hydroregime and the carbonates content of
the soil. It has been proved that an alliance range is a such
syntaxonomic categary as reflects an ecosystem peculiarity for complex
of environmental factors. 12 species found are enrolled in the European
Red List, 27 spesies – in the Red Data Book of Ukraine, 34 ones need
regional protection, 10 formations are listed in the Green Data Book of
Ukraine. It has been proposed to form 56 protected subjects a total area
of which are ca 2,7 th ha with the participation of the author.

Key words: Left-Bank Forest-Steppe south part of Ukraine, steppe, flora,
analysis, vegetation, floristic classification, phytoindication of
environmental factors, rare species, rare communities.

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