1

IAOe?IIAEUeIA AEAAeAI?ss IAOE OE?A?IE

?INOEOOO A?IEIA?? I?AAeAIIEO II??A ?i. I.I.EIAAEAANUeEIAI

IAO?INssI AIIA AAI?A??AIA

OAeE 502.55:551.35 (262.5)

A?IOANOOAAIIss II?NUeEI? AIAeE OA AeIIIEO

A?AeEEAAe?A I?AI?*II-CAO?AeII? *ANOEIE

*I?IIAI II?ss

03.00.17 — a?ae?ia?ieia?y

Aaoi?aoa?ao

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Naaanoiiieue — 1998

Aeena?oaoe??th ? ?oeiien.

?iaioo aeeiiaii a Iaeanuee?e o?e?? ?inoeoooo a?ieia?? i?aaeaiieo ii??a
?i. I.I. Eiaaeaanueeiai IAI Oe?a?ie

Iaoeiaee ea??aiee:Aeaaeai?e IAI Oe?a?ie,

aeieoi? a?ieia?/ieo iaoe, i?ioani?

Caeoeaa THaaiae?e Iao?iae/,

Iaeanueea o?e?y ?inoeoooo a?ieia?? i?aaeaiieo ii??a IAI Oe?a?ie,

aieiaiee iaoeiaee ni?a?ia?oiee.

Io?oe?ei? iiiiaioe:Aeieoi? a?ieia?/ieo iaoe, i?ioani?

Ie?iiia Ieaa Ae?aiae/,

?inoeooo a?ieia?? i?aaeaiieo ii??a IAI Oe?a?ie,

caa. a?aeae?eii ii?nueei? nai?oa?ii? a?ae?ia?ieia??;eaiaeeaeao
aaie.-i?i. iaoe

Niaaa Ieaia ?aaaiiaia,

Ii?nueeee a?ae?io?ce/iee ?inoeooo IAI Oe?a?ie,

noa?oee iaoeiaee ni?a?ia?oiee

I?ia?aeia onoaiiaa -Iaeanueeee aea?aeaaiee oi?aa?neoao
?i.?.?.Ia/ieeiaa, a?ieia?/iee oaeoeueoao, i.IaeanaCaoeno a?aeaoaeaoueny
“_28_”_a?oaeiy_ 1998 p. i__14.00_ aiaeei? ia can?aeaii? niaoe?ae?ciaaii?
a/aii? ?aaee Ae 50.214.01 ?ino?oooo a?ieia?? i?aaeaiieo ii??a
?i.I.I.Eiaaeaanueeiai IAI Oe?a?ie ca aae?anith 335011, i.Naaanoiiieue,
i?.Iao?iiaa, 2.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? ?ino?oooo a?ieia??
i?aaeaiieo ii??a ?i.I.I.Eiaaeaanueeiai IAI Oe?a?ie: 335011,
i.Naaanoiiieue, i?.Iao?iiaa, 2.

Aaoi?aoa?ao ?ic?neaiee “_2_”__a?oaeiy_ 1998 ?.

A/aiee nae?aoa? niaoe?ae?ciaaii? a/aii? ?aaee Ae50.214.01

eaiaeeaeao a?ieia?/ieo iaoe
I.E. Iaa?iaa

caaaeueia oa?aeoa?enoeea ?iaioe

Aeooaeuei?noue i?iaeaie. Aieiaiith iaoith aea/aiiy ? eiio?ieth
caa?oaeiaieo aiaeieo aeinenoai ? ioe?iea ?nioth/i? oa iioaioe?eii?
iaaaciaee oeo ci?i, ui a?aeaoaathoueny i?ae aieeaii oiene/ieo oaeoi??a
oa i?aaenoaaeythoue caa?ico aeeooth ia Caie? a oe?ieiio ?icoi?ii?
oeueiai neiaa (A?aa?inueeee, 1995).

Ieoaiiy i?i iaiao?aei?noue oi?o?eaoe?? a?ieia?/ieo iaoiae?a ioe?iee
yeino? aiaee aoei iinoaaeaii a?eueoa 10 ?ie?a oiio (Oieoeoee?iaaiiua
iaoiaeu, 1983), aea ua e nueiaiaei? aiii aeaeaei a?ae ae??oaiiy.
Iacaaaeath/e ia oea, a?i?iaeeeaoe?y ? a?ioanooaaiiy oe?iei
aeei?enoiaothoueny o iii?oi?eiao caa?oaeiaiiy a?ae?inoa?e (Ao?aeei,
1982; Aeyoeia e ae?.,1994; Malins, 1989; Stebbing et al., 1980 ).

I?ae eiioeaio?aoe??th caa?oaeithth/eo ?a/iaei o aiaeieiao oa ?oiueith
oiene/i?noth aeey a?ae?ia?iio?a iaia? iaeiicia/iiai ca`yceo. Aeyaeoe
iioaioe?eio iaaaciaeo aeey aeeaeo i?aai?ci?a caa?oaeithth/eo ?a/iaei, ui
?ic/eiai? o aiae? /e aaeni?aiaai? o aeiiieo a?aeeeaaeao iiaeia o?eueee c
aeei?enoaiiyi niaoe?aeueieo a?ieia?/ieo iaoiae?a. O ii?aeiaii? c
iaoiaeaie aiae?oe/ii? o?i??, iaoiaee a?ioanooaaiiy aeicaieythoue ye?nii
? e?euee?nii ioe?ieoe aoaeoe, ye? aeieeathoue o caa?oaeiai?e aeinenoai?.

I?eioeeiiaith a?aei?iith a?ioanooaaiiy a?ae ?ioeo a?ieia?/ieo iaoiae?a
ioe?iee yeino? aiaeiiai na?aaeiaeua ? aeei?enoaiiy /ooeeaeo eaai?aoi?ieo
eoeueoo? oano-ia`?eo?a ??ciiai nenoaiaoe/iiai oa o?io?/iiai ??ai?a. Aei
oiai ae, c on?o a?ieia?/ieo iaoiae?a ioe?iee yeino? iaaeieeoiueiai
na?aaeiaeua naia a?ioanooaaiiy iaeeaaoa i?aeaea?oueny noaiaea?oecaoe??.
E?euee?noue iaoe?iiaeueieo, ?aa?iiaeueieo oa i?aeia?iaeieo noaiaea?oieo
iaoiae?a a?ioanooaaiiy iaooeeueii c?inoa?. Iaeacii I?i?no?a ioi?iie
iaaeieeoiueiai i?e?iaeiiai na?aaeiaeua oa yaea?ii? aaciaee Oe?a?ie
caoaa?aeaeaia oe?ea iecea ea??aieo ii?iaoeaieo aeieoiaio?a c
a?ioanooaaiiy ii?nueeeo oa i??nieo aiae (EIAe 211.1.4.047-95, EIAe
211.1.4.055-97, EIAe 211.1.4.054-97).

Iayai?noue noaiaea?oieo iaoiae?a a?ioanooaaiiy ?icoe?th? iiaeeeaino? ?o
i?aeoe/iiai canoinoaaiiy, ui aaea a?aeia?aaeaii o ii?iaoeaieo
aeieoiaioao. Oae, iai?eeeaae, Iinoaiiaa EI Oe?a?ie a?ae 11 aa?aniy 1996
?ieo ? 1100 ia?aaeaa/a?, ui i?e ?ic?iaoe? a?aie/ii aeiionoeieo neeae?a
(AAeN) iaia`yceiai iathoue aooe o?aoiaai? ?acoeueoaoe a?ioanooaaiiy. C 1
n?/iy 1999 ?ieo aoaea aaaaeaiee o ae?th I?aea?aeii/ee ii?iaoeaiee
aeieoiaio “Iaoiaeeea ioe?iee aeieia?/ii? yeino? iiaa?oiaaeo aiae ca
a?aeiia?aeieie eaoaai??yie”, o yeiio /?oei iicia/aia noa?a aeei?enoaiiy
?acoeueoao?a a?ioanooaaiiy.

Ana oea ciaia`yco? aeine?aeiee?a ?ic?iaeyoe iia?, ua a?eueo /ooeea? oa
iia?aoeai? iaoiaee ioe?iee oiene/iino? aiaee oa aeiiieo a?aeeeaae?a, a
oaeiae ie?aieo caa?oaeithth/eo ?a/iaei oa ?o noi?oae, ye? neeaeathony o
aiaeieie.

Iaoa oa caaaeaiiy aeine?aeaeaiiy. Iaoith aeena?oaoe?? aoea ?ic?iaea oa
ai?iaaoe?y iaoiae?a a?ioanooaaiiy aiaee, aeiiieo a?aeeeaae?a ? ie?aieo
caa?oaeithth/eo ?a/iaei a oiiaao i?ai?/ii-cao?aeii? /anoeie *i?iiai
ii?y. O a?aeiia?aeiino? c iaoith aeine?aeaeaiiy aoee iinoaaeai? oae?
caaaeaiiy:

— ?ic?iaeoe iaoiaee a?ioanooaaiiy ii?nueeiai na?aaeiaeua noiniaii aei
oiia i?ai?/ii-cao?aeii? /anoeie *i?iiai ii?y;

— anoaiiaeoe ae?aiacii eieeaaiue nieiiino?, yeee iiaeooue aeo?eioaaoe
eaai?aoi?i? oano-ia`?eoe o?eaaeee /an;

— ioe?ieoe ye?noue ii?nueei? aiaee a Iaeanuee?e caoioe? oa i?eeaaeeo
ae?eyieao ii?y c iaoith aeyaeaiiy iaea?eueo e?oiieo aeaea?ae
caa?oaeiaiiy;

— aecia/eoe oiene/i?noue ii?nueeeo aeiiieo a?aeeeaae?a o aeaaoi??yo c
iaiaeiaeiaith i??ith aio?iiiaaiieo iaaaioaaeaiue;

— i?iaanoe aenia?oio ioe?ieo caa?oaeithth/eo ?a/iaei (neio/eo aaioaae?a,
ieth/eo cania?a) aeey ioe?iee iiaeeeaeo iane?aee?a ?o iaaeoiaeaeaiiy o
ii?nueea na?aaeiaeua.

Iaoeiaa iiaecia io?eiaieo ?acoeueoao?a. Aia?oa ca aeiiiiiaith eiiieaena
iaoiae?a ? oano-ia`?eo?a aoea i?iaaaeaia ioe?iea oiene/iino? i?e?iaeii?
ii?nueei? aiaee, no?/ieo aiae ??ciiai iioiaeaeaiiy, ye? neeaeathoueny o
i?ai?/ii-cao?aeio /anoeio *i?iiai ii?y, ie?aieo caa?oaeithth/eo ?a/iaei,
aeiiieo a?aeeeaae?a o caa?oaeiaieo ii?nueeeo aeaaoi??yo, a oaeiae
aeaye?o neio/eo aaioaae?a, ui ia?aaicyoueny o aaeeeeo ianyaao.

Anoaiiaeai? naciii? ? i?inoi?ia? eieeaaiiy ??aiy oiene/iino? i?e?iaeii?
ii?nueei? aiaee ? aeiiieo a?aeeeaae?a o Iaeanuee?e caoioe? oa i?eeaaeeo
ae?eyieao ii?y. Ia i?aenoaa? aenia?eiaioaeueieo niinoa?aaeaiue
cai?iiiiiaaia oeaea aecia/aiiy oiene/iinoe na?aaeiaeu, ye? i?aeeyaathoue
a?ioanooaaiith. Aia?oa ca ?acoeueoaoaie a?ioanooaaiiy anoaiiaeai?
“aa?y/? oi/ee”, o yeeo oiene/i?noue caa??aaeanue i?ioyaii o?eaaeiai
/ano.

I?aeoe/ia cia/aiiy io?eiaieo ?acoeueoao?a. I?aenoieii aeine?aeaeaiue
caeiaoaa/a ? ?ic?iaeai? aeey ii?nueeeo aiae c nieii?noth, yea
ci?ith?oueny, ? ai?iaiaai? c caeo/aiiyi aaeeeiai oaeoe/iiai iaoa??aea
iaoiaee a?ioanooaaiiy ii?nueei? aiaee ? aiaeieo aeno?aeo?a aeiiieo
a?aeeeaae?a, o oiio /ene? e iaoiaee ioe?iee oiene/iino? ia eaai?aoi?ieo
eoeueoo?ao Phaeodactylum tricornutum Bochlin oa Artemia salina L. Oe?
iaoiaee aeeth/ai? aei ea??aiiai ii?iaoeaiiai aeieoiaioo EIAe
211.1.4.047-95 “Aiioanooaaiiy ii?nueei? aiaee oa noi/ii?, yea
aiaeaiaeeoueny a ii?a”, caoaa?aeaeaiee iaeacii I?ii?e?iaee Oe?a?ie ? 46
a?ae 30 o?aaiy 1995 ?.

Cai?iiiaaiee aaoi?ii eiiieaen iaoiae?a aeey ioe?iee oiene/iino? aiaee,
no?/ieo aiae ??ciiai iioiaeaeaiiy, ie?aieo caa?oaeithth/eo ?a/iaei oa
aeiiieo a?aeeeaae?a, ye? i?aaenoaaeythoue iioaioe?eio iaaaciaeo aeey
ii?y, a oaeiae aeai? i?i ?ioaa?aeueio oiene/i?noue aiaee oa aeiiieo
a?aeeeaae?a aeei?enoiaothoueny Aea?ae?iniaeoe??th ioi?iie *i?iiai ii?y o
niinoa?aaeaiiyo ca ye?noth ii?nueeiai na?aaeiaeua.

Iniaenoee aianie aaoi?a. Aeena?oaoe?eia ?iaioa ? naiino?eiei iaoeiaei
aeine?aeaeaiiyi, o yeiio aaoi?ii aoea ?ic?iaeaia iaoiaeieia?y
aeine?aeaeaiiy, i?iaaaeai? eaai?aoi?i? aenia?eiaioe, aiae?c,
ocaaaeueiaiiy io?eiaieo aeaieo oa ?o ni?anoaaeaiiy c e?oa?aoo?ieie
aeaea?aeaie.

Ai?iaaoe?y ?iaioe. Iniiai? iieiaeaiiy aeena?oaoe?? aeiiia?aeaeeny ?
iaaiai?thaaeeny ia I-io oa II-io c`?caeao oiaa?enoaa a?ae?iaeieia?a
Oe?a?ie, iaoeiai-i?aeoe/ieo eiioa?aioe?yo “Aeieia?/i? i?iaeaie
Iaeanueeiai ?aa?iio oa ?o ae??oaiiy” (Iaeana, 1995), “Aiaea ? caei?ia`y”
(Iaeana, 1998), ia i?aenoieiaeo nan?yo IO ?iAII IAI Oe?a?ie oa
Oe?a?inueeiai iaoeiaiai oeaio?o aeieia?? ii?y.

Ioaeeeaoe??. Ca iaoa??aeaie aeena?oaoe?? iioae?eiaaii 15 ?ia?o.

No?oeoo?a oa ianya ?iaioe. Aeena?oaoe?y aeeeaaeaia ia 179 noi??ieao
ae?oeiaaiiai oaenoo ? neeaaea?oueny c anooio, iaeyaeo e?oa?aoo?e, 6
?icae?e?a aeanieo aeine?aeaeaiue, cae?i/aiiy oa aeniiae?a. Ia?ae?e
aeei?enoaii? e?oa?aoo?e aeeth/a? 210 aeaea?ae. ?iaioa i?noeoue 12
oaaeeoeue oa 32 ?enoiee.

ci?no ?iaioe

A?ieia?/i? iaoiaee eiio?ieth yeino?

ii?nueeiai ca?aaeiaeua. Iaeyae e?oa?aoo?e

Aeieeaaeii aena?oeai? iaoiaee oa i?aeoiaee aei ioe?iee oiene/iino?
ii?nueei? aiaee, aeiiieo a?aeeeaae?a oa ie?aieo caa?oaeithth/eo ?a/iaei
c aeei?enoaiiyi ieaieoiiieo aiaei?inoae, ?aeiiiae?aieo oa iaei?ino?oeo.
I?eaiaeyoueny e?oa?aoo?i? aeai? uiaei e?eoa???a aeai?o oano-ia`?eo?a ?
oano-ooieoe?e aeey a?ioanooaaiiy ii?nueeiai na?aaeiaeua.

Ia`?eoE OA iaoiaeE a?ioanoOaaiIy ii?nUeei? aiaeE,

aeiiiEo A?AeEEAAe?A ? iE?AIEO caA?OAeITHTH*EO ?A*IAEI

I?eaaaeai? iniiai? i?eeiie ? iaoiaee eaai?aoi?iiai oo?eiaiiy eoeueoo?
ii?nueeeo ieaieoiiieo aiaei?inoae ? ?iooci??e, io?eiaiiy oa aeei?enoaiiy
neio?ii?ciaaieo iaoie?on?a a?oai?? ia ?aii?o noaae?yo ?icaeoeo.
?ic?iaeai? oiiae, i?ioeaaeo?a a?ioanooaaiiy ? iieacieee oiene/iino? aeey
i?iaaaeaiiy aenia?eiaioaeueii? ioe?iee ii?nueeeo i?e?iaeieo aiae,
no?/ieo aiae i??niiaiaeiiai iioiaeaeaiiy, ye? neeaeathoueny a ii?a,
caa?oaeiaieo aeiiieo a?aeeeaae?a oa ie?aieo caa?oaeithth/eo ?a/iaei.
Cai?iiiiiaaia oeaea aecia/aiiy oiene/iino? na?aaeiaeu, ye? i?aeeyaathoue
a?ioanooaaiith (oaae. 1).

Oaaeeoey 1

Oeaea aecia/aiiy oiene/iino? oanoo?iiai

na?aaeiaeua aeey aiaei?inoae oa ?aeiiiae?aieo

Ph. TricornutumA. salinaIieacieee

a?ioanoo-aaiiy,

% a?ae

eiio?iethOa?aeoa-?enoeea

na?aaeiaeuaAaee

oie-ne/-iino?Iaiao?aeia e?aoi?noue

picaaaeaiiyAaee

oie-ne/-iino?Iaiao?aeia e?aoi?noue

?icaaaeaiiy0-10Eaoaeueia-3 >1:30511-24 1:16-1:304>1:2525-49Oiene/ia-2
1:9-1:1531:11 — 1:2550-74 1:2-1:721:3 — 1:1075-89Neaai-oiene/ia-1
1:111:1 — 1:290-110Iaoiene/ia0-0-111-150Noeioeyoe?y

ae?eaiiy ee?oei

aiaei?inoae+1 1:1-1:3—151-300+2>1:3—>301+3—Ai?iaiaai? iaoiaee
ia?aa??ee /ooeeaino? eoeueoo? ii?nueeeo eaai?aoi?ieo oano-ia`?eo?a c
aeei?enoaiiyi noaiaea?oiiai oieneeaioa a?o?iiaoo eae?y. Aeai? i?i
/ooeea?noue eaai?aoi?ieo oano-ia`?eo?a aei a?o?iiaoo eae?y i?eaaaeai? o
oaae. 2.

sse iieacieee /ooeeaino? oano-ia`?eo?a cai?iiiiiaaii aeei?enoiaoaaoe
ii??a /ooeeaino? iaoiaea a?ioanooaaiiy, a oaeiae iaae?aiio aoaeoeaio
(EC5024/) /e iaae?aiio eaoaeueio eiioeaio?aoe?th noaiaea?oiiai
oieneeaioa a?o?iiaoo eae?y.

Aenia?eiaioaeueii anoaiiaeaia ia??iaee/i?noue ia?aa??ie eoeueoo?
aiaei?inoae (Ph. tricornutum) oa ?iooci??e (Euplotes patella lemani
Dragesco).

I?eaaaeai? aeai? i?i i?aeaeuaiiy /ooeeaino? a?oai?? i?ae /an ia?aoiaeo
c? noaae?? i?oiiaoie?ono aei noaae?? iaoaiaie?ono, a oaeiae i?i ??ciee
ae?aiacii /ooeeaino? aeaio ?an a?oai?? (neaaonueei? oa eoyeueieoeueei?)
aei noaiaea?oiiai oieneeaioo.

Oaaeeoey 2

*ooeea?noue eaai?aoi?ieo oano-ia`?eo?a aei a?o?iiaoo eae?y

Oano-ia`?eoOano- ooieoe?yYenii-ceoe?y,

aiaeAe?aiacii

eiioeaio?aoe?e,Iieacieee

/ooeeaino?ia/eAN0,

ia/eAN50,

ia/eLN50,

ia/ePh.

Tricornutum??no

eoeueoo?240,1-10,00.252,50,5A.salina

(eoyeueieoeeay)Aeaee-aaiiy240,5-40,08,0027,73,6A. salina

(neaaoneay)Aeaee-aaiiy244,0-10,05,007,61,8E.
patellaAeaee-aaiiy2410,0-140,012,5072,510,6

Oaeoi? nieiiino? a a?ioanooaaii?

no?/ieo aiae, ye? neeaeathoueny A II?A

Aenia?eiaioaeueii anoaiiaeai? cia/aiiy nieiiino?, ye? aeicaieythoue
io?eioaaoe no?ee? eoeueoo?e ia i?ioyc? 60-oe oa a?eueoa ae?a o
noaiaea?oieo oiiaao. A ae?aiacii? nieiiino?, yeee ? oa?aeoa?iei aeey
?ici??niaii? i?ai?/ii-cao?aeii? /anoeie *i?iiai ii?y, eoeueoo?e
ieaieoiiieo aiaei?inoae Ph. tricornutum caa??aathoue aeeoo?ae?yeuei?noue
? aenieee eiao?oe??io ae?eaiiy o ae?aiacii? 14-16. I?e a?ioanooaaii?
nieiiiaaoiaiaeieo aiae o ae?aiacii? nieiiino? 5-8 ? aeioe?eueiith
aee?iaoe?y aiaei?inoae ia na?aaeiaeuao c a?aeiia?aeiith nieii?noth.

Ne?ae a?aoiaoaaoe, ui i?e cia/aiiyo nieiiino? ieae/a 5 eoeueoo?e Ph.
tricornutum caa??aathoue aeeoo?caeaoi?noue i?ioyaii iaiaaeaiiiai /ano
(a?ey 16 ae?a) ?, ca aecia/aiiyi A.A.Oeaaiae/a (1974), oaea na?aaeiaeua
aeey eoeueoo? Ph. tricornutum iiaeia aaaaeaoe “ciiith oei/aniai?
aaeaioaoe??”.

Eaai?aoi?i? eoeueoo?e ?iooci??? E. patella lemani oni?oii aee?iothoueny
aei ae?aiaciio nieiiino? 5-20, iaeiae iaeneiaeuei? cia/aiiy
eiao?oe??io?a ae?eaiiy a?aeiia?aeathoue nieiiino? 5-10.

Nieiiiniee iioeioi aeey ii?iaeueiiai ?icaeoeo A. salina ia noaae?yo
i?oiiaoie?ona ? iaoaiaoie?ona i?ioyaii 72 aiaeei ia ciaioo?oueny
ieae/a 10. I?iaeaia nieiiinieo aaeaioaoe?e aeey A. salina ia ?
nooo?aith, ai oeae ia`?eo ia i?aeo?eio?oueny o?eaaeee /an o eoeueoo??, a
i?e iaiao?aeiino? iaoie?one iiaeia eaaei io?eiaoe c y?oeue.

Aenia?eiaioaeueia ioe?iea ssEINO? aiae

IaeanUeei? caOIEE OA i?eeaaEeo Ae?EssIIE ii?y

I?eaaaeai? ?acoeueoaoe aeine?aeaeaiue oiene/iino? aiae Iaeanueei?
caoiee a 1993-1996 ??., ye? i?iaaaeai? c iaoith aeyaeaiiy iaea?eueo
iaaeaaiiieo/ieo ae?eyiie aeaaoi???. Oiene/i?noue aiaee ioe?ithaaee a
aaeao ca oeaeith oiene/iino?.

Aeine?aeaeaiiy ia?oiai aoaio, ye? aoee i?iaaaeai? o ea?oi?-aa?ani? 1993
?., iieacaee iayai?noue cia/ieo eieeaaiue iieaciee?a ?inoo o
oano-eoeueoo?ao aiaei?inoae iaa?oue o iaaeao iaei??? noaioe?? o i?iaao,
ui aoee a?ae?a?ai? c ??cieo ai?eciio?a (iiaa?oiaaiai ? i?eaeiiiiai).
Iaeiae aiae?c yeino? i?e?iaeii? aiaee, ui aoa ae?aaeaiee o aaeao
oiene/iino?, iieacaa iayai?noue caaaeueieo iciae, ye? iiaeooue
ioa?aeoa?ecoaaoe aeaaoi??th a oe?eiio (oaae. 3). Oae, i?iae aiaee, ye?
aoee a?ae?a?ai? o aeaaoi??? caoiee a ea?oi?, ni?e/eiyee noeioethth/o
ae?th ia ?inoia? iieacieee eoeueoo? aiaei?inoae, oiae? ye o /a?ai?
a?eueo?noue i?ia aoee iaoiene/ieie, a a na?ii? on? i?iae ca oeaeith
oiene/iino? aoee a?aeianai? aei eaoaai??? iaoiene/ieo aai
neaaioiene/ieo.

Oaaeeoey 3

Oiene/i?noue aiaee Iaeanueei? caoiee a ea?oi?-aa?ani? 1993 ?.

(a /enaeueieeo — iiaa?oiaaee ai?eciio, a ciaiaiieeo — i?eaeiiiee
ai?eciio, e?euee?noue noaioe?e a % a?ae caaaeueii? ?o e?eueeino?)

Aeaoa, eiee a?aeae?aeeny

I?iaeAaee oiene/iino? -10+1+2+328.04.1993-0

16,716,7

16,783,3

500

16,725.05.199350

10033,3

016,7

0—28.06.1993-66,6

87,533,3

16,7—28.07.199314,3

042,9

85,728,6

14,314,3

0-29.08.199342.9

57,157,1

42,9—30.09.19930

14,328,6

28,671,4

57,1—I?aenoiee oanoe aan?iiuei-in?ii?o ceiiie, aeicaieeee aeyaeoe
“aa?y/? oi/ee” — ae?eyiee aeaaoi???, ye? ciaoiaeyoueny i?ae iino?eiei
i?anii caa?oaeiaiiy, aea i?e?iaeia ii?nueea aiaea iiaea aooe
ioa?aeoa?eciaaia ye oiene/ia ca aeaieie a?ioanooaaiiy ? ia yeeo
eiioeaio?aoe?? iiethoaio?a aeua ca oiiia?, /e aiie ciaoiaeyoueny o
iaaacia/ieo niieo/aiiyo, ui aeeeeea? a?ieia?/i? aoaeoe caa?oaeiaiiy. Ca
ia??iae niinoa?aaeaiue (ea?oaiue-aa?anaiue 1993 ?.) aaniethoii
aeaaiiieo/ieo ae?eyiie a aeaaoi??? Iaeanueei? caoiee ia aoei.

“Aa?y/? oi/ee” oa i?eeaae? aei ieo ae?eyiee aeaaoi??? aoee aeeth/ai? o
ceiieo ae?oaiai aoaio, yea aoea i?iaaaeaia 1-3.06.1994 ?. Aei eeano
neaaioiene/ieo ca oeaeith oiene/iino? aeey aiaei?inoae a?aeianai?
i?eaeiii? ai?eciioe aiaee a ?aeii? Iaeanueeiai ii?oo (Iaooiaaaaiue,
ciai?oiee ?aeae ii?oo), i?noea aeioneo no?/ieo aiae c NAI “I?ai?/ia” oa
i?eeaae? aei ia? aeaaoi???. Oiene/ieie ca oeaeith oiene/iino? aoee
i?iae aiaee, ye? a?ae?a?ai? o ii?? a i?eaeiiiiio ai?eciio? a ?aeii?
A?eaae?enueeiai aaa??eiiai ceeaiaiai eieaeoi?a (iieacieee ?inoo
aiaei?inoae neeaaeaee 59,2 % a?ae io eiio?ieth). A?aeiiniei
aeaaiiieo//yi ia iiiaio aeine?aeaeaiiy a?ae??ciyany ?aeii aeioneo ?
?icn?thaaiiy no?/ieo aiae c NAI “I?aaeaiia”. Ae?eyiee aeaaoi??e, ye?
aoee a?aeianai? aei o?ueio eean?a ca oeaeith oiene/iino?:
neaaioiene/iiio, iaoiene/iiio ? oaeiio, yeee aeeeeea? noeioeyoe?th
ae?eaiiy aiaei?inoae, a ia??iae niinoa?aaeaiue ?iciiae?eyeeny a
Iaeanuee?e caoioe? oa i?eeaaeeo aeaaoi??yo ieyienoi ? ia aoee i?eo?i/ai?
aei a?aeiia?aeiiai ai?eciioo /e ?aeiio.

Iie?aiii? ?iaioe, ye? aoee i?iaaaeai? o eeii? 1994 ?., aeicaieeee
iaeii/anii ioe?ieoe oiene/i?noue aiaee o caieiaieo aaaaiyo oa ia
ciai?oiueiio ?aeae? Iaeanueeiai ii?oo (iie?aii “Ii?o”) ii??aiyii c
a?aeiinii /enoith aeaaoi???th a ?aeii? a?inoaioe?? Iaeanueeiai
aea?aeoi?aa?neoaoo (iie?aii “Iaeanueea caoiea”) ? c a?aeaeaeaiei a?ae
aa?aaiaeo aeaea?ae caa?oaeiaiiy iie?aiiii “Iaeanueea aaiea”. I?iae
aiaee c iie?aiio «Iaeanueea aaiea» aeeeeeaee ia?aaeuaiiy ?inoiaeo
iieaciee?a a eoeueoo?ao ii??aiyii c eiio?ieai. Iie?aii “Ii?o”
oa?aeoa?ecoaaany a?eueoei ae?aiaciiii eieeaaiue iieaciee?a ?inoo a
eoeueoo?ao aiaei?inoae, ui aeicaieeei c?iaeoe aeniiaie i?i ieyienoee
oa?aeoa? ?iciiae?eo aiae c oiene/ieie aeanoeainoyie a caieiaieo aaaaiyo
Iaeanueeiai ii?oo.

Noaioe?y i/enoee aaeanoieo aiae (NIAA), ?icoaoiaaia ia oa?eoi???
Iaeanueeiai ii?oo, a?aeianaia aei oi/a/ieo aeaea?ae caa?oaeiaiiy oa e
ia i?ioyc? aaaaoueio ?ie?a iii?oii aieeaaea ia caa?oaeiaiiy Iaooiaaaai?
? Iaeanueei? caoiee iaooiaeie aoaeaaiaeiyie (Aeyoeia, Iao?inyi, 1994).
Aeine?aeaeaiiy c ioe?iee yeino? i/enoee no?/ieo aiae NIAA aoee
i?iaaaeai? o ea?oi? 1993 ?., eiee NIAA i?aoethaaea ua c i?i?iaeueiei
iaaaioaaeaiiyi. Eiioeaio?aoe?y iaooiaeo aoaeaaiaei?a a no?/ieo aiaeao
NIAA neeaaeaea 0,19-1,0 ia/e. E?eoa???i oiene/iino? no?/ieo aiae NIAA
aoea aaee/eia i?i?iaeueii? e?aoiinoe ?icaaaeaiiy, yea iaiao?aeia aeey
oiai, aae i?iaa, yea i?aeeyaa? oanooaaiith ia aeeeeeaea a??ia?aeieo
a?aeoeeaiue a?ae eiio?ieueieo iieaciee?a o oano-ia`?eo?a.

Ieaieoiii? aiaei?ino? Ph. tricornutum aeyaeeeny iaea?eueo /ooeeaei
oano-ia`?eoii aei iaooiaiai caa?oaeiaiiy. C iaoith onoiaiiy oiene/ii?
ae?? aiaee c ?aca?aoa?a-iaeiie/oaa/a ia aiaei?ino?, ?? o?aaa aoei
?icaaaeoe /enoith ii?nueeith aiaeith o 31 ?ac (?en.1). Aeanyoee?aoiiai
?icaaaeaiiy i?iae c ?aca?aoa?a — iaeiie/oaa/a aeyaeeiny aeineoue aeey
oiai, uia ?inoia? iieacieee eoeueoo? ?iooci??e A. patella ia
a?ae??ciyeeny a?ae eiio?ieth (?en. 1).

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a00040000002d01080004000000f00106000400000004010d00040000000201020007000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?en.1. Aeeiai?ea

c?inoaiiy /enaeueiino? Ph.tricornutum a caeaaeiino? a?ae e?aoiino?
?icaaaeaiiy no?/ieo aiae NIAA, ye? i?noyoue iaooo.
picscalex28010009000003440c00000d002d00000000000500000009020000000005000
0000102ffffff000400000004010d000400000002010200050000000c026c041c0703000
0001e00040000002701ffff030000001e00040000002701ffff050000000b02000000000
30000001e00050000000102ffffff000500000009020000000012000000fb02c4ff00000
00000009001000000cc00000020417269616c2043797200cc00040000002d01000004000
0002d010000040000002d01000012000000fb02c9ff0000000000009001000000cc00000
020417269616c2043797200cc00040000002d010100040000002d010000040000002d010
000040000002d010000040000002d01010012000000fb02bfff000000000000900100000
0cc00000020417269616c2043797200cc00040000002d010200040000002d01000004000
0002d010000040000002d01000010000000fb021000070000000000bc02000000cc01020
22253797374656d00cc040000002d010300040000002701ffff030000001e00040000002
d010000050000000102ffffff00050000000902000000000700000016046c041c0700000
000040000002d010300040000002701ffff030000001e00040000002d010000050000000
102ffffff00050000000902000000000700000016046c041c0700000000040000002d010
300040000002701ffff030000001e00040000002d010000050000000102ffffff0005000
000090200000000070000001604620412070a000a0007000000fc0200008080800000000
40000002d01040009000000fa02050000000000000000002200040000002d01050004000
00004010d0004000000020102000e00000024030500330184036a015a03bd065a0386068
4033301840307000000fc020100000000008020040000002d0106000e000000240305003
3018403330152006a0128006a015a03330184030e000000240305006a015a036a012800b
d062800bd065a036a015a03070000001504740172022c00460109000000fa02000000000
000000000002200040000002d01070007000000fc020000ffffff000000040000002d010
800040000002d010300040000002701ffff030000001e00040000002d010000040000002
d010600040000002d010500050000000102ffffff0005000000090200000000070000001
604620412070a000a0007000000fc020100000000008020040000002d01090004000000f
001060009000000fa02000000000000000000002200040000002d01060004000000f0010
5000400000004010d0004000000020102000e00000024030500bd065a038606840333018
4036a015a03bd065a0309000000fa02050000000000000000002200040000002d0105000
4000000f0010600070000001504740172022c0046010e000000240305003301840333015
2006a0128006a015a03330184030e000000240305006a015a036a012800bd062800bd065
a036a015a03040000002d010700040000002d010800040000002d010300040000002701f
fff030000001e00040000002d010000040000002d010900040000002d010500050000000
102ffffff0005000000090200000000070000001604620412070a000a0009000000fa020
00002000000000000002200040000002d01060004000000f001050007000000fc0200008
08066008020040000002d01050004000000f00109000400000004010d000400000002010
2000e00000024030500b5017703b501ab02cb019a02cb016703b501770307000000fc020
000ffffcc008020040000002d01090004000000f00105000e00000024030500740177037
401ab02b501ab02b50177037401770307000000fc020000000000000000040000002d010
50004000000f001090007000000fc020000bfbf99008020040000002d0109000e0000002
4030500b501ab02cb019a028a019a027401ab02b501ab0209000000fa020000020000000
00000002200040000002d010a0004000000f001060007000000fc0200000080800000000
40000002d01060004000000f00109000e00000024030500f6017703f60163020c0253020
c026703f601770307000000fc02000000ffff008020040000002d0109000e00000024030
500b5017703b5016302f6016302f6017703b5017703040000002d01050007000000fc020
00000bfbf008020040000002d010b000e00000024030500f60163020c025302cb015302b
5016302f601630209000000fa02000002000000000000002200040000002d010c0004000
000f0010a0007000000fc020000808066008020040000002d010a0004000000f0010b000
e00000024030500980277039802ab02ae029a02ae0267039802770307000000fc020000f
fffcc008020040000002d010b0004000000f0010a000e00000024030500570277035702a
b029802ab029802770357027703040000002d01050004000000f0010b0007000000fc020
000bfbf99008020040000002d010a000e000000240305009802ab02ae029a026d029a025
702ab029802ab0209000000fa02000002000000000000002200040000002d010b0004000
000f0010c00040000002d01060004000000f0010a000e00000024030500d9027703d902a
600ef029500ef026703d9027703040000002d0109000e00000024030500980277039802a
600d902a600d902770398027703040000002d01050007000000fc02000000bfbf0080200
40000002d010a000e00000024030500d902a600ef029500ae0295009802a600d902a6000
9000000fa02000002000000000000002200040000002d010c0004000000f0010b0007000
000fc020000808066008020040000002d010b0004000000f0010a000e000000240305007
b0377037b03ab0291039a02910367037b03770307000000fc020000ffffcc00802004000
0002d010a0004000000f0010b000e000000240305003a0377033a03ab027b03ab027b037
7033a037703040000002d01050004000000f0010a0007000000fc020000bfbf990080200
40000002d010a000e000000240305007b03ab0291039a0251039a023a03ab027b03ab020
9000000fa02000002000000000000002200040000002d010b0004000000f0010c0004000
0002d01060004000000f0010a000e00000024030500bc037703bc03c000d203af00d2036
703bc037703040000002d0109000e000000240305007b0377037b03c000bc03c000bc037
7037b037703040000002d01050007000000fc02000000bfbf008020040000002d010a000
e00000024030500bc03c000d203af009103af007b03c000bc03c00009000000fa0200000
2000000000000002200040000002d010c0004000000f0010b0007000000fc02000080806
6008020040000002d010b0004000000f0010a000e000000240305005f0477035f04ab027
5049a02750467035f04770307000000fc020000ffffcc008020040000002d010a0004000
000f0010b000e000000240305001e0477031e04ab025f04ab025f0477031e04770304000
0002d01050004000000f0010a0007000000fc020000bfbf99008020040000002d010a000
e000000240305005f04ab0275049a0234049a021e04ab025f04ab0209000000fa0200000
2000000000000002200040000002d010b0004000000f0010c00040000002d01060004000
000f0010a000e000000240305009f0477039f046401b6045301b60467039f04770304000
0002d0109000e000000240305005f0477035f0464019f0464019f0477035f04770304000
0002d01050007000000fc02000000bfbf008020040000002d010a000e000000240305009
f046401b6045301750453015f0464019f04640109000000fa02000002000000000000002
200040000002d010c0004000000f0010b0007000000fc020000808066008020040000002
d010b0004000000f0010a000e00000024030500420577034205ab0258059a02580567034
205770307000000fc020000ffffcc008020040000002d010a0004000000f0010b000e000
00024030500010577030105ab024205ab024205770301057703040000002d01050004000
000f0010a0007000000fc020000bfbf99008020040000002d010a000e000000240305004
205ab0258059a0217059a020105ab024205ab0209000000fa02000002000000000000002
200040000002d010b0004000000f0010c00040000002d01060004000000f0010a000e000
000240305008305770383057402990563029905670383057703040000002d0109000e000
000240305004205770342057402830574028305770342057703040000002d01050007000
000fc02000000bfbf008020040000002d010a000e0000002403050083057402990563025
8056302420574028305740209000000fa02000002000000000000002200040000002d010
c0004000000f0010b0007000000fc020000808066008020040000002d010b0004000000f
0010a000e00000024030500250677032506ab023b069a023b0667032506770307000000f
c020000ffffcc008020040000002d010a0004000000f0010b000e00000024030500e4057
703e405ab022506ab0225067703e4057703040000002d01050004000000f0010a0007000
000fc020000bfbf99008020040000002d010a000e000000240305002506ab023b069a02f
a059a02e405ab022506ab0209000000fa02000002000000000000002200040000002d010
b0004000000f0010c00040000002d01060004000000f0010a000e0000002403050066067
703660690027c0680027c06670366067703040000002d0109000e0000002403050025067
70325069002660690026606770325067703040000002d01050007000000fc02000000bfb
f008020040000002d010a000e00000024030500660690027c0680023b068002250690026
6069002040000002d01050004000000f0010a00040000002d01070004000000f0010b000
9000000fa02000000000000000000002200040000002d010a00050000001402840333010
400000002010100050000001302520033010500000014028403330105000000130284032
5010500000014021e0333010500000013021e032501050000001402b8023301050000001
302b80225010500000014025102330105000000130251022501050000001402eb0133010
50000001302eb01250105000000140285013301050000001302850125010500000014021
f0133010500000013021f012501050000001402b8003301050000001302b800250105000
0001402520033010500000013025200250109000000320a6303fb0001000000300021000
a000000320afd02da00020000003530210021000c000000320a9702b9000300000031303
0002100210021000c000000320a3002b90003000000313530002100210021000c0000003
20aca01b90003000000323030002100210021000c000000320a6401b9000300000032353
0002100210021000c000000320afe00b90003000000333030002100210021000c0000003
20a9700b90003000000333530002100210021000c000000320a3100b9000300000034303
000210021002100040000002d010000040000002d010000040000002d010700040000002
d010300040000002701ffff030000001e00040000002d010000040000002d010a0005000
0000102ffffff0005000000090200000000070000001604cc02ab000a014a00040000000
4010d00040000000201010012000000fb02c4ff0000840384039001000000cc001000204
17269616c2043797200cc00040000002d010b000e000000320ac30257000e0000002520e
2b3e420eaeeedf2f0eeebfe040000002d01000004000000f0010b00040000002d0100000
40000002d010700040000002d010300040000002701ffff030000001e00040000002d010
000040000002d010a00050000000102ffffff00050000000902000000000700000016046
20412070a000a00050000001402840333010400000004010d00040000000201010005000
000130284038606050000001402840333010500000013029203330105000000140284031
602050000001302920316020500000014028403f9020500000013029203f902050000001
4028403dc030500000013029203dc030500000014028403bf040500000013029203bf040
500000014028403a3050500000013029203a305050000001402840386060500000013029
2038606040000002d01010012000000320aa003400107000000c2e8f5b3e4ede00025001
f001c000c0020001e001f000f000000320ae303570105000000eff0eee1e0001e001f001
f0020001f000c000000320aa003710203000000493a49000f000f000f000c000000320aa
0034d0303000000493a33000f000f001f000c000000320aa003300403000000493a35000
f000f001f000c000000320aa003130503000000493a37000f000f001f000d000000320aa
003e60504000000493a31330f000f001f001f00040000002d010700040000002d0103000
40000002701ffff030000001e00040000002d010100040000002d010a00050000000102f
fffff00050000000902000000000700000016046c041c0700000000040000002d0107000
40000002d010300040000002701ffff030000001e00040000002d010100040000002d010
a00050000000102ffffff00050000000902000000000700000016046c041c07000000000
40000002d01020009000000fa02050000000000000000002200040000002d010b0004000
000f0010a00040000002d0108000400000004010d000400000002010200070000001b047
60173022c004601040000002d010700040000002d010800040000002d010300040000002
701ffff030000001e00040000002d010200040000002d010800040000002d010b0005000
0000102ffffff0005000000090200000000070000001604740172022c004601040000002
d010700040000002d010800040000002d010300040000002701ffff030000001e0004000
0002d010200040000002d010800040000002d010b00050000000102ffffff00050000000
90200000000070000001604740172022c00460109000000fa02000002000000000000002
200040000002d010a0004000000f0010b0007000000fc020000ffffcc008020040000002
d010b000400000004010d000400000002010200070000001b0498008d017000650104000
0000201010009000000320a5d009c010100000030002400040000002d010700040000002
d010800040000002d010300040000002701ffff030000001e00040000002d01020004000
0002d010b00040000002d010a00050000000102ffffff000500000009020000000007000
0001604740172022c004601040000002d010700040000002d010800040000002d0103000
40000002701ffff030000001e00040000002d010200040000002d010b00040000002d010
a00050000000102ffffff0005000000090200000000070000001604740172022c0046010
9000000fa02000002000000000000002200040000002d010c0004000000f0010a0004000
0002d01090004000000f0010b000400000004010d000400000002010200070000001b043
c018d0114016501040000000201010010000000320a01019c0106000000343820f7e0f12
40024001200220024002100040000002d010700040000002d010800040000002d0103000
40000002701ffff030000001e00040000002d010200040000002d010900040000002d010
c00050000000102ffffff0005000000090200000000070000001604740172022c0046010
40000002d010700040000002d010800040000002d010300040000002701ffff030000001
e00040000002d010200040000002d010900040000002d010c00050000000102ffffff000
50000000902000000000700000016046c041c0700000000040000002d010700040000002
d010800040000002d010300040000002701ffff030000001e00040000002d01020004000
0002d010900040000002d010c00050000000102ffffff000500000009020000000007000
00016046c041c0700000000040000002d010500040000002d01070004000000f0010c000
400000004010d002d000000f700000314000000000080000000008000008080000000008
0008000800000808000c0c0c000c0dcc000a6caf000fffbf000a0a0a40080808000ff000
00000ff0000ffff00000000ff00ff00ff0000ffff00ffffff000400000034020a0007000
000fc020000ffffcc000000040000002d010b0009000000fa02000002000000000000022
200040000002d010c00070000001b04ca00990160005c01040000002d01050004000000f
0010b00040000002d01070004000000f0010c00040000002d01090009000000fa0200000
2000000000000022200040000002d010b00070000001b0470019901fa005c01040000002
d010500040000002d01070004000000f0010b0005000000090200000000050000000102f
fffff00040000002d010500040000002701ffff050000000c026c041c07030000001e000
50000000102ffffff0005000000090200000000040000002701ffff050000000b0200000
000030000001e00050000000102ffffff000500000009020000000004000000f00101000
4000000f0010000040000002701ffff0300000000000000160456025a033f00700104000
0002d0107000400000000000000?en.2. Aeeiai?ea

c?inoaiiy /enaeueiino?

E. patella a caeaaeiino? a?ae e?aoiino? ?icaaaeaiiy no?/ieo aiae NIAA,
ye? i?noyoue iaooo.

Nia?oi?noue iaoie?on?a A. salina ia ia?aaeuoaaea eiio?ieueieo cia/aiue
i?e ?icaaaeaii? i?iae c ?aca?aoa?a-iaeiie/oaa/a o o?noue ?ac?a.

A?ioanooaaiiy ii?nueei? aiaee a ?aeii? neeaeo no?/ieo aiae c NIAA
iieacaei, ui a?aeoeeaiiy a?ae eiio?ieth (o a?e ca?eueoaiiy iieaciee?a
?inoo) niinoa??aaee o?eueee aeey eaai?aoi?ieo eoeueoo? ieaieoiiieo
aiaei?inoae. Aeanoeaino? aiaee, ye? noeioethaaee iiae?ee ee?oei, cieeaee
i?e ?? ?icaaaeaii? o 4 ?ace. Aoaeo noeioeyoe?? iieaciee?a ?inoo o
oano-ia`?eo?a a i?iaao c ?aca?aoa?o-iaeiie/oaa/a, a oaeiae a?aenooi?noue
aino?i? oiene/iino? a ?aeii? ii?nueeiai neeaeo no?/ieo aiae NIAA ia
iiaea neoaeeoe e?eoa???i iaaaciaee iaooiaiai caa?oaeiaiiy a oe?eiio aeey
caieiaieo aeaaoi??e, aei yeeo iino?eii neeaeathoueny no?/i? aiaee, ye?
i?noyoue iaooo.

A?ioanoOAAIIss i??niiaiaeieo

no?*IEO AIAe, ssE? nEEAeATHOUeNss a ii?a

Iniiaiee iio?e caa?oaeithth/eo ?a/iaei aio?iiiaaiiiai iioiaeaeaiiy
iaaeoiaeeoue o ii?a ?c i??niiaiaeiei noieii. Ia aeaeue, aein? ia
ciaeaeaiee oa e iaa?yae /e ?aaeueii ?nio? o i?e?iae? oi?aa?naeueiee
oano-ia`?eo /e oano-ooieoe?y, ye? iaeii/anii iiaeia aeei?enoiaoaaoe ye
aeey i??nieo, oae ? aeey ii?nueeeo aiae.

A ca`yceo c oeei, ia iaoo aeoieo, oi/eia? i??niiaiaei? aa?aaia?
aeaea?aea ii?nueeiai caa?oaeiaiiy iaiao?aeii eiio?iethaaoe ca iieacieeii
“oiene/i?noue” c aeei?enoaiiyi i??niiaiaeieo noaiaea?oieo oano-ia`?eo?a
(Daphnia magna Straus, Ceriodaphnia affinis Lilljeborg) ua aei ?o
iaaeoiaeaeaiiy aei ii?nueeiai na?aaeiaeua. ? o?eueee i?ney
iaaeoiaeaeaiiy no?/ieo aiae aei ii?y, oiaoi i?ney ?o ?icaaaeaiiy,
aeioe?eueii i?iaiaeeoe ioe?ieo oiene/iino? aiaee c aeei?enoaiiyi
ii?nueeeo oano-ia`?eo?a (Ph. tricornutum, A. salina).

Oeae i?aeo?ae aoa caiii?iiiaaiee aaoi?ii i?e i?iaaaeaii? eiiieaenii?
aeieia?/ii? ceiiee o Iaeanuee?e caoioe? ia iie?aii? “Aaeeeee Oiioai” o
aa?ani? 1996 ?. (Aeaenaiae?ia e ae?., 1998). Oiiiaa ceiiea a ii??
aeeiioaaeany iaeii/anii c a?aeai?ii i?ia no?/ieo aiae c? neeaeiiai
eieaeoi?a NAI “I?aaeaiia”. A?ioanooaaiiy neeaeieo aiae NAI “I?aaeaiia”
aoei aeeiiaii o a?aeiia?aeiino? aei EIAe 211.1.4.055-97. No?/ia
aiaea i?ae /an neeaeo o ii?a aeyaeeany aaniethoii eaoaeueiith aeey C.
affinis. O?eueee o?ee?aoia ?icaaaeaiiy no?/ii? aiaee aeyaeeiny
aeinoaoi?i aeey oiai, uia i?iaa, yea oanooaaeany, iicaaaeeany a
oiene/ieo aeanoeainoae. Iaiao?aeii iii?oeoe, ui NAI «I?aaeaiia» iiaeia
a?aeianoe aei e?oiieo aeaea?ae aa?aaiaiai caa?oaeiaiiy Iaeanueei?
caoiee.

Ca niinoa?aaeaiiyie inoaii?o i`yoe ?ie?a, oiene/i?noue aiae, ye?
neeaeathoueny c oeueiai ia`?eoo, ?a?no?o?oueny iino?eii i?e i?i?iaeuei?e
e?aoiino? ?icaaaeaiiy o 2-8 ?ac?a.

Ioe?iea oiene/iino? no?/ieo aiae NAI “I?aaeaiia” a cii? ?icn?thaaiiy
a ii?? aoea aeeiiaia o a?aeiia?aeiino? aei EIAe 211.1.4.047-95 c
aeei?enoaiiyi eaai?aoi?ii? eoeueoo?e ii?nueeeo ieaieoiiieo aiaei?inoae
Ph. tricornutum. ?acoeueoaoe a?ioanooaaiiy iieacaee, ui a iiaa?oiaaiio
ai?eciio? aaciina?aaeiuei a oi/oe? neeaeaiiy ? ia i?eeaaeeo noaioe?yo
noi?ioaaeany ieyia, yea oa?aeoa?ecoaaeany i?aeaeuaiieie iieacieeaie
?inoo eoeueoo? aiaei?inoae. I?eeiaiaeia i?eaa?aaeia ciia, iaaiaee,
yaeyea niaith iaoe?io ae?eyieo, yea oa?aeoa?ecoaaeany i?eai?/aiiei
noaiii oano-eoeueoo? aiaei?inoae, ui aeicaieeei a?aeianoe i?iae aiaee c
oaeeie oa?aeoa?enoeeaie ca oeaeith oiene/iino? aei eean?a
neaaioiene/iiai oa oiene/iiai a?aeiia?aeii.

Aenia?eiaioaeueia ioe?iea yeino? aeiiieo a?aeeeaae?a i?ai?/ii-cao?aeii?
/anoeie *i?iiai ii?y

Aei ?aeii?a, ye? aeo?eiothoue cia/i? aio?iiiaaii? iaaaioaaeaiiy, o ia?oo
/a?ao, iiaeia a?aeianoe aeaaoi??? ii?o?a, i?noey i?iaaaeaiiy
aeiiiiaeeaethaaeueieo ? ao?iaeo ?ia?o, ii?nuee? caaeeua a?oio?a. ?ieue
aeiiieo a?aeeeaae?a a i?ioeanao, ye? iia`ycai? c oi?ioaaiiyi
oieneeieia?/ii? neooaoe?? a oeeo ?aeiiao aei oaia??oiueiai /ano aea/aia
ua iaaeinoaoiuei. Iaea?eueoee aieea ia a?ae?ia?iio?a iiaeooue ni?e/eiyoe
aiaei?ic/eii? niieoee aeiiieo a?aeeeaae?a.

Iaeanueeee ii?o. Aeine?aeaeaiiy oiene/iino? aiaeieo aeno?aeo?a aeiiieo
a?aeeeaae?a Iaeanueeiai ii?oo ca iaoa??aeaie aaniyii? ceiiee 1993 ?.
aoee i?iaaaeai? ia eaai?aoi?ieo eoeueoo?ao ii?nueeeo ieaieoiiieo
aiaei?inoae Ph. tricornutum, iaoie?onao A. salina, ?iooci??yo E. patella
lemani.

Iieacaii, ui aiaei? aeno?aeoe aeiiieo a?aeeeaae?a Iaeanueeiai ii?oo iaee
aenieo oiene/i?noue aeey on?o aeei?enoaieo oano-ia`?eo?a. O aeayeeo
aeiaaeeao (Noa?ee oaee?eii, Iaooiaaaaiue) oiene/i?noue aiaeieo
aeno?aeo?a aeiiieo a?aeeeaae?a c?inoaea c aeeaeiith caeyaaiiy i?iae.
Oiene/ia ae?y i?iyaeyeany o i?eai?/aii? ?inoo, aeieoaaii? iaoai?ci?a
ae?eaiiy oa caaeaae? ee?oei a eoeueoo?ao aiaei?inoae oa ?iooci??e,
nia?oiino? iaoie?on?a a?oai?e i?e ia?aoiae? c aoaio II i?oiiaoie?ona
ia aoai I iaoaiaoie?ona. A ie?aieo aeiaaeeao a?aei?/aia noeioeyoe?y
ee?oeiieo iiae?e?a a eoeueoo?ao ?iooci??e (aeiii? a?aeeeaaee Iaeanueeiai
noaeii?aiiioiiai caaiaeo).

Aiaei? aeno?aeoe aeiiieo a?aeeeaae?a ia an?o noaioe?yo a?aeai?o i?ia a
Iaeanueeiio ii?oo aeeeeeaee a?aeoeeaiiy a?ae eiio?ieth o oano-ia`?eo?a.
Oeae oaeo i?aeoaa?aeaeo? oaeiae e aaioinia ceiiea, yea aoea i?iaaaeaia
iaeii/anii c iaoeie aeine?aeaeaiiyie ? eiinoaooaaea i?eai?/aiiee noai
aeiiieo oa?oiiaaiue, a o aeayeeo aeiaaeeao e iiaio a?aenooi?noue
iae?iciiaaioino.

Iiaa Iaeanueea caaeeua a?oio?a. Aeaii?ia a?oio?a, ui aoee caeiaoo? i?e
aeiiiiaeeaethaaeueieo ?iaioao a ii?nueeeo ii?oao, i?ecaiaeeoue aei
eieaeueieo ci?i oiia ?nioaaiiy aeiiieo oa?oiiaaiue. A?aeaoaa?oueny
caneiaiiy i?aai?ci?a ?a/iaeiaie, ye? neeaeathoueny, ciaioo?oueny
/enaeuei?noue ? a?iiana aeae?a, ye? ? /ooeeaeie aei caneiaiiy a?oio?a,
ca?eueoo?oueny ai?no caaeneeo ?a/iaei o i?eaeiiieo oa?ao aiaee,
ci?ith?oueny oa?aeoa? noano?aoo. O aeiiieo a?aeeeaae?a c`yaey?oueny
iiaa aeanoea?noue — oiene/i?noue, yea aeeeeeaia iaaeoiaeaeaiiyi ?
iino?eiith i?enooi?noth a aeiiieo a?aeeeaaeao caa?oaeithth/eo ?a/iaei
??ciiiai?oii? o?i?/ii? i?e?iaee oa iioiaeaeaiiy. C iaoith ioe?iee
oiene/iino? aeiiieo a?aeeeaae?a a ?aeii? Iiaiai Iaeanueeiai caaeeua
a?oio?a (IICA) aoea cae?eniaia ceiiea o o?aai? 1993 ?.

A?ioanooaaiiy aeiiieo a?aeeeaae?a IICA, yea aoei i?iaaaeaii c
aeei?enoaiiyi eaai?aoi?ii? eoeueoo?e Ph. tricornutum, iieacaei, ui
aaciina?aaeiuei ia caaeeu? aeiii? a?aeeeaaee aoee a?aeianai? aei eeano
iaoiene/ieo: ?o aiaei? aeno?aeoe ia aeeeeeaee a?aeoeeaiue o ?icaeoeo
eoeueoo? ieaieoiiieo aiaei?inoae. Iaeneiaeueiee i?e??no ?ioaa?aeueii?
a?iiane aiaei?inoae (220,0 e 171,7% ii??aiyii c eiio?ieai) aoa
a?aei?/aiee ia aeaio noaioe?yo, ye? ?icoaoiaai? ia ?ic??c? o iai?yieo
aei ?ee?/?anueeiai ii?oo, aea o aeiiieo a?aeeeaaeao ciaeaeai? ne?aee
a?aeneiee iaoa??ae?a aeaii?iao.

Aenia?eiaio c ioe?iee oiene/iino? aiaeieo aeno?aeo?a aeiiieo a?aeeeaae?a
IICA aeey eoeueoo? ii?nueeeo ?iooci??e aeyaea oe?eo ieceo caaaeueieo
?en o ?aaeoe?? oeueiai oano-ia`?eoa ? eoeueoo? ii?nueeeo aiaei?inoae,
oiaoi o noeioeyoe?? iiae?eo ?oi?o ee?oei.

Ia i?aenoaa? i?iaaaeaiieo aeine?aeaeaiue aoa c?iaeaiee aeniiaie, ui
neeaeaiiy caa?oaeiaieo a?oio?a ia ni?aiaaea? c iaaeith a?aeaaaeaiii?
aeey oe??? iaoe ae?eyiee.

Aenia?oia ioe?iea oiene/iino? caA?OAeITHTH*EO

?A*IAEI iaoiaeaie a?ioanooaaiiy

Aieneoe ? aeeiicai. Aeine?aeaeaiiy oiene/iino? aieneo?a oa aeeiicaia,
ye? iiaeooue iio?aieyoe aei ii?y i?ae /an o?ainii?ooaaiiy oa
ia?aaaioaaeaiiy a /i?iiii?nueeeo ii?oao ia? cia/iee ?ioa?an. Aeia?a
a?aeiii, ui niieoee aethi?i?th, ye? aoiaeyoue aei neeaaeo aeeiicaia
iathoue iaaaeeeo oiene/i?noue, ui e iaoiiaeth? ?o oe?iea aeei?enoaiiy
a i?ioeanao i/euaiiy ieoii? aiaee, a oa?/ia?e oa iaaee/i?e
i?iieneiaino?. I?e oeueiio aenia?eiaioaeueii aeiaaaeaii, ui aoaeoe
iaaaoeaiiai aieeao caaeneeo /anoie aieneo?a oa aeeiicaio aeey
a?ae?ia?iio?a aeyaeythoueny ia ??ai? oaeeo eiioeaio?aoe?e, ye? ?aaeueii
iiaeooue aeieeaoe o ii?? a ia??iae ia?aaaioaaeaiiy ia eieaeueieo
ae?eyieao aeaaoi??? ? ia iao?eaaeee /an.

Ae?y caaeneeo /anoie oeeo niieoe ia a?ae?ia?iio?a ia? iaoai?/iee,
iioeiaeaeoth/ee oa?aeoa?. Aeey aiaei?inoae i?e eiioaeo? c caaeneeie
/anoeaie aieneo?a ? aeeiicaio aoei oa?aeoa?iei ceeiaiiy ee?oei ?
ooai?aiiy ee?oeiieo eiiaeiia?ao?a, ao?aoa ieie ieaao/ino? (?en. 3).

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00000000000000000000000000000000000000000000000000000000000?en. 3. Aieea
aeeiicaio ? aieneo?a ia /enaeueiino? ee?oei a eoeueoo?ao Ph.tricornutum
(aeniiceoe?y

24 aiaeeie)Aieneoe ? aeeiicai, ye? ciaoiaeyoueny o caaeneiio noai?,
aeoeaii iiaeeiaeeny ?aeiiiae?aieie. A aenia?eiaio? niinoa??aaee
cieaeaiiy ieaao/ino? oano-ia`?eo?a, ci?io oa?aeoa?a ?oo?a aiane?aeie
iae?iaiiy caaaeaieo /anoie ia aioaie ? iaiaeeaoee.

Iaea?eueo /ooeeaeie aei i?enooi?no? o aiae? caaaeaieo /anoie aeyaeeeny
ieaieoiii? aiaei?ino?: LC5048/an aeeiicaio ? aieneo?a neeaaeaee 3,0 ?
1,5 a/e a?aeiia?aeii. Aeey ieaieoiiieo ?aeiiiae?aieo LC5048/an aeeiicaio
? aieneo?a aei??aithaaee 8,0 e 6,5 a/e a?aeiia?aeii.

Aeaaia?. Iiaee i?aia?ao aeaaia?, aeoeaiei eiiiiiaioii yeiai ?
iiaa?oiaai-aeoeaia ?a/iaeia (IA?), i?ecia/aiee aeey onoiaiiy iaooe c?
noi/ieo aiae, i/euaiiy iiaa?oiae oa a?oio?a, ye? caa?oaeiai?
iaooii?iaeoeoaie ? aee?aie. I?aia?ao aeicaieaii neeaeaoe ia noaioe?th
a?ieia?/ii? i/enoee o eiioeaio?aoe?? 50 ia/e (a ia?a?aooieo ia
eiioeaio?aoe?th IA? — 5 ia/e), aea a?i i?aeaea?oueny aeano?oeoe?? aei
iaoiene/ieo eiiiiiaio?a.

A oiae? aecia/aiiy oiene/iino? i?aia?aoo ioe?ithaaee ne?aeoth/?
ia?aiao?e: aino?o oiene/i?noue aeaaia?o aeey eaai?aoi?ieo oano-ia`?eoia,
caeaaei?noue oiene/iino? a?ae oa?i?io caa??aaiiy ?ic/ei?a,
oiene/i?noue i?e noi?ni?e ae?? aeaaia?o ? aeecaeueiiai iaeeaa.

Na?aeii?eaioiaaiee ?ic/ei aeaaia?o a eiioeaio?aoe??, yea aei??aith? AAeE
? yeo aeicaieaii neeaeaoe a eaiaeecaoe?th ? aino?ioiene/iith
niieoeith, yea aeeeeea? caaeaaeue iaoie?on?a a?oai?? ? cieaeo? ?inoia?
iieacieee a eoeueoo?ao ii?nueeeo ieaieoiiieo aiaei?inoae (?en. 4).

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50000000102ffffff000500000009020000000004000000f0010100040000002701ffff0
3000000000000050500000014020e0238050400000000000000?en.4. Aieea
na?aeii?eaioiaaieo ?ic/ei?a aeaaia?o ia iaoie?on?a A.salina (/an
100%-ai? caaeaae? oano-ia`?eo?a) ?

?h.tricornutum (?ioaa?aeueia a?iiana ee?oei i?ney

48-e aiaeeiii? aeniiceoe??).Aenia?eiaioaeueii aeiaaaeaia iayai?noue
neia?a?/iiai aoaeoo ?ic/ei?a aeaaia?o ? iiaooiaeo no?/ieo aiae NAI
“I?aaeaiia”, i?e yeiio noia?iee aoaeo oiene/iino? aeey a?ae?ia?iio?a
aeua, i?ae ae?y eiaeiiai ie?aiiai eiiiiiaioa. I?e naiee?aoiiio
?icaaaeaii? no?/ieo aiae oeae aoaeo cieeaa.

Iieacaii cieaeaiiy oiene/iino? aeaaia?o a caeaaeiino? a?ae oa?i?io
caa??aaiiy ?ic/ei?a (?en. 5).

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50000000102ffffff00050000000902000000000700000016040f03f7063e00100104000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?en.5. ?ioaa?aeueia a?iiana ee?oei a
eoeueoo?ao Ph.tricornutum i?ney 48-e aiaeeiii? aeniiceoe?? a ?ic/eiao
aeaaia?o c ??cieie oa?i?iaie caa??aaiiy.Ieaieoiii? aiaei?ino? o
na?aeii?eaioiaaieo ?ic/eiao aeaaia?o (400 ia/e) aie?aee, aea i?ney
aeo?eiaiiy ?ic/eio i?ioyaii 6-e ae?a a oano-eoeueoo?ao a?aei?/aii
iiyeaiiy ee?oei, yee ae?eeeeny. Eiioeaio?aoe?? 5-100 ia/e aieeaaee ia
eoeueoo?e oaeei naiei /eiii: caa??aaiiy ?ic/ei?a i?ecaiaeeei aei
cieaeaiiy ?o oiene/iino?, aea a aeiaeiiio c aa??aio?a (ca aeeeth/aiiyi
5 ia/e) o aeo?eiaieo ?ic/eiao a?iiana ee?oei ia aeinyaaea eiio?ieueieo
cia/aiue. Aino?a oiene/i?noue ?ic/eio aeaaia?o 400 ia/e aeey
?aeiiiae?aieo caa??aaeany i?ioyaii 9-e ae?a. Na?aeii?eaioiaaiee ?ic/ei
aeaaia?o 50 ia/e aeeeeeaa 100 %-ao caaeaaeue iaoie?onia a?oai?? /a?ac
1,5 aiaeeie, i?ney 6-e ae?a caa??aaiiy ?ic/eio caaeaaeue ?aeiiiae?aieo o
iueiio neeaaeaea 50%. I?e aea/aii? noi?nii? ae?? aeaaia?o (25 ia/e) ?
aeecaeueiiai iaeeaa (25 ia/e) aoei anoaiiaeaii, ui noi?o, yea aoea
aeo?eiaia i?ioyaii 1,5 aiaeei caa??aaea oiene/i? aeanoeaino?, a i?ney
24-o aiaeeiiiai a?aenoithaaiiy oiene/i?noue noi?o? aeey ?aeiiiae?aieo ?
aiaei?inoae cia/ii ciaiooaaeany.

AENIIAEE

1. Aeey ioe?iee yeino? i?e?iaeieo ii?nueeeo aiae, no?/ieo aiae
ii?nueeiai iioiaeaeaiiy, ie?aieo ?ic/eiieo o?i?/ieo niieoe, aeiiieo
a?aeeeaae?a oa caaeneeo ?a/iaei cai?iiiiiaai? ye iaia`yceia?
oano-ia`?eoe eoeueoo?e ii?nueeeo ieaieoiiieo aiaei?inoae Ph.
tricornutum oa ?aii? iaoie?one A. salina, a ye aeiaeaoeia? — eoeueoo?e
ii?neeo ?iooci??e E. patella lemani.

2. Cai?iiiiiaaia oeaea aecia/aiiy oiene/iino? ii?nueeeo aiae.
Aaaaoi??/i? niinoa?aaeaiiy ca ye?noth aiae Iaeanueei? caoiee oa
i?eeaaeeo aeaaoi??e aeiaaee, ui ii?nueea aiaea oa?aeoa?ecoaaeany ??ciei
nooiaiai oiene/iino? (a?ae -2 aei +3 aae?a ca oeaeth oiene/iino?).

3. Cai?iiiiiaaiee iiaee i?aeo?ae aei ioe?iee oiene/iino? i??niiaiaeieo
noie?a, ye? neeaeathoueny a ii?a. Ia ia?oiio aoai? i??niiaiaeiee no?e
i?aeeyaa? a?ioanooaaiith c aeei?enoaiiyi noaiaea?oieo i??niiaiaeieo
oano-ia`?eo?a C. affinis /e D. magna. Ia ae?oaiio aoai?, i?ney
?icaaaeaiiy noiea a ii?nueeiio noai??, a?ioanooaaiiy aeeiiothoue ia
ii?nueeeo oano-ia`?eoao — eaai?aoi?ieo eoeueoo?ao aiaei?inoae
Ph.tricornutum oa ?aii?o iaoie?aeueieo noaae?yo A. salina.

4. O i?ai?/ii-cao?aei?e /anoei? *i?iiai ii?y aei “aa?y/eo ieyi”
a?aeianai? aaaai? Iaeanueeiai ii?oo, ?aeiie neeaeaiiy no?/ieo aiae c
NAI “I?ai?/ia” ? “I?aaeaiia”. Ia??iaee/ii oiene/i? aeanoeaino? ii?nueei?
aiaee a?aei?/aeeny a ?aeii? aeioneo A?eaae?enueeiai ceeaiaiai eieaeoi?a.
Ia iie?aii? “Iaeanueea caoiea” (a?inoaioe?y Iaeanueeiai
aea?aeoi?aa?neoaoo) aiaea, ca aeaieie aaaaoi??/ieo niinoa?aaeaiue,
oa?aeoa?ecoaaeany ye iaoiene/ia ca oeaeith oiene/iino?.

5. Ioe?iea oiene/iino? no?/ieo aiae NIAA (ca noaiii ia ea?oaiue 1993
?.) iieacaea, ui no?/i? aiaee, ye? i?noyoue iaooo (0,19-0,1 ia/e),
aeeeeeaee a?aeoeeaiiy a?ae eiio?ieth a iieacieeao ?inoo eoeueoo?
aiaei?inoae ? ?iooci??e, caaeaae? iaoie?on?a a?oai??. Iaea?eueo
/ooeeaith aei iaooiaiai caa?oaeiaiiy aoea aiaei?inoue Ph. tricornutum.
Anoaiiaeai? cia/aiiy i?i?iaeueii? e?aoiino? ?icaaaeaiiy aiae, o yeeo
i?noeoueny iaooa, aeey onoiaiiy oiene/ieo aoaeo?a aeey oano-ia`?eoia.

6. Ca aeaieie aa?aniaai? 1996 ?. ceiiee, i??niiaiaeiee no?e c aa?aaiaiai
aeaea?aea caa?oaeiaiiy NAI “I?aaeaiia” aoa aino?ioiene/iei aeey
oano-ia`?eoo C. affinis. Oiene/i? aeanoeaino? no?/ii? aiaee iiaeia
aoei onoiooe i?e o?ee?aoiiio ?? ?icaaaeaii?. A oi/oe? ii?nueeiai neeaeo
oa e cii? ?icn?thaaiiy no?/ieo aiae NAI «I?aaeaiia» a?aei?/aia
noeioeyoe?y ee?oeiieo iiae?e?a a oano-eoeueoo?ao ii?nueeeo aiaei?inoae
Ph. tricornutum, ui aeicaieeei ieiioo?eoe ?aeii ?iciianthaeaeaiiy
no?/ieo aiae NAI “I?aaeaiia”.

7. Aiaei? aeno?aeoe aeiiieo a?aeeeaae?a aaaaiae Iaeanueeiai ii?oo
oa?aeoa?ecoaaeeny aino?ith oiene/i?noth aeey oano-ia`?eo?a. A ?yae?
aeiaaee?a oiene/i?noue aiaeieo aeno?aeo?a aeiiieo a?aeeeaae?a a aaaaiyo
Iaeanueeiai ii?oo ci?ithaaeanue a caeaaeiino? a?ae aeeaeie ?o caeyaaiiy.
A Iaooiaaaai? iaea?eueo oiene/ieie aoee aiaei? aeno?aeoe aeiiieo
a?aeeeaae?a c ai?eciioo 10-20 ni, o Noa?iai oaee?eiia — c ai?eciioo
30-50 ni, a a aeaaoi??? N?C — c iiaa?oiaaiai ai?eciioo (0-10 ni).

8. Aiaei? aeno?aeoe aeiiieo a?aeeeaae?a Iiaiai Iaeanneiai caaeeua
a?oio?a aoee iaoiene/ieie aeey eaai?aoi?ieo eoeueoo? Ph. tricornutum. Ca
iieacieeaie ?inoo eoeueoo? aiaei?inoae ? ?iooci??e, ye? cia/ii
ia?aaeuoaaee eiio?ieuei? cia/aiiy, aoee anoaiiaeai? i?noey aeaii?iao
caa?oaeiaieo a?oio?a ca iaaeaie IICA.

9. Aenia?oia ioe?iea oiene/iino? neio/eo aaioaae?a — aieneo?a ?
aeeiicaio iieacaea, ui aoaeoe iaaaoeaiiai aieeao caaeneeo /anoie oeeo
?a/iaei ia a?ae?ia?iio?a iineee oa?aeoa? iaoai?/iiai iioeiaeaeaiiy ?
aeyaeyeeny ia ??ai? eiioeaio?aoe?e, ye? ?aaeueii aeieeathoue o ii?? a
ia??iae ia?aaaioaaeaiiy (0,1-5,0 a/e). I?e eiioaeo? c caaeneeie /anoeaie
oeeo ?a/iaei aeey aiaei?inoae oa?aeoa?iith aoea ao?aoa ieaao/ino? ?
ooai?aiiy ee?oeiieo eiiaeiia?ao?a, aeey ?aeiiiae?aieo — aeoeaia
caeiaooaaiiy /anoie, ci?ia oa?aeoa?o ?ooo ? ao?aoa ieaao/ino?.

10. Iiaa?oiaai-aeoeaia ?a/iaeia aeaaia? a eiioeaio?aoe??, yea aei??aith?
AAeE (5 ia/e), iaea aino?ioiene/i? aeanoeaino? aeey a?ae?ia?iio?a. A
i?enooiino? no?/ieo aiae NAI “I?ai?/ia” oiene/ia ae?y aeaaia?o
iineethaaeany. I?e noi?ni?e ae?? aeaaia?o ? aeecaeueiiai iaeueiiai
na?aeii?eaioiaaia noi?o caa??aaea oiene/i? aeanoeaino?, i?e caa??aaii?
i?ioyaii 24 aiaeei ?? oiene/i?noue aeey aiaei?inoae ? ?aeiiiae?aieo
iii?oii cieaeoaaeany.

11. Iaoiaee a?ioanooaaiiy, ye? aoee cai?iiiiiaai? aeey ioe?iee
oiene/iino? ii?nueeiai na?aaeiaeua, aoee ai?iaiaai? i?e i?iaaaeaii?
iii?oi?eiao a i?ai?/ii-cao?aei?e /anoei? *i?iiai ii?y a ?aeiiao c?
ci?iiith nieii?noth, i?e aecia/aii? oiene/iino? no?/ieo aiae, aiaeieo
aeno?aeo?a aeiiieo a?aeeeaae?a, caaaeaieo /anoie, ie?aieo ?ic/eiieo o
aiae? o?i?/ieo niieoe. Iaoiaee iieacaee aenieo /ooeea?noue oano-ia`?eo?a
aei noaiaea?oiiai oieneeaioa a?o?iiaoo eae?y, a oaeiae aenieo
ca?aei?noue oa a?aeoai?thaai?noue ?acoeueoao?a i?e i?iaaaeaii?
i?aeeaai?aoi?ieo eaeea?iaie.

IOAE?EAOe?? II OAI? AeENA?OAOe??

1. Aeyoeia N.A., Iao?inyi A.A., Oiaeaeia E.A., Aeioeaiei O.A.,
Yeuenoa? A.I. Yenia?eiaioaeueiay ioeaiea ea/anoaa i?ea?aaeiuo aiae e
aeiiiuo ioeiaeaiee iaoiaeaie aeioanoe?iaaiey // Enneaaeiaaiea yeinenoaiu
*a?iiai ii?y: Na. iao/iuo o?oaeia Oe?IOeYI. — Aui. 1. — Iaeanna:
E?aiiieea?ao, 1994. — N. 141-148.

2. Iao?inyi A.A., Aeyoeia N.A., Aeioeaiei O.A., Oiaeaeia E.A.
Iaoiaee/aneea ?aeiiaiaeaoeee ii ii?neei aeieiae/aneei oanoai // ?aae.
Aeae?iaeie. ae. IAI Oe?aeiu. — Eeaa, 1995. — 56 n. Aeai. a AEIEOE
22.07.96, ? 2480-A96.

3. E?aeitheiaa A.I., Oeueyiiaa I.I., Onaiei I.A., Aeyoeia N.?., Iao?inyi
A.A. Aeioeaiei O.A, Oiaeaeia I.A., Oaae?iia E.A., Nieieiaa N.I.,
A?icaeia A.I., Eociaoeia I.I. Aiioanooaaiiy ii?nueei? aiaee oa
noi/ii?, yea aiaeaiaeeoueny a ii?a. Iaoiaeeea. EIAe 211.1.4.047-95. —
Aeaeaiiy io?oe?eia. — E., 1995. — 37 n.

4. Oiaeaeia E.A., Aeyoeia N.A., Iao?inyi A.A. Eniieueciaaiea ?aiieo
noaaeee yia?eiiaeueiiai ?acaeoey /a?iiii?neie ieaeee Mytilus
galloprovincialis Lam. aeey aeioanoe?iaaiey i?e?iaeiuo e noi/iuo aiae //
Aeae?iaeie. aeo?i. — 1996. — O. 32, ? 5. — N. 67-77.

5. Iao?inyi A.A., Aeyoeia N.A., Aeioeaiei O.A. Eioaa?aeueiay
ioeaiea ea/anoaa aiaeu e aeiiiuo inaaeeia i?ea?aaeiuo ?aeiiia ii?y //
Iao. iao/ii-i?aeo. eiio. “Yeieiae/aneea i?iaeaiu Iaeanneiai ?aaeiia
e eo ?aoaiea”. — Iaeanna, 1995. — N.131-135.

6. Aeyoeia N.A., Iao?inyi A.A. Nia?aiaiiua iaoiaeu aeioanoe?iaaiey
noi/iuo, i?e?iaeiuo e ieoueaauo aiae // Iao. iaaeaeoia?. iao/ii-i?aeo.
eiio. “Aiaea e caei?iauea-98” — Iaeanna: Ano?ii?eio, 1998. — N. 324-329.

7. Aeaenaiae?ia A.A., Ai?iaueaaa E.A., Aeyoeia N.A., Ianoa?iaa Ae.A.,
Iao?inyi A.A., Iieeuoe E.I., ?yneioeaaa I.E., Neiaaoa E.A., Oaieeineay
I.A., Oo?iaa I.I. naieoa?ii-yeieiae/aneea aniaeou oi?ie?iaaiey ea/anoaa
ii?neeo aiae a ?aeiia auionea eieeaeoi?a noaioeee aeieiae/aneie i/enoee
“thaeiay” // Iao. iaaeaeoia?. iao/ii-i?aeo. eiio. “Aiaea e
caei?iauea-98” — Iaeanna: Ano?ii?eio, 1998. — N. 263-269.

8. Iao?inyi A.A. Iiuo eniieueciaaiey iaeiieeaoi/iuo ieaieoiiiuo
aiaei?ineae aeey aeioanoe?iaaiey caa?yciaiiuo ii?neeo aeiiiuo inaaeeia
// O?. Anan. eiio. “Iaoiaeieiaey yeieiae/aneiai ii?ie?iaaiey”. — *.2. —
Oa?ueeia: A.e., 1990. — N. 141.

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Aeena?oaoe?y ia caeiaoooy a/aiiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
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Cai?iiiiiaai? iaoiaee ioe?iee oiene/iino? i?e?iaeii? ii?nueei? aiaee,
aeiiieo a?aeeeaae?a oa ie?aieo caa?oaeithth/eo ?a/iaei c aeei?enoaiiyi
oano-ia’?eo?a — eaai?aoi?ieo eoeueoo? ii?nueeeo ieaieoiiieo aiaei?inoae,
?iooci??e ? iaoie?aeueieo noaae?e ?aeiiiae?aieo. I?iaaaeaia aoanoaoe?y
iaoiae?a a?ioanooaaiiy a Iaeanuee?e caoioe? oa i?eeaaeeo aeaaoi??yo ii?y
c iaoith aeyaeaiiy iaea?eueo aaeeeeo aeaea?ae caa?oaeiaiiy.

Eeth/ia? neiaa: a?ioanooaaiiy, oiene/i?noue, oano-ia’?eo, ii?nueea
aiaea, aeiii? a?aeeeaaee.

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Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.17 — aeae?iaeieiaey. — Einoeooo aeieiaee
thaeiuo ii?ae ei. A.I.Eiaaeaaneiai IAI Oe?aeiu, Naaanoiiieue, 1998.

I?aaeeiaeaiu iaoiaeu ioeaiee oiene/iinoe i?e?iaeiie ii?neie aiaeu,
aeiiiuo ioeiaeaiee e ioaeaeueiuo caa?yciythueo aauanoa n eniieueciaaieai
oano-iauaeoia — eaai?aoi?iuo eoeueoo? ii?neeo ieaieoiiiuo aiaei?ineae,
eiooci?ee e iaoieeaeueiuo noaaeee ?aeiia?aciuo. I?iaaaeaia ai?iaaoeey
iaoiaeia aeioanoe?iaaiey a Iaeanneii caeeaa e i?eeaaathueo aeaaoi?eyo
ii?y n oeaeueth auyaeaiey iaeaieaa e?oiiuo enoi/ieeia caa?yciaiey.

Eeth/aaua neiaa: aeioanoe?iaaiea, oiene/iinoue, oano-iauaeo, ii?neay
aiaea, aeiiiua inaaeee.

Petrosyan A.G. Biotesting of marine water and bottom sediments of the
North-Western part of the Black Sea. — Manuscript.

The dissertation on competition of a scientific degree of the candidate
of biological sciences on a speciality 03.00.17 — hydrobiology. —
Institute of Biology of Southern Seas named by A.O.Kovalevskiy, National
Academy of Sciences of Ukraine, Sevastopol, 1998.

The new methods of toxicity assessment of water, bottom sediments and
individual substances using test-objects — laboratory cultures of the
marine plankton algae, infusoria and nauplii of the Crustaceans.
Approbation of these methods of biotesting in the Odessa Bay and
adjoined aquatories of the sea with the purpose to find the largest
sources of pollution.

Key words: biotesting, toxicity, test-object, marine water, bottom
sediments.

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