2

Aei?i?iiao?ianueeee aea?aeaaiee

oi?aa?neoao

AEoeia Na?a?e Iao?iae/

OAeE 574.3:581.524:581.55 (477.60)

Aio?iiiaaiia noeoean?y ?ineeiiino?

a?aeaae?a aoa?eueieo oaoo Aeiiaano

03.00.16 — aeieia?y

Aaoi?aoa?ao

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

Aei?i?iiao?ianuee — 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia o Aeiiaoeueeiio aioai?/iiio naaeo IAI Oe?a?ie

o a?oi? aeieia?? ?ineei a?aeae?eo i?iieneiai? aioai?ee

Iaoeiaee ea??aiee: aeieoi? a?ieia?/ieo iaoe, i?ioani?

Ao?aea ?a?na ?aai?aia, i?ia?aeiee iaoeiaee
ni?a?ia?oiee

?inoeoooo aa?iaeieia?? oa a?ioaoiieia?? OAAI (i. Ee?a)

Io?oe?ei? iiiiaioe:

aeieoi? a?ieia?/ieo iaoe, i?ioani? Aeieaiaa E?ae?y A?eai??aia, i?ioani?
eaoaae?e o?c?ieia?? ?ineei oa aeieia?? Aei?i?iiao?ianueeiai
aea?aeoi?aa?neoaoo (i. Aei?i?iiao?ianuee);

eaiaeeaeao a?ieia?/ieo iaoe, aeioeaio A?eiaa Iaoae?y Aiaoie??aia,
aeioeaio eaoaae?e oaoi?/ieo cania?a ieoiiai eiio?ieth Aeaaeai?? ieoii?
neoaeae Oe?a?ie (i. Aei?i?iiao?ianuee)

I?ia?aeia onoaiiaa: ?inoeooo aioai?ee ?i. I.A. Oieiaeiiai IAI

Oe?a?ie (i. Ee?a)

Caoeno a?aeaoaeaoueny “ 1 ” a?oaeiy 1999 ?. i _16__ aiaeei?

ia can?aeaii? niaoe?ae?ciaaii? a/aii? ?aaee Ae 08 051 04 ii
i?enoaeaeaiith iaoeiaiai nooiaiy aeieoi?a a?ieia?/ieo iaoe a
Aei?i?iiao?ianueeiio aea?aeaaiiio oi?aa?neoao? ca aae?anith: 320625,
INI, Aei?i?iiao?ianuee, i?iaoeie Iaoeiaee, 13, oi?aa?neoao,
a?ieiai-aeieia?/iee oaeoeueoao, ei?i. 17, aoae. 611.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? Aei?i?iiao?ianueeiai
aea?aeaaiiai oi?aa?neoaoo

Aaoi?aoa?ao ?ic?neaiee “ 29 ”__aeiaoiy_____1999 ?.

A/aiee nae?aoa? niaoe?ae?ciaaii? a/aii? ?aaee,

eaiaeeaeao a?ieia?/ieo iaoe, aeioeaio ________
A.I. Aeoa?ia1

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie

A?aeaaee aoa?eueieo oaoo neeaaeathoue cia/io /anoeio aio?iiiaaiieo oi?i
?aeue?oo Aeiiaano ? iii?oii caa?oaeiththoue iaaeieeoi? oa?eoi???. ?o
e?euee?noue aeinyaa? 1134-1270, a ciaoiaeaeaiiy a aonoiianaeaieo i?noeyo
a?aeiinii iineeth? aeieia?/i? iane?aeee ?o aieeao.

A inoaii?e /an, o cacyceo c ?ioaa?aoe??th Oe?a?ie aei na?oiai?
ni?eueiioe ca?eueoeeenue aeiiae aei noaio iaaeieeoiueiai na?aaeiaeua.
Aiane?aeie ?ano?oeoo?ecaoe?? aoa?eueii? i?iieneiaino? cia/ia e?euee?noue
a?aeaae?a aeaaaeaia c aenieoaoaoe?? ? i?aeeyaa? ?aeoeueoeaaoe??.

?aeoeueoeaaoe?y ?icaeyaea?oueny ye oaoiieia?y a?aeiiaeaiiy i?e?iaeieo
?ano?n?a, aiaeia eaiaeoaooieo noeoean?e (Caeoeaa, 1976). Caaaeueia
cae?naiiy oa?eeii?a ia a?aeiia?aea? ?o ciaoiaeaeaiith o noaiia?e cii?.
Iioeiaeueiei ia? aooe ii?aeiaiiy aea?aaii-/aaa?ieeiai? oa o?aacyienoi?
?ineeiiino? c oe?ieei aeei?enoaiiyi o?oiiae?i?aoeaiiai iioaioe?aeo
i?noeaai? oei?e (Oa?aa?ei, Oa?oioa, 1989). Ooai?aiiy iica?/ii?
no?oeoo?e, iiae?aii? aei oaei? i?e?iaeieo oa?oiiaaiue, aoaea
ia?aoeiaeaeaoe iioe?aiith ii?ooaiue, ine?eueee iayai?noue aioo??oi?o
iaae niia?eueith? ?o ?iciianthaeaeaiiy (Landscape heterogeneity…,
1987). Aeey oni?oiiai ae??oaiiy oe??? i?iaeaie iaiao?aeiith aeiiaith ?
aea/aiiy oa aeei?enoaiiy caeiiii??iinoae i?ioiaeaeaiiy aio?iiiaaiii?
noeoean?? ?ineeiiino? a oiiaao a?aeaae?a aoa?eueieo oaoo o noaiia?e
cii?.

Aiane?aeie cia/iiai ni?iuaiiy neeaaeo oa no?oeoo?e oa?oiiaaiue a?aeaae?a
aoa?eueieo oaoo, aiie ? c?o/iei iac?eoii aeey ae??oaiiy
caaaeueiioai?aoe/ieo ieoaiue o?oioeaiieia??, aei?iaoaaiiy oa
a?aei?aoethaaiiy iiaeo iaoiaee/ieo i?aeoiae?a aei aea/aiiy ?ineeiiino?.
Aei iaoeo aeine?aeaeaiue no/ani? iaoiaee o?oioeaiieia??, ye?
a?oioothoueny ia noaoenoeoe? oa nenoaiieia??, uiaei ?ineeiiino?
a?aeaae?a iaeaea ia canoiniaoaaeenue.Nii/aoeo noeoean?? ?ineeiiino? ?
aaeaoioiio a?aeaae?a aoa?eueieo oaoo aeine?aeaeoaaeenue c iaoith ?o
oaeaeeiai aea/aiiy oa aeei?enoaiiy aeyaeaieo caeiiii??iinoae i?e ?o
?aeoeueoeaaoe?? (Ieeueeia, 1978; I?iiuoeaiiay aioaieea, 1980; ?anoaiey e
i?iiuoeaiiay n?aaea, 1976-1989; O?aaeaaa, Aeueaeoeea e ae?., 1980).

Oi?ioaaiiy ?ineeiiiai iie?eao ia a?aeaaeao oaoo a?aeaoaa?oueny caiiai ia
noano?aoao, aea iaia? ?ineei, ia?aeaeueii c ooai?aiiyi aaeaoioiio. sseui
a i?e?iae? i?e?inoeoean?? ?ineeiiino? i?e eieaeueiiio ii?ooaii?
i?e?iaeieie /eiieeaie ? aoaiii ?? iinooiiaiai iiiaeaiiy ?
i?aeo?eiothoue a?ieia?/io ??ciiiai?oi?noue ca ?aooiie ni?a?nioaaiiy
aeae?a ??cieo noeoean?eieo noaae?e, oi a oaoiiaaiieo aeioiiao oaea
aiane?aeie ?o ?ici??o a?aeaoaa?oueny ye?niee ia?ao?ae, yeee ia?aaiaeeoue
oeae i?ioean ia aeuee aaiethoe?eiee ??aaiue. ?ai?oa a noeoean?ei?e
nenoai? ?ineeiiino? ?aa?iio aoee a?aenooi? oa?oiiaaiiy, i?enoiniaai? aei
aeieia?/ieo oiia a?aeaae?a. Oi?ioaaiiy iiaeo oeaiic?a, ui aaaea aei
aaiethoe?eieo ia?aoai?aiue, a?aeaoaa?oueny ia iaoeo i/ao ? aeinooiia
aeey aea/aiiy.

Iaoa e caaaeaiiy aeine?aeaeaiiy

Iaoith iaoeo aeine?aeaeaiue ? aecia/aiiy iine?aeiaiino? ci?i ?ineeiiino?
a?aeaae?a aoa?eueieo oaoo, iai?yio ?icaeoeo oa?oiiaaiue a?aeaae?a oa
coiiaethth/eo oe? i?ioeane i?e/ei.

A oiae? aeine?aeaeaiue ia?aaeaa/aeinue ae??oeoe ianooii? caaaeaiiy:

· aea/eoe ??ciiiai?oi?noue i?e?iaeii-ee?iaoe/ieo oiia ?aeiia
aeine?aeaeaiue ? ?ic?iaeoe eeaneo?eaoe?th aeioii?a a?aeaae?a;

· aeyaeoe iine?aeiai?noue i?ioiaeaeaiiy noaae?e noeoean?? ?ineeii?noth
a?aeaae?a;

· aecia/eoe oei?enoe/i? oa oeaiioe/i? oa?aeoa?enoeee noaae?e noeoean??;

· ii??aiyoe noeoean?ei? oa?oiiaaiiy a?aeaae?a c aiaeia?/ieie
o?oioeaiicaie a i?e?iaeieo i?noeaciaoiaeaeaiiyo ? oaeei /eiii aeyaeoe
iai?yi ?icaeoeo ?ineeiiino? a?aeaae?a;

· aecia/eoe noeoean?eiee noaoon aeae?a oei?e a?aeaae?a;

· ioe?ieoe ??ciiiai?oi?noue ?ineeiiino? a?aeaae?a c aeei?enoaiiyi
oei?enoe/ii? eeaneo?eaoe?? oa?oiiaaiue a?aeaae?a;

· aea/eoe i?aeaeaeiao ni?yaeai?noue ? aeyaeoe ni?yaeai? a?oie aeae?a;

· ?ic?iaeoe oai?aoe/i? iieiaeaiiy ? noai?eoe aaco aeaieo aeey
i?aeoe/iiai aeei?enoaiiy caeiiii??iinoae noeoean?? ?ineeiiino? a?aeaae?a
i?e ?aeoeueoeaaoe??.

Iaoeiaa iiaecia ?iaioe:

1) Aia?oa iiaoaeiaaia aii??eei-noaoenoe/ia iiaeaeue aio?iiiaaiii?
noeoean?? ?ineeiiino? a?aeaae?a aoa?eueieo oaoo – oeaiio?iiieeei
?ineeiiino? a?aeaae?a, c canoinoaaiiyi noaoenoe/iiai i?aeoiaeo aei
aea/aiiy oa c iiaeeo?eaoe??th aeei?enoaieo iaoiae?a ? aeei?enoaieo
?iaeaen?a (iiae?aiino?, o?oioeaiioe/ii? cia/ouino? oa ?i.) a?aeiia?aeii
aei oiia a?aeaae?a.

2) Aea/aii i?inoi?iao no?oeoo?o ia ??ai? ye ie?aieo aeae?a, oae ?
oa?oiiaaiue ? ?o ci?ie o oiae? noeoean??. Cai?iiiiiaaii iiaee can?a
aecia/aiiy aiiiaai?oaoa oa?oiiaaiue ? iiao noaio ci?ie ?iciiae?eo aeaeo
a caeaaeiino? a?ae noeoean?eiiai /ano ? a?aeiia?aeiino? oiia
i?noeaciaoiaeaeaiiy aeiiaai aeaeo.

3) Aia?oa ?ic?iaeaia iaoiaeeea e?euee?niiai aecia/aiiy a?aeiinii?
eiieo?aioicaeaoiino? aeae?a o aaaaoiaeaeiaeo o?oioeaiicao, yea
a?oioo?oueny ia aiae?c? i?aeaeaeiaeo ei?aeyoe?e oa ia?oe?aeueieo
ei?aeyoe?e.

4) Aecia/aiee noeoean?eiee noaoon aeae?a ? aia?oa cai?iiiiiaaia
iaoiaeeea o?oi?iaeeeaoe?? noeoean?eiiai noaio oa oaiaeaioe?e ?icaeoeo
oa?oiiaaiue ca noeoean?eiei noaoonii ?o aeae?a.

5)I?iaaaeaia oei?enoe/ia eeaneo?eaoe?y ?ineeiiino? a?aeaae?a, ? a ??
neeaae? aeyaeaii 7 anioe?aoe?e 4-o nithc?a, 3-o ii?yaee?a ? 3-o eean?a,
c ieo 2 anioe?aoe?? iienaii aia?oa

(Persicario — Obernetum behenii; Poo compressae — Hieracietum virosae).

I?aeoe/ia cia/aiey io?eiaieo ?acoeueoao?a

I?aeoe/iei cia/aiiyi ?acoeueoao?a aeine?aeaeaiiy ? ae??oaiiy ieoaiue, ui
noinothoueny ?aeoeueoeaaoe?? ii?iaeieo a?aeaae?a oaoo ? iie?ioaiiy
aeieia?/ieo ianoaaei o i?iieneiaiio ?aa?ii?, aea aeeaa a?ey 20%
ianaeaiiy Oe?a?ie. ?acoeueoaoe aeei?enoai? a oiae? aeeiiaiiy 2
athaeaeaoieo ? 4 ainiaeiaia??ieo oai, iiacycaieo c icaeaiaiiyi ii?iaeieo
a?aeaae?a oaoo. ?ic?iaeai? ia i?eeeaae? ?ineeiiino? a?aeaae?a iaoiaee
i?eaeaoi? aeey aea/aiiy ?ineeiiino? ?ioiai aaiaceno.

?iaioa aeeiioaaeanue a eaai?aoi??? aeieia?? a?aeae?eo i?iieneiai?
aioai?ee Aeiiaoeueeiai aioai?/iiai naaeo IAI Oe?a?ie a iaaeao oai
“Iiieoi?eia aio?iiiaaiiie noeoeannee ia ii?iaeiuo ioaaeao oaieueiuo oaoo
e ?ac?aaioea eiioeaioeee eo oeoiiioeiecaoeee” (? aea?ae?a?no?aoe??
0193U024297) ? “?ac?aaioea yeioeceieiae/aneie eiioeaioeee oi?ie?iaaiey
oeoioeaiicia ia ii?iaeiuo ioaaeao oaieueiuo oaoo Aeiiaanna” (?
aea?ae?a?no?aoe?? 0196U01215)

Aaoi?ii naiino?eii ?ic?iaeai? i?ia?aia ? iaoiaeeea aea/aiiy ?ineeiiino?
a?aeaae?a, i?iaaaeai? iieueia? ?iaioe, eaia?aeueia ia?iaea iaoa??ae?a ?
aiae?c aenia?eiaioaeueiiai iaoa??aeo, ocaaaeueiaia io?eiaia ?ioi?iaoe?y,
c?iaeai? oai?aoe/i? aeniiaee.

Ai?iaaoe?y ?iaioe

Iniiai? ?acoeueoaoe aeine?aeaeaiue aeiiia?aeaeenue ia i?aeia?iaeieo
eiioa?aioe?yo “I?iieneiaa aioai?ea: noai oa ia?niaeoeae ?icaeoeo”
(E?eaee ??a, 1993; Aeiiaoeuee, 1998), eiioa?aioe?yo iieiaeeo a/aieo
“Aeooaeuei? ieoaiiy aioai?ee oa aeieia??” (Aeiiaoeuee, 1995; Oa?e?a,
1996; Iaeana, 1997, Oa?nii, 1998), /eoaiiyo, i?enay/aieo iai’yo?
i?ioani?a I. E. ?aae (Aeiiaoeuee, 1998), a oaeiae ia can?aeaiiyo
Aeiiaoeueeiai a?aeae?eaiiy Oe?a?inueeiai aioai?/iiai oiaa?enoaa
(1992-1998).

Ioae?eaoe??

Ca aeena?oaoe?eiith oaiith iioae?eiaaii 5 noaoae ? 4 oace. C ieo 1
noaooy ? 1 oace a ni?aaaoi?noa?.

Ianya ? no?oeoo?a ?iaioe

Aeena?oaoe?y neeaaea?oueny c anooio, 7 ?icae?e?a, aeniiae?a, nieneo
e?oa?aoo?e ? /ioe?ueio aeiaeaoe?a. ?iaioa aeeeaaeaia ia 208 n.
iaoeiiieniiai oaenoo, aeeth/a? 26 oaaeeoeue (27 n.), 25 ?enoie?a (15
n.). Nienie e?oa?aoo?e neeaaea?oueny c 160 aeaea?ae, o oiio /ene? 25
?iiiiaieo. A aeiaeaoeao iaaiaeeoueny aiioiaaiee nienie ?ineei a?aeaae?a
(14 n.), noaia eeaneo?eaoe?? ?ineeiiino? (4 c.), iaoa??aee iieueiaeo
iien?a ( 39 n.) oa aeai?eoie ?o ia?iaee o canoiniaaieo i?ia?aiieo
iaeaoao (2 n.).

I?IA?AIA, IA’?EO OA IAOIAeE AeINE?AeAEAIUe

Iniiaith aeine?aeaeaiiy ? a?iioaca, ui aio?iiiaaiia noeoean?y
?ineeiiino? a?aeaae?a nii/aoeo ia? cia/i? a?aei?iiino? a?ae i?ioean?a o
i?e?iaeiiio na?aaeiaeu?. Aea aeae?, c iaaeeaeaiiyi oiia a?aeaae?a aei
i?e?iaeieo, ?ineeii?noue a?aeaae?a iaaoaa? ciiaeueiiai oa?aeoa?o. Oiaoi
a noeoean?ei?e nenoai? ?aa?iio a?aeaoaa?oueny oi?ioaaiiy c aaeaaioeaieo
oa aai?eaaiieo aeaiaio?a iiai? i?aenenoaie, caeaoii? anei?ethaaoe
aeioiie a?aeaae?a.

I?ia?aiith aeine?aeaeaiue ia?aaeaa/aeinue aeyaeoe a?aeaaee, ia yeeo
noeoean?? ia ia?aoeiaeaeaea iiaeaeueoa ae?yeuei?noue ethaeeie. Oea
aeicaieeei a?aeoai?eoe iae?iia??i?oo iine?aeiai?noue naiino?eiiai
?icaeoeo oa?oiiaaiue. Ia io?eiaiiio oeaiio?iiieeei? ca ci?iith
oei?enoe/ieo oa oeaiioe/ieo iieaciee?a aecia/aii o?e noeoean?eieo
noaae??. Aiae?c ?ineeiiino? noaae?e aeicaiey? aeyaeoe ?o iniaeeaino?, a
ii??aiyiiy c i?e?iaeiith ?ineeii?noth — aecia/eoe iai?yi ?icaeoeo.
I?inoi?iaa no?oeoo?a aea/aeanue iaoiaeaie aiae?ca ?iciiae?eo.
I?inoi?iaee oa /aniaee ?iciiae?e aeae?a aecia/a? ?o noeoean?eiee
noaoon. Eeaneo?eaoe?y o?oioeaiic?a a?aeaae?a aeyaeea ?o
??ciiiai?oi?noue. Aea/aiiy ni?yaeaiino? aeae?a aeicaiey? aecia/eoe ?o
aca?iia?aeiioaiiy.

Aeine?aeaeoaaeenue a?aeaaee ?c iaii?ooaiith iiaa?oiath oe?a?inueei?
/anoeie Aeiiaano ia oa?eoi??? Aeiiaoeueei? oa Eoaainueei? iaeanoae.
A?aeiia?aeii aei i?iaaaeaii? eeaneo?eaoe?? aeioii?a ia 47 eii?/ieo
a?aeaaeao c?iaeaii iiene 312 i?iaieo ae?eyiie ?ici??ii 100 i2 oa
aeecueei 4000 iae?eiaeo ae?eyiie iaioiai ?ici??o, ui a?aeaeaathoue
aa?’thaaiiy ?ineeiiino? o /an? oa i?inoi??.

Aeei?enoiaoaaeenue ia?o?ooiee iaoiae aeine?aeaeaiue, iaoiae i?iaieo
ae?eyiie, aeieiai-ii?oieiai-aaia?ao?/iee oiui, aea aieiaia oaaaa
iaaeaia iaoiaee/iei i?aeoiaeai, ye? a?oioothoueny ia nenoaiieia?? oa
iaoaiaoe/i?e noaoenoeoe? (Ie?eei, ?icaiaa?a, Iaoiiaa, 1989), ui ?ai?oa
iaeaea ia aaeeaaeenue uiaei oe??? ?ineeiiino?.

?aeiino?oeoe?th ?icaeoeo oa?oiiaaiue a?aeaae?a o noeoean?eiiio /an?
aeeiiaii ia?aoai?aiiyi i?inoi?iaiai ?o aa?’thaaiiy ia /aniaa. Aeey
oeueiai ia?aii ?i?oe?aeuei? oa oa?i?iaeuei? oa?oiiaaiiy, a ?aooa
iine?aeiaiino? a?aeoai?thaaeanue iaoiaeii eiiiiceoe?eii? i?aeeiaoe??,
iiaeeo?eiaaiei a?aeiia?aeii aei oiia a?aeaae?a. Iiaoaeiaaiee
oeaiio?iiieeei ?ineeiiino? a?aeaae?a ? aii??eei-noaoenoe/iith iiaeaeeth
?? noeoean??, ui noaiiaeoue iaeei c aoai?a nenoaiiiai aiae?ca
(?icaiaa?a, 1984) oa a?aee?eaa? iiaeeeaino? iiaeaeueoiai ?? aea/aiiy.
I?iaiae?ciaaii ?iaeaene iiae?aiino? oa o?oioeaiioe/ii? cia/ouino?.
Cai?iiiiiaaii oi?ioee oeeo ?iaeaen?a, i?enoiniaai? aei oiia a?aeaae?a.
?iciiae?e ?iaeaena o?oioeaiioe/ii? cia/ouino? aeae?a acaeiaae a?n?
noeoean?eiiai /ano aea/aany a?aae??ioiei aiae?cii c aeei?enoaiiyi
iaoaiaoe/iiai caeaaeaeoaaiiy.

Iaae? noeoean?eieo noaae?e aecia/aii ca ci?iaie aeaeiaiai neeaaeo oa
eiai iiae?ai?noth ia ae?eyieao. Nenoaiaoe/iee oa oeiieia?/iee aiae?c
oei?enoe/ieo eiiieaen?a noaae?e aeeiiaii ca aeiiiiiaith aaoiiaoeciaaii?
aace aeaieo oei?e ?aa?iio “Aio?iiiaaiia o?ainoi?iaoe?y oei?e” (Ao?aea,
Inoaiei, 1993); a- oa b-??ciiiai?oi?noue, oei ?iciiae?eo aeae?a o
i?inoi?? oa ?o ni?yaeai?noue ioe?ithaaeenue ca a?aeiia?aeieie
?iaeaenaie. Iica?/i?noue aea/aeanue iaoiaeii I. A?aea-Ni?oa ca ci?iith
aeenia?n?? o?aieyiiy aeae?a ia ae?eyieao ieiuath a?ae 0,1 aei 4 i2 , oa
ia o?ainaeoao. Aieea i?noeaciaoiaeaeaiiy ia oi?ioaaiiy ?ineeiiiai
iie?eao aecia/aii aeenia?n?eiei aiae?cii. Aeey eeaneo?eaoe?? aaeeoi
iaoiaee A?aoi-Aeaiea, “Ceoi/aiiy-4” oa A.?. Aaneeaae/a — A.I. Ii??ia
(Ie?eei, ?icaiaa?a, 1978), ye? a?aeiia?aeathoue oa?aeoa?o ?ineeiiino?.
Eeanoa?iee aiae?c aeae?a ca iiae?ai?noth ?o aeieia?? c?iaeaii iaoiaeii
ei?aeyoe?eieo ieayae (Aaneeaae/, 1969) oa iiaoaeiaith aeaiae?ia?ai
iaoiaeii a?oiiaiai na?aaeiueiai (Ie?eei, ?icaiaa?a, 1978).
Aca?iia?aeiioaiiy aeae?a aea/aeenue ca ci?iaie ?o ni?yaeaiino? i?e
ca?eueoaii? ae?eyiie (A?aea-Nieo, 1968), oa ei?aeyoe?eiei aiae?cii, ia
aac? yeiai ?ic?iaeaiee iaoiae aecia/aiiy eiieo?aioini?iiiaeiino? aeae?a
a aaaaoiaeaeiaeo o?oioeaiicao.

Aeei?enoiaoaaeenue iaeaoe i?ia?ai “Statgraphics plus” (SGC, 1996)
oa “MS Excel” (Microsoft, 1997).

EI?IOEEE O?CEEI-AAIA?AO?*IEE IA?EN ?AEIIA OA AEIEIA?*IEO OIIA A?AeAAE?A
AOA?EUeIEO OAOO

?aeii aeine?aeaeaiue ia noiae? iaiaaeaiee aea?aeaaiei ei?aeiiii, a ia
caoiae? nyaa? *a?aiiia?i?enueea. Ia i?aaei? a?i aeioiaeeoue aei
Aieiiaaoe, ia i?aii/? ioiieth? Eene/ainuee oa Eoaainuee. Oeae ?aeii i?e
?aeiioaaii? ca oaeoii?/ieie, aaieia?/ieie, aaiii?oieia?/ieie,
o?ceei-aaia?ao?/ieie, aaiaioai?/ieie, oei?enoe/ieie, e?eoa??yie
noaiiaeoue ?aeeio oa?eoi??th. Iaoai?ieia?/i? iieacieee ci?iththoueny
iinooiiai, ia ooai?thth/e ??ceeo a?aae??io?a (iai?eeeaae, ia iaaeao
?aeiia aeine?aeaeaiue ca?a?no?iaai? oae? oaiia?aoo?e: iaeneiaeueia — 39,
40, 41°N; i?i?iaeueia — -36, -37, -38, -39, -41°N (Ni?aai/iee ii eeeiaoo
NNN?, 1966-1969). Cia/ia /anoeia oa?eoi??? caeiyoa ooo/ieie
ianaaeaeaiiyie aai ?ioei /eiii aeei?enoiao?oueny ethaeeiith. Iacaeiaia
noaiiaa ?ineeii?noue, ui ?ai?oa ia?aaaaeaea, caa?aaeany eeoa o
caiia?aeieeao. Aaeeea e?euee?noue aeae?a iio?aao? ioi?iie. I?ioa a?ioa
caa?aaea nai???aei?noue ? ua aeineoue ??ciiiai?oia (Aeoaiaee, 1964;
Ao?aea, 1991).

Aeieia?/i? oiiae a?aeaae?a, cie?aia aaeao?/i?, aeineoue aeia?a aea/ai?,
? aiie ??cei a?aei?ii? a?ae oiia a i?e?iaeieo aeioiiao oa eieeaathoueny
a oe?ieeo iaaeao (I?iiuoeaiiay aioaieea, 1980). Oea a?aeaeaa?oueny o
neeaae? oa no?oeoo??, aeeiai?oe? ?ineeiieo oa?oiiaaiue a?aeaae?a
caeaaeii a?ae ?o ?icoaooaaiiy. I?iai? ieiu? caeeaaeaeenue ca noaiith, ui
a?aoiaoaaea eeaneo?eaoe?th aeioii?a a?aeaae?a ?

Oaaeeoey 1 Noaia ?icoaooaaiiy i?iaieo ae?eyiie o nenoai? aeioii?a ia
oaooieo a?aeaaeao

Iieiaeai- Aeniiceoe?y ? iieiaeaiiy ia iiia?a/iiio i?io?ethiy ia
iiac-aeiaaeiueiio
I?ai?/I?aaeaiueNo?aeCao?aei?io?eth-a*aaaaaaaaaaaOnueiaioa?no—14—7—10-
-1041na?aaeeia
1213111367710811227127o?iio161717111091467161011144Onueiai28304224162321
1625273228Onueiai100636287312· a – aa?o, a – na?aaeeia, a – iec noeeo

·

?icoaoiaai? oae, ui cia/iith i??ith ioiieththoue ??ciiiai?oi?noue
aeioii?a (oaae.1).

?INEEII?NOUe A?AeAAE?A OA ?? NOEOeAN?EIA AeEIAI?EA

Iaiao?aeiith oiiaith aea/aiiy ?ineeiiino? ? ciaiiy oei?e. A oiae?
aeine?aeaeaiue ia a?aeaaeao aeyaeaii 262 aeaee noaeeiieo ?ineei c 175
?iae?a oa 49 ?iaeei, 3 eean?a, 2 oei?a. Ca nenoaiaoe/iith no?oeoo?ith
oei?a a?aeaae?a a?ae??ciy?oueny ca?eueoaiiyi /enaeueiino? ?iae?a oa
?iaeei, ui neeaaeathoueny c ?oaea?aeueieo aeae?a c oe?ieith aeieia?/iith
aiie?ooaeith. Oea Chenopodiaceae Polygonaceae, Chenopodium, Hieracium,
Plantago, Poa, Rumex. ?iaeiaee eiao?oe??io oei?e a?aeaae?a noaiiaeoue
0,67, ui aeaco? ia cia/io ?ieue i?a?aoe?eieo aeaiaio?a a ?? noaiiaeaii?.
Aaia?ao?/iee aiae?c na?ae/eoue, ui e?euee?nii ia?aaaaeathoue aaeaaioeai?
oa oe?ieia?aaeuei? aeaee: aiea?eoe/i?, iaeaa?eoe/i?, ?a?iiaenuee?. Ca
aeeoo?aith oi?iith iaea?eueoa aai?e??ioio?o?a oa oa?io?o?a. Ca
oei?ioeaiioeiaie iaeaaaeeea?oeie ? noaiia? oa neiaio?iii? aeaee.

?ineeii?noue a?aeaae?a neeaaea?oueny c ?oaea?aeueieo oa
iai?a?oaea?aeueieo oa?oiiaaiue, caeaa?eueoiai c aeii?ioaaiiyi
iaei??/iee?a. Acaaae? aiia ca?aeiaia ca aeaeiaei neeaaeii,
a?ae??ciy?oueny ca oei?enoe/iith no?oeoo?ith a?ae ?aa?iiaeueii?, ui
aeeeeeaii, iaaii, niaoeeo?/ieie aeieia?/ieie oiiaaie.

Uia ?icaeyiooe oei?o oa ?ineeii?noue a?aeaae?a o i?ioean? ?o
?icaeoeo, 312 iien?a ?icoaoiaai? iaoiaeii eiiiiceoe?eii? i?aeeiaoe?? o
oeaiio?iiieeei, ui a?aeoai?th? noeoean?eio iine?aeiai?noue ?icaeoeo
oa?oiiaaiue a?aeaae?a. Ca ci?iaie aeaeiaiai neeaaeo a?ae??ciythoueny o?e
noaae?? noeoean?? c iaaeaie ia 25 oa 75 iaeeieoeyo i?aeeiaoe??. C ieo
ae?oaa noaae?y ioiieth? a?eueoo /anoeio iien?a. Aeey ia?aa??ee
oe?e?niino? oe??? noaae?? aecia/aia iiae?ai?noue i?ae aeieaie iien?a
ai?iaeiaae oeaiio?iiieeeia. Iiaoaeiaaiee aeaiae?eo a?aeiia?aea?
iine?aeiaiino? oeaiio?iiieeeio oa ia? a?eueo? cia/aiiy iiae?aiino? a
iaaeao noaae??. Iiae?e noeoean?eii? iine?aeiaiino? ia o?e noaae??
iniaeeai aeia?a iii?oiee a ?iciiae?e? aeae?a aeniei? ?yniino?. Ia ia?o?e
noaae?? iaeneiaeuei? cia/aiiy ?iaeaena niinoa??aathoueny o Oberna behen
(L.) Ikonn., Polygonum aviculare L.oiui,a iaeneioi ?iaeaena o aeae?a,
oa?aeoa?ieo aeey ae?oai? noaae??, iii?oii ci?uaiee aeae? ia
oeaiio?iiieeei?, acaeiaae yeiai ci?ith?oueny noeoean?eiee /an, ye o
Echium vulgare L. aai Centaurea diffusa Lam. Iaaea i?ae ae?oaith ?
o?aoueith noaae?yie ? iaio iii?oiith, ii/eia? i?iyaeyoeny
eiioeioaeuei?noue ?ineeiiino?. Aeae?a c aenieeie cia/aiiyie ?iaeaena
o?oioeaiioe/ii? cia/ouino? noa? a?eueoa, oea, iai?eeeaae, Galium
humifusum Bieb., Thesium arvense Horvatovsky, Achillea nobilis L. oiui.

Aeey ii??aiyiiy, ye eiio?ieueiee c?acie aeei?enoaii oei?o-icieyo
o?i/eua Iaeaeaiia aaeea (Ao?aea, 1991). Nooiiiue aio?iiiaaiiiai
iaaaioaaeaiiy ia ia? (ianeaaeueiiai, ?ae?aaoeieiiai) aaaaea?ii oiiiaei
aeey ?aaiiio. Ieiua oei?e-?cieyoa (10 aa) oa /enei ca?a?no?iaaieo aeae?a
(151) aeinoaoii aeey ii??aiyiiy c aiaeia?/ieie iieacieeaie a oiiaao
aoa?eueieo a?aeaae?a.

Aeey ioeiiee iiaeiaiinoi nenoaiaoe/ii? no?oeoo?e aeei?enoaii
eiaoioei?io ?aiaiai? ei?aeyoei? Eaiaeaea. Iiaoaeiaaiee ca iei aeaiae?eo
ia? eiiieio oi?io, aiaeiiaiaeii aei /a?aoaaiiy noaaeie. Oei?a-icieyo
Iaeaeaiia aaeea ia? iaeaieueoee ei?aeyoe?eiee ca’ycie c oei?ith
oa?oiiaaiue o?aouei? noaaei? (0,274), a ca’ycie c iiia?aaeiiie
noaaeiyie iineaaeth?oueny i?iii?oeieii aei ?o aiaeaeaeaiinoi
(0,116; -0,033). Aeaii oeiieiai/iiai aiaeico, ui anaai/ii
oa?aeoa?ecothoue i?ioean ?icaeoeo ?ineeiiiai iie?eao aiaeaaeia,
iiaeoaa?aeaeothoue aeyaeaio iineiaeiaiinoue ciii (oaae. 2). Iai?eeeaae,
ca aaia?aoi/iith no?oeoo?ith ii/aoeia? noaae?? noeoeani?
oa?aeoa?ecothoueny ia?aaaaeaiiyi aeae?a c oe?ieeie a?aaeaie, aea aeaei
niinoa?iaa?oueny iaaeeaeaiiy aei eiio?ieth. Iaoaii/iee neeaae ai?nueeeo
ii?iae, c yeeo neeaaea?oueny aiaeaae, iiiioii aieeaa? ia aeieiai/io
no?oeoo?o noeoeanieieo oa?oiiaaiue. *anoea eioioioia oaeai? aeua, i?ae o
eiio?ie? ? iaaioue ia o?aoie noaaei? noeoeani? cieaeo?oueny iacia/ii.
Aeieiai/iee niaeo? oei? noeoeanieieo noaaeie oe?oee, iiae o eiio?iei. Ca
a?iii?oieia?/ieie iieacieeaie, aoaeiaith iaaii?a, caaa?aeaiiyi
ioea?oeie, ni?aa?aeiioaiiyi oei?ioeaiioei?a oaeiae niinoa??aa?oueny
caeeaeaiiy i?ae ?ineeii?noth a?aeaae?a oa Iaeaeaiii? aaeee. A 40
aeiaaeeao oea i?iyaey?oueny a iiae?aiino? ca e?euee?noth aeae?a oa ca ?o
ni?aa?aeiioaiiyi a?aeiinii caaaeueii? e?eueeino?, a o 8 aeiaaeeao — ca
e?euee?noth aeae?a iaaiiai aeaiaioo, iai?eeeaae, ca iaeaa?eoe/iei
aaia?ao?/iei aeaiaioii. A?aenooi?noue /?oe?oi? iine?aeiaiino? ?ieiee
iaoiiaeaia noioanoe/ieie “ooiaie” oa iacia/iith e?euee?noth aeae?a oiai
/e ?ioiai oeio a?aaea aai ?ioiai aeaiaioa aiae?ca. Iiaeia c?iaeoe
aeniiaie, ui ?icaeoie ?ineeiiinoi aiaeaaeia aoaieueieo oaoo,
aiaeaoaa?oueny o iai?yio oi?ioaaiiy no?oeoo?e, i?eoaiaiii? ciiaeueiie
?ineeiiinoi, ui cacia? aio?iiiaaiiiai aieeao ia oiiiaiio aeey ?aaiiia
?iaii, c iacia/ieie

Oaaeeoey 2 Oeiieia?/ia no?oeoo?a oei?e noeoean?eieo noaae?e

I noaae?yII noaae?yIII noaae?yIaeaeaiia aaeeaIciaeaE?eue-e?noue
aeae?a%E?eue-e?noue aeae?a%E?eue-e?noue aeae?a%E?eue-e?noue
aeae?a%Aaia?ao?/iee aeaiaioIeth???aa?iiaeueiee2 11,8
3 3,5 2 2,6
1 0,7 Aiea?eoe/iee3 17,6
8 9,4 6 7,8
5 3,4 Iaeaa?eoe/iee7 41,2
34 40 34 44,1
57 38,3 ?a?iiaenueeee0 0
11 13 10 13
17 11,4
?a?iiaenueei-na?aaecaiiiii?nueeee1 5,9 5
5,9 4 5,2 4
2,6 I?e/i?iiii?nueei-na?aaecaiiiii?nueeee0
0 3 3,5 4 5,2
10 6,7 I?e/i?iiii?nueeee0
0 8 9,4 8
10,4 28 18,8
Oeaio?aeueii?a?ac?e-nueeee1 5,9 5
5,9 3 3,9 20
13,4 A?oia aaeaaioeaieo ?ineei3 17,6
8 9,4 6 7,8 7
4,7 Na?aaeiaeua aeeooy
Aa?iiaaeioio15 88,2 74 87,1
70 90,9 13993,3 Inaiioio1
5,9 4 4,7 2
2,6 6 4 Eioioio1
5,9 5 5,9 4
5,2 4 2,7
Aa?ia?ae?iiaaeio?o0 0 2 2,3
1 1,3 0 0
Aiaeiee ?aaeeiAoena?io?o3 17,6
10 11,8 9 11,7 17
11,4 Iaciena?io?o2 11,8
25 29,4 26 33,8 40
26,8 Ena?iiacio?o7 41,2
33 38,8 30 38,9
71 47,7 Aoiacio?o4 23,5
14 16,5 11 14,3
21 14,1 A?ae?iiacio?o1 5,9
3 3,5 1 1,3
0 0

aiaeoeeaiiyie, eio?? iiyniththoueny niaoeeoieith aeioiiia.

Iaeiei c iaeaaaeeea?oeo ia?aiao??a o?oioeaiic?a, ye neeaaeieo
aeeoocieo nenoai, ? ?o i?inoi?iaa no?oeoo?a. O a?eueoino? cae/aeieo
aeae?a a?aeaae?a i?e oeueiio ianooaa? ?iciiae?e ?aaoey?iee aai
aeiaaeeiaee, ui aeicaiey? aeei?enoaoe eiai i?e oi?ioaaii? ooo/ieo
o?oioeaiic?a. Eiioaa?iciee ?iciiae?e e?eueeio aeae?a iiaeia iiynieoe
iniaeeainoyie ?iciiiaeaiiy. No?oeoo?e a?eueoiai ?ici??o a
?iciiae?eaoaeae?a aeine?aeaeoaaeenue ia aeieia?/ieo i?io?eyo ??cieo
iai?yie?a. A oe?eiio niinoa??aa?oueny caeiiii??ia ci?ia iaeieo a?oi
aeae?a ?ioeie ca iniiaieie a?aae??ioaie na?aaeiaeua, ui aieeaa? ? ia
i?aeaeaeia? noinoiee.

C iaoith aeyaeaiiy aeeiai?ee ?iciiae?eo aeae?a ca noeoean?eiei
/anii aecia/aii ?iaeaen ?iciiae?eo aeey aeae?a ia ??cieo noaae?yo.
Noai?aia ocaaaeueiaia noaia ci?ie ?iciiae?eo aeae?a a?ae ?o iiyae aei
cieeiaiiy c oeaiica (?en.1). Nii/aoeo ?iciiae?e aeiaaeeiaee, aeae?,
ca?aeii c aeaeiaeie aeanoeainoyie oa a?aeiia?aeiino? oiia aei aeiia
aeaeo,a?i iiaea noaoe ?aaoey?iei aai, /ano?oa, eiioaa?iciei. A ia?aae
cieeiaiiyi aeaeo eiai ?iciiae?e ciiao noa? aeiaaeeiaei.

?en. 1 Noaia ci?ie ?iciiae?eo aeaeo a caeaaeiino? a?ae oace eiai
?nioaaiiy a oa?oiiaaii?

Aeey aea/aiiy aiiiaai?oaoo o?oioeaiic?a aeine?aeaeoaaany ?iciiae?e
aaee?aeiai? aeenoaioe?? i?ae iienaie ae?eyiie ?ici??ii 1i2 a iaaeao
oeueiai oeaiico. sse ? i/?eoaaeinue, aiiiaai?oao ?ineeiieo oa?oiiaaiue
a?aeaae?a iaeaeuee ia o?ao?e noaae?? (h=1,59). A?aeiinii aenie? cia/aiiy
ia ia?o?e noaae?? (h=1,4) coiiaeai? iacia/iith e?euee?noth aeae?a oa
ni?iuaiiyi no?oeoo?e.

Noeoean?eiee noaoon aeae?a oei?e a?aeaae?a

Ca ciaoiaeaeaiiyi oa aeeiai?eith ?iaeaena o?oioeaiioe/ii?
cia/ouino? ia oeaiio?iiieeei? a?aeaae?a, ?iciiae?eii, iiae?ai?noth
aeieia?? oa ?ioeie aeanoeainoyie aecia/aii noeoean?eiee noaoon aeae?a
a?aeaae?a, yeee oa?aeoa?eco? /aniaee aniaeo iiaaae?iee aeae?a. Aeey
oeueiai iaoiaeii aeenia?n?eiiai aiaeeco ia?aa??aii, yeith i??ith
i?noeaciaoiaeaeaiiy oeaiico aieeaa? ia oi?ioaaiiy ?ineeiiino?. Caaaeueia
neea aieeao aeniiceoe??, aenioe ?icoaooaaiiy oa nooiaith aio?iiiaaiii?
o?ainoi?iaoe?? ioi/oth/i? ?ineeiiino? ia ia?aaeuo? 20 a?aenioe?a. Oea
aeicaiey? ?icaeyaeaoe oeaiio?iiieeei ?ineeiiino? a?aeaae?a ye ?aeeiee
noeoean?eiee ?yae.

Noeoean?eiee noaoon aeae?a iicia/aii oae: ? — (i?iia?e) aeaee
ia?oi? noaae??, I — (iaae?oie) aeaee ae?oai? (na?aaeiuei?) noaae?? oa F
— (o?iae?noe) aeaee o?aouei? noaae??, s — (noioanoe/i?) aeiaaeeia?
aeaee. Aaeeeeie e?oa?aie iicia/athoueny aeaee, caeaoi? aeii?ioaaoe a
oiiaao a?aeaae?a, iaeeie — on? ?io?. A iienao ia oeaiio?ieiieei?
ia?aaaaeathoue aeaee, ye? cono??/athoueny ai?iaeiaae e?eueeio noaae?e
(57): ?ioaa?oth/? (aeii?iaioe) oa iaeo?aeuei? (anaeoaoi?e). Aeiaaeeiaeie
? 48 aeae?a, oa 18 — aeeoa?aioe?aoi?e (oa?aeoa?i? aeey ie?aii? noaae??
aeaee). Noeoean?eiee noaoon on?o aeae?a oeaiio?iiieeeio iicia/a?oueny
oi?ioeith: 8P9p16M60m25F37f48s, yea a?aeaeaa? iioaioe?ei? iiaeeeaino?
noeoean?eii? nenoaie oa?eoi??? ia on?o noaae?yo.

Ia iniia? aeei?enoaiiy noeoean?eiiai noaoono ?ic?iaeaia iaoiaeeea
o?oi?iaeeeaoe?? no/aniiai noeoean?eiiai noaio oa eiai ianooiii?
aeeiai?ee. Oea aea? ciiao c’ynoaaoe i?e/eie, ui aieeaathoue ia ?icaeoie
?ineeiiino?.

EEANEO?EAOe?ss ?INEEIIEO OA?OIIAAIUe A?AeAAE?A

?ineeii?noue a?aeaae?a aoa?eueieo oaoo o aaaaoueio aeiaaeeao
oa?aeoa?eco?oueny eeeiaeuei?noth ca iniiaieie a?aae??ioaie na?aaeiaeua
oa eiiieaeni?noth, ui coiiaeaia neeaaeiei ?aeue?oii, eieaeueieie
a?aei?iiinoyie neeaaeo ii?iaee, ina?aaeeaie ai??iiy oiui. Aiane?aeie
oeueiai iiaeeea?noue eeaneo?eoaaoe ?ineeii?noue a?aeaae?a ? iaaenieith.
Oae, canoinoaaiiy iaoiaea A.?. Aaneeaae/a — A. I. Ii??ia aeey iien?a
ia?oi? oa ii/aoeo ae?oai? noaae?? aeaei iaeiiaa?oeiio e?eao ?iciiae?eo,
ai aiie ooai?ththoue iaeio a?oio. A oa?oiiaaiiyo e?ioey noeoean??
e?euee?noue a?oi c?inoa?. I?i oea na?ae/aoue ? ?acoeueoaoe canoinoaaiiy
aaoiiaoe/ii? eeaneo?eaoe?? ca iaoiaeii “Ceoi/aiiy-4”, ui aeaco? ia
iineeaiiy aeene?aoiino? ?ineeiiino? c /anii.

?ai?oa aeey eeaneo?eaoe?? ?ineeiiino? a?aeaae?a canoiniaoaaany eeoa
aeii?iaioiee i?aeo?ae. A?i ia ? iaee?auei aeey ?ineeiieo oa?oiiaaiue
a?aeaae?a, ye? ooai?ai? caeaa?eueoiai iaeii- oa iaei??/ieeaie.
I?ieea?noue ?o ni?aa?aeiioaiue a ??ci? ?iee oa ai?iaeiaae iaeiiai
aaaaoaoe?eiiai naciio a?aei?/aeanue ? aeey ?ioeo oei?a aio?iiiaaiii?
?ineeiiino?. Aeaeiaee neeaae oa?oiiaaiue a?eueo iino?eiee. Oiio aeey
eeaneo?eaoe?? ia?aii iaoiae A?aoi-Aeaiea, ui oe?iei aaeeaa?oueny ca?ac
i?e aea/aii? neiaio?iiii? ?ineeiiino?. Aeyaeaii 7 anioe?aoe?e, 4 nithce,
ui a?aeiinyoueny aei 3 ii?yaee?a 3 eean?a (Nieiiaoa, 1995), 2
anioe?aoe?? iaaiaeyoueny aia?oa (Persicario — Obernetum behenii; Poo
compressae — Hieracietum virosae). A?aei?ii?noue oiia a?aeaae?a a?ae
i?e?iaeieo aeeeeeaea ci?io /anoeie ae?aaiinoe/ieo aeae?a neioaenii?a.

Ia ia?o?e oa ii/aoeo ae?oai? noaae?? noeoean?? ia?aaaaeathoue
aeaee, ye? ia ? ae?aaiinoe/ieie aeey yeiainue eeano ?ineeiiino? Oe?a?ie,
iai?eeeaae Oberna behen, Persicaria maculata oiui. Aeae? ii/eiathoue
iaioaaoe aeaee eeano Artemisietea vulgaris oa ii?yaeeo
Meliloto-Artemisietalia absinthii, ui ?ieiee a?aeie?aieth?oueny a eean
Meliloto-Artemisietea absinthii, oa?oiiaaiiy yeiai ? noaae?yie noeoean??
o iai?yi? oi?ioaaiiy oa?oiiaaiue eeano Festuco-Brometea. Ae?aaiinoe/i?
aeaee oeueiai eeano ? eean?a Agropyretea repentis oa Festucetea
vaginatea ia?aaaaeathoue ia caeeth/i?e c iayaieo noaae?e noeoean??.

NI?ssAEAI?NOUe I?AE AEAeAIE AEUEO ?INEEI O OIAe? AIO?IIIAAI-II?
NOEOeAN?? ?INEEIIINO? IA A?AeAAEAO AOA?EUeIEO OAOO

Aea/aiiy ca’yce?a i?ae aeaeaie ? iaeiei c aoai?a aiae?ca
o?oioeaiioe/ieo nenoai. Oea aea? aeo?aei? aeai? aeey aea/aiiy ?o
no?oeoo?e, aca?iia?aeiioaiue aeae?a, aeyaeaiiy aeieia?/ii iiae?aieo a?oi
aeae?a oa aeei?enoiao?oueny i?e eeaneo?eaoe??.

Aeeoa?aioe?aoe?th aeae?a ia aeieia?/ii iiae?ai? a?oie cae?eniaii
iaoiaeii ei?aeyoe?eieo ieayae, ui aeicaieeei ia’?aeiaoe a a?oie aeaee
ia?oeo aeaio noaae?e noeoean??, oa ca’ycaoe ?o eaioethaaie c i?ii?aeieo
aeae?a. Aea aeae? iaaeoi aaeeea e?euee?noue ca’yce?a oneeaaeith? aiae?c,
oaeiae ia aeei?enoiaothoueny a?ae’?ii? cia/aiiy eiao?oe??io?a. Oiio
iiaeaeueoee aiae?c ?iciiae?eo aeae?a ia aeieia?/ii iiae?ai? a?oie ca ?o
ni?yaeai?noth i?iaiaeeany iaoiaeii aeaiae?ia?aie.

Aeaiae?ia?aia iiaoaeiaaia aeey 64 iaeiioe?ai?oeo aeae?a. Iniiaiei
/eiieeii, ui aieeaa? ia aeeoa?aioe?aoe?th aeae?a ia a?oie ca
iiae?ai?noth ?o aeieia?? ? ?iciiae?e aeae?a a i?inoi?? oa noeoean?eiiio
/an?. Ia aeaiae?ia?ai? iiaeia, iai?eeeaae, ?aeaioeo?eoaaoe a?oie aeae?a,
ui aeii?iooue ia ie?aieo noaae?yo, aai ae?oai?yaei? aeaee oeeo noaae?e.

Aeei?enoaiiy ia?oe?aeueieo eiao?oe??io?a ei?aeyoe?? aeicaieeei
?icaeyiooe ca’ycie i?ae aeaeaie i?e a?aenooiino? aieeao ?ioeo aeae?a.
Aeaiae?ia?aia, iiaoaeiaaia ca oeeie aeaieie, a?aeaeaa? iiae?ai?noue
aooaeieia?/ieo, a ia o?oioeaiioe/ieo, aeiia aeae?a.

I?e aea/aii? ?ineeiiino? ia ae?oa?e noaae?? noeoean??
(na?aaeiueia?/i? a?aeaaee) iaoiaeii I. A?aea-Ni?oa (ca ni?yaeai?noth ia
ae?eyieao ??ciiai ?ici??o) onoaiiaeaii, ui a?ia?ianya aeieia?/ii? i?o?
Artemisia absinthium L. a cia/i?e i??? ni?aiaaea? c a?ia?i?inoi?ii
aea/aieo ae?eyiie ? oeae aeae caeaoai ni?a?nioaaoe c a?eueo?noth
aeine?aeaeaieo aeae?a. Ia ao?a/athoue a?ae’?iii? ei?aeyoe?? c
a?eueo?noth ?ioeo aeae?a i?e aeei?enoaieo ?ici??ao ae?eyiie Melilotus
officinalis (L.) Pall. oa Hieracium robustum Fries. O a?eueoino? ae
aeiaaee?a ?c ca?eueoaiiyi ae?eyiie c?inoa? ? ei?aeyoe?y i?ae aeaeaie.
Oea na?ae/eoue, ui ca’ycie aeae?a coiiaeaiee ia?aaaaeii ?o aeieia?/ieie
aeiiaaie.

Uia iaoe ia o?eueee ye?nio, a e e?euee?nio oa?aeoa?enoeeo
aca?iia?aeiioaiue aeae?a ?ic?iaeaii iaoiae aea/aiiy a?aeiinii?
eiieo?aioicaeaoiino? aeae?a a oiiaao aaaaoiaeaeiaeo o?oioeaiic?a. Aeey
oeueiai aeei?enoai? eiao?oe??ioe ei?aeyoe?? oa ia?oe?aeueii? ei?aeyoe??
i?ae aeaeaie, ui aeicaiey? onoiooe aieea on?o ?ioeo aeae?a. ??cieoey
i?ae ei?aeyoe?yie oa ia?oe?aeueieie ei?aeyoe?yie i?iii?oe?eia /ooeeaino?
aeaeo aei oeueiai aieeao, ? yeui ??cieoey aei??aith? ioeth, oi
/ooeea?noue aeaeo (Ksen) i?i?iaeueia. Aeey iaea?eueo /ooeeaiai aeaeo
iaeneiaeueia ??cieoey iiaeiia aei??aithaaoe noi? eiai ei?aeyoe?e (rij)
oa ia?oe?aeueieo ei?aeyoe?e (rij·n). Aieea ie?aiiai aeaeo ia
aca?iia?aeiioaiiy ?ioeo aeae?a (Kinf) ciaoiaeeoueny ye ??cieoey
na?aaei?o /ooeeainoae aeae?a i?e onoiaiiio aieea? on?o aeae?a (`Ksen) oa
i?ae aieeaii oeueiai aeaeo (`Kss). Cai?iiiiiaai? oi?ioee iathoue aeaeyae
:

n
n

Ksen(i) = S1/2 rij — rij·n1/2 / S(1/2rij1/2+ 1/2rij·n1/2),
Kinf(i) = 1/2`Ksen -`Kss1/2

j=1 j=1

Noi?nii oe? eiao?oe??ioe neeaaeathoue oa?aeoa?enoeeo
eiieo?aioini?iiiaeiino? aeae?a.

Aaaaeaiei iaoiaeii aea/aeenue aca?iia?aeiioaiiy aeae?a ia ia?o?e oa
o?ao?e noaae?yo noeoean?? ? ia ii/aoeo, a na?aaeei? oa a e?ioe? ae?oai?
noaae??. Oaeiae ?icaeyaeaeanue aeeoa?aioe?aoe?y aeae?a ia aeieia?/ii
iiae?ai? a?oie.

?en. 2 Aeaiae?ia?aie eeanoa??caoe?? aeae?a ia ia?o?e noaae??. A — ca
ei?aeyoe?yie, A

— ca ia?oe?aeueieie ei?aeyoe?yie,

1.Oberna behen (L.) Ikonn.

2.Persicaria maculata (L.)

3.Polygonum aviculare L.

4.Fallopia convolvulus (L.) A.Love7. Rumex crispus L.

8. Reseda lutea L.

9. Erucastrum armoracioides (Czern.ex Turcz.) Cruchet5.Ambrosia
artemisiifolia L.

6.Digitaria sanguinalis (L.) Scop.10.Amaranthus albus L.

11.Salsola australis R.Br.

Ia ia?o?e noaae?? iiaoaeiaai? aeaiae?ia?aie ca neeaaeii eeanoa??a
ia aeoaea a?ae??ciythoueny (?en.2). Oea, ye ? iaaenie? cia/aiiy
eiao?oe??io?a /ooeeaino? oa aieeao, aeaco? ia a?aenooi?noue iii?oiiai
aieeao i?aeaeaeiaeo aca?iia?aeiioaiue ia oi?ioaaiiy ?ineeiiiai iie?eao
ia oe?e noaae??. Ia ii/aoeo ae?oai? noaae?? oiiae aeaui iie?ioothoueny,
?icoe?th?oueny aeaeia?e neeaae oa?oiiaaiue, c?inoa? ? cia/aiiy oeeo
eiao?oe??io?a — c?inoa? aieea i?aeaeaeiaeo aca?iia?aeiioaiue. Aea
iioe?aiiy aeae?a ia?oi? noaae?? ooo a iacia/i?e i??? coiiaeaii ?o
aca?iiae??th. A?eueo eiieo?aioini?iiiaei? a oeeo oiiaao aeaee iino?eii
ci?iththoueny. Ia o?ao?e noaae?? c?inoa? iai?oaeai?noue i?aeaeaeiaeo
aca?iia?aeiioaiue, na??aeia cia/aiiy eiao?oe??ioo /ooeeaino? ia?aaeuo?
0,5. Aeaee, ui aeii?ioaaee ?ai?oa, aeoeneathoueny o ii?ooai?
i?noeaciaoiaeaeaiiy, aiane?aeie /iai aiie noathoue iaio /ooeeaeie, aea
aieea ?o iaaea? ? iaaeae?. A?oie iiae?aieo ca aeieia??th aeae?a iino?eii
ci?iththoueny. Iiyaa iiaeo, a?eueo eiieo?aioicaeaoieo aeae?a caeeaeo?
aeaee, ui aoee ?ai?oa a?aeiinii a?aeaeaeaieie ca aeieia??th.

Eaneaaeiee aoaeo aio?iiiaaiii? noeoean?? ?ineeiiino? a?aeaae?a
aoa?eueieo oaoo

Ai?iaeiaae ia?aeiii? noeoean?? ia a?aeaaeao aoa?eueieo oaoo
niinoa??aa?oueny oae caaiee “eaneaaeiee aoaeo”, yeee iieyaa? a
ia??aiii??iino? iiyae iiaeo aeae?a (?en.3) oa iniaeeainoyo ci?ie ?o
o?oioeaiioe/ieo ?ieae. Iia? aeaee c’yaeythoueny oa aeinyaathoue
iaeneiaeueii? o?oioeaiioe/ii? cia/ouino? a?oiaie, a yeeo aiie iathoue
aeieia?/i? aeiiae, ye? aeiiiaiththoue iaeei iaeiiai ? iieaaoothoue ?o
ni?a?nioaaiiy o o?oioeaiic?. Aei oeueiai ni?e/eiththoue e?eueea
aca?iiiia’ycaieo i?ioean?a. Nii/aoeo ae??oaeueia cia/aiiy iathoue
aa?ioe/i? /eiieee, a c oi?ioaaiiyi o?oioeaiic?a, ia ?o no?oeoo?o ana
a?ae/ooi?oa aieeaathoue aca?iia?aeiioaiiy ?ineei.

Iaa?oue iacia/ia iaiaeii??aei?noue a noeoean?eieo ci?iao ?ineeiiiai
eiioeiooio aoaea iineethaaoeny oa /?oe?oa i?iyaeyoeny i?e aca?iiae??
aeae?a o o?oioeaiic? oa c aeioiiii. I?noeaciaoiaeaeaiiy ? aeeiai?/ia
nenoaia c i?e?inaeo?a, ia ??ai? yeeo i?iyaeythoueny aeieia?/ii cia/ou?
aeey ?ineei a?aei?iiino? /eiiee?a. O eiaeai iiiaio /ano ?iciiae?e
cia/aiue aeieia?/ieo /eiiee?a ca oeeie i?e?inaeoaie a?aeiia?aea?
aeca?iiiiae?ai?e oi?i?. O i?ioean? ?icaeoeo aeioiio e?euee?noue
i?e?inaeo?a c a?aie/ieie cia/aiiyie /eiieea, ye? a?aeiia?aeathoue
iai?yio noeoean?eiiai ?icaeoeo, ca?eueoo?oueny, a c ?ioeie cia/aiiyie —
ciaioo?oueny. Aeca?iiiiae?aia e?eaa ?iciiae?eo cia/aiue aeieia?/ieo
/eiiee?a ca i?e?inaeoaie, “ia?ai?uo?oueny” a iai?yio, a yeiio
ci?iththoueny oe? /eiieee. Eiee eiiieaen i?e?inaeo?a i?ioiaeeoue e??cue
iaeanoue aeieia?/iiai iioeioio ie?aiiai aeaeo c aeca?iiiiae?aiei
?iciiae?eii ?ineei ca ?o aeiiaaie, oe? e?ea? iaeeaaeathoueny ? e?ioeaaa
e?i?y, ui aecia/a? iaeneiaeueii iiaeeeao ?yni?noue aeaeo, ia? a?eueo
e?oo? /anoee i?aeeiio oa niaaeo, i?ae aoaeue-yea c aeo?aei?o e?i?e,
ine?eueee eiia??iino?, ye? aiie a?aeia?aaeathoue, i?e oeueiio
ia?aiiiaeothoueny.

?en. 3 ?iciiae?e ?iaeaena o?oioeaiioe/ii? cia/ouino? (Y) acaeiaae a?n?
eiiiiceoe?e-

ii? i?aeeiaoe?? (O) aeey aeae?a: A — ia?oi? oa B — ae?oai?
noaae?? noeoean??

?ineeiiino?

Oiiai? iicia/aiiy: 1 — Oberna behen, 2 — Ambrosia artemisiifolia, 3 —
Erucastrum armoracioides, 4 — Polygonum aviculare, 5 — Fallopia
convolvulus, 6 — Persicaria maculata, 7 — Rumex crispus, 8 — Salsola
australis., 9 — Reseda lutea, 10 — Amaranthus albus, 11 — Digitaria
sanguinalis, 12 — Echium vulgare, 13 — Centaurea diffusa Lam., 14 —
Artemisia absinthium, 15 — Melilotus officinalis, 16 — Chenopodium
album L., 17 -Anisantha tectorum (L.) Nevski, 18 — Rumex stenophyllus
Ledeb., 19 — Sisymbrium altissimum L., 20 — Diplotaxis tenuifolia (L.)
DC, 21 — Hieracium virosum Pall., 22 — Fumaria schleicheri Soy.-Willem.,
23 — Acer negundo L., 24 — Lactuca serriola L., 25 — Sinapis arvensis
L., 26 — Sisymbrium loeselii L.

I?aeaeaeiaa eiieo?aioe?y, ye ? ci?ie aeieia?/ieo oiia, i?ecaiaeeoue
aei oi?ioaaiiy eaneaae?a o iiya?, aeii?ioaaii? oa cieeiaii? iiaeo a?oi
aeae?a. O i?iia?ieo oa?oiiaaiiyo i?e iae?e e?eueeino? aeae?a, ?o
noeeueiino? aei aa?aaaoe?? oa a iaiaeii??aeiiio na?aaeiaeu? i?e
eiieo?aioe?? aeaee iiaeooue ni?a?nioaaoe i?e cia/iiio ia?ae?eaaii?
aeieia?/ieo i?o. C /anii e?euee?noue aeae?a c?inoa? ? eiieo?aioe?y noa?
aeeoociith. I?e eiieo?aioe?? c iiaeie aeaeaie, o?oioeaiioe/ia ?ieue
aeae?a, ui aeii?ioaaee ?ai?oa, ciaioo?oueny. Eiieo?aioini?iiiaei?o?
aeaee aeoeneothoue ?o o aeayeeo aei??ao a?ia?i?inoi?o, aiane?aeie /iai
aea?eueiy?oueny a?aeiia?aeiee a?ia?ia’?i ? a ?ioeo aei??ao. Ana oea
coiiaeth? iiyao iiaeo aeae?a aai iineeaiiy ?ie? oeo, ui ?nioaaee ?ai?oa,
iai?eeeaae, Linaria vulgaris L., Daucus carota L., ye?, c’yaeyth/enue ua
ia ia?oeo aeaio noaae?yo noeoean??, caa??aathoue iacia/io ?yni?noue aei
o?aouei? noaae??. Aiie, aoaeooue, iiaeeeai, aoaeoeai?oa aeei?enoiaoaaoe
?ano?ne o yeiionue ?ioiio aei???, i?ae aeaee, ui aeii?ioaaee ?ai?oa.
Aiane?aeie oeueiai o?oioeaiioe/ia ?ieue aeae?a, ye? iaee ?ai?oa cia/io
?yni?noue, i?iaeiaaeo? cieaeoaaoeny. Aiaeii/an a?aeaoaa?oueny
aiaeia?/iee i?ioean aca?iiae?? c ?ioeie aeaeaie o ?aoo? aei???a
a?ia?i?inoi?o. Caeaaeii a?ae naciiieo aai aeiaaeeiaeo eieeaaiue aeayeeo
ia?aiao??a aeioiio, aeieia?/i? /eiieee noathoue e?i?ooth/eie a?aeiinii
ie?aieo aeae?a, ui i?ecaiaeeoue aei ciaioaiiy ?o ?yniino?.
Aiaeia?/iiai, aea i?ioeeaaeiiai ca iai?yiii aieeao, caciathoue aeaee, ui
o?eueee-ii c’yaeeeny o o?oioeaiic?, c aieo aeae?a c? noaeeie
aca?iia?aeiioaiiyie. Ana oea i?iyaey?oueny o ii?ooaiiyo ??aiii??iino?
ci?ie aeae?a o noeoean??, oiaoi iiya? ?o eaneaae?a — noeoean?eieo
noaae?e.

Aeniiaee

1. Aiae?c no/aniiai noaio aea/aiino? ?ineeiiiai iie?eao a?aeaae?a oaoo
Aeiiaano iieacaa oe?e?ni?noue oe??? aio?iiiaaiii ia?aoai?aii? o?oia?ioe,
ui aeicaiey? aeei?enoaoe iaoiaee aea??eiaiai aeine?aeaeaiiy c ianooiiith
aeno?aiieyoe??th ia anth oa?eoi??th Aeiiaano. sse?ni? iaoiaee ioe?iie
?ineeiiiai iie?eao, ui aeei?enoiaoaaeenue aei oeueiai, ae/a?iaee naaa, ?
aeey iiaeeaeaiiai aeine?aeaeaiiy iaiao?aeii aeei?enoaiiy no/anieo
noaoenoe/ieo iaoiae?a oa nenoaiiiai aiae?ca.

2. Ia iniia? aea/aiiy 312 i?iaieo ieiu ia 47 oa?eeiiao, ye?
ioiieththoue onth ??ciiiai?oi?noue aeioii?a a?aeaae?a, iiaoaeiaaii
oeaiio?iiieeei ?ineeiiino?, ui a?aeia?aaea? /aniaee aniaeo ci?ie
?ineeiiino? c o?aooaaiiyi o?ainoi?iaoe?? ?o i?inoi?iaiai aa??thaaiiy. Ia
oeaiio?iiieeei? ca oei?enoe/ieie ? oeaiioe/ieie e?eoa??yie aeae?eai? o?e
noaae?? noeoean??, ui ? aeaiaioaie iaeiiai ??aiy oe?e?niino?.

3. Io?eiai? i?e nenoaiaoe/iiio ? oeiieia?/iiio aiae?c? noeoean?eieo
noaae?e aeai?, ii??aiyiiy c i?e?iaeieie oei?aie oa ?acoeueoaoe
oei?enoe/ii? eeaneo?eaoe?? ?ineeiiinoi iieacaee, ui ?ineeii?noue
aiaeaaeia ?icaeaa?oueny o iai?yi? oi?ioaaiiy no?oeoo?e, i?eoaiaiii?
ciiaeueiie ?ineeiiinoi, ui cacia? aio?iiiaaiiiai aieeao ia oiiiaiio
aeey ?aaiiia ?iaii.

4.I?inoi?iaee ?iciiae?e aeae?a o o?oioeaiicao ia a?aeaaeao ia?
eiioaa?iciee oa?aeoa? ca oiiae, yeui ianooaa ?icaeyaeo ni?aiaaea? c
ieyieno?noth ?iciiae?eo /eiiee?a na?aaeiaeua. *ano?oa oea ni?aiaaea? c
?ici??aie ae?eyiie, iaeiioeiieo ca oa?aeoa?ii ?aeue?oa. Ona?aaeei? oeeo
ae?eyiie oa?aeoa? ?iciiae?eo aecia/a?oueny nooiaiai aaeaioaoe?? aeaeo,
oa?aeoa?ii ?iciiiaeaiiy oa /aniaith oacith ?nioaaiiy aeaiiai aeaeo o
oeaiic?; yeui oiiae i?noeac?inoaiiy iaaeeaeathoueny aei iioeiaeueieo,
?iciiae?e aeaeo oyae?? aei aeiaaeeiaiai aai ?aaoey?iiai. Ia ??ai? oe?eeo
oa?oiiaaiue aiiiaai?oao i?aeaeuo?oueny i?ae e?iaoeue noeoean??.

5. Ca ?acoeueoaoaie aeenia?n?eiiai aiaeeco oeaiio?iiieeei ?ineeiiino?
a?aeaae?a oyaey? ?aeeiee noeoean?eiee ?yae, oa?oiiaaiiy yeiai a ??cieo
i?noeac?inoaiiyo iia’ycai? iinooiiaeie ia?aoiaeaie. Noeoean?eiee noaoon
aeae?a aecia/aii ye i?noea ? cia/ei?noue o oeueiio ?yae?. A?i
coiiaeth?oueny oaiaeaioe?yie ?icaeoeo oa?oiiaaiiy oa /eiieeaie, ui ia
ieo aieeaathoue.

6. Eeanoa?iei aiae?cii aeae?eai? aeieia?/ii noiae? a?oie aeae?a,
iiae?ai?noue yeeo i?e aeei?enoaii? ei?aeyoe?eii? iao?eoe? cia/iith
i??ith coiiaeaia ?o ?iciiae?eii o /an? oa i?inoi??, a ia iniia?
ia?oe?aeueieo ei?aeyoe?e aeyaey?oueny iiae?ai?noue aeieia?? aeae?a i?e
aeaeaeaiiio aieea? i?aeaeaeiaeo a?aeiioaiue. Ci?ia aeieia?/ieo oiia a
oiae? noeoean?? a?aeia?aaea?oueny oaeiae a iiae?e? aeae?a ia a?oie.
Iiyaa aeae?a ianooiii? noaae?? noeoean?? a?aeiinii caeeaeo? aeaee
iiia?aaei?o noaae?e.

7. Ia ??cieo noaae?yo noeoean?? aecia/ai? iieacieee /ooeeaino? ? aieeao
iae?yni?oeo aeae?a, ui a noeoiiino? aea? ioe?ieo ?o a?aeiinii?
eiieo?aioicaeaoiino?. Ci?ie ?o o oiae? noeoean?? iiacycai? c? ci?iith
noaio aeaeo a oa?oiiaaiiyo: aai ccyaeythoueny a?eueo eiieo?aioicaeaoi?
aeaee, aai ae ci?ia aeieia?/ieo oiia i?ecaiaeeoue aei i?aeaeuaiiy
eiieo?aioicaeaoiino? iayaieo aeae?a. Oeueiio a?aeiia?aea? caaaeueia
c?inoaiiy iai?oae i?aeaeaeiaeo aca?iia?aeiioaiue, ye? ia o?ao?e noaae??
a?ae?a?athoue aecia/aeueio ?ieue o no?oeoo?? oa?oiiaaiue.

8. ?ineeii?noue a?aeaae?a oa?aeoa?eco?oueny eiioeioaeuei?noth ?
ia/?oeeie iaaeaie oa?oiiaaiue, i?aaenoaaeaia 7 anioe?aoe?yie, ?c yeeo
aea?: Persicario — Obernetum behenii ? Poo compressae — Hieracietum
virosae – iienai? aia?oa, ui a?aeia?aaea? nai???aei?noue ii/aoeiaiai
aoaio oi?ioaaiiy ?ineeiiino? ia a?aeaaeao. Ianooiia iaaeeaeaiiy aei
ciiaeueii? ?ineeiiino? ae?aaea?oueny o i?aeaeuaii? oe?e?niino?
oa?oiiaaiue oa a o??ciiiai?oiaii? neeaaeo anioe?aoe?e.

9. Noeoean?y ?ineeiiino? ia a?aeaaeao oaoo eiio?ieth?oueny aeeiai?/iei
eiiieaenii iayaieo aa?ioe/ieo /eiiee?a ? i?aeaeaeiaeo aca?iia?aeiioaiue
a oa?oiiaaiiyo, ui i?ecaiaeeoue aei iiyae eaneaaeiiai yoaeoo a
oi?ioaaii? ?ineeiiino?, iaeyne?aa?oei i?iyaii yeiai ? noeoean?ei?
noaae??.

10. ?ineeii?noue a?aeaae?a oaoo ? c?o/iei ia’?eoii aeey a?aei?aoethaaiiy
oa ia?aa??ee iaoiaee/ieo ieoaiue aea/aiiy neiaio?iiii? ?ineeiiino?.
?ic?iaeai? a ?? oiiaao iaoiaee (aiae?ca aiiiaai?oaoo, i?aeaeaeiaeo
aca?iia?aeiioaiue oa ?i.), iiaeia canoiniaoaaoe aeey aea/aiiy
?ineeiiino? ?ioiai aaiacena.

11. Aeyaeai? a oiae? aeine?aeaeaiue caeiiii??iino? ?icaeoeo ?ineeiiiai
iie?eao ia a?aeaaeao aoa?eueieo oaoo ? iaa?iiaaeaeaia aaca aeaieo ?
iniiaa aeey iaoeiai iaa?oioiaaiiai oi?ioaaiiy noaeeo ooo/ieo
o?oioeaiic?a ? eiio?ieth ?o noeoean?eieo ci?i.

Iioae?eiaai? ca oaiith aeena?oaoe?? i?aoe?:

1. Iiao A. I., AEoeia N. I. I oi?ie?iaaiee niiauanoa aunoeo ?anoaiee ia
ioaaeao oaieueiuo oaoo // Eio?iaeoeoeey e aeeeeiaoecaoeey ?anoaiee.-
1995.- Aui. 22.- N. 89-91 .

2. AEoeia N. I. Canoinoaaiiy eiao?oe??io?a iiae?aiino? i?e aea/aii?
?ineeiiino? a?aeaae?a aoa?eueieo oaoo // Oe?. aioai. aeo?i. — 1998. — ?
5. — N. 542-545.

3. AEoeia N. I. Noeoeanneiiiue noaoon aeaeia a ?anoeoaeueiuo niiauanoaao
ioaaeia e aai eniieueciaaiea a oeaeyo oeoieiaeeeaoeee // I?iiuoeaiiay
aioaieea: ninoiyiea e ia?niaeoeau ?acaeoey. Iaoa?eaeu o?aoueae
iaaeaeoia?iaeiie iao/iie eiioa?aioeee (Aeiiaoee, 3-5 naioya?y 1998 a.).
— Aeiiaoee: Ioeueoei?ann,1998. — N. 36-40.

4. AEoeia N. I. Eaneaaeiee aoaeo ia?aeiii? noeoean?? ia a?aeaaeao
aoa?eueieo oaoo Aeiiaano // Oe?. aioai. aeo?i. — 1999. — ? 1. — N.
5-10.

5. Iiao A. I., AEoeia N. I. Eciaiaiea i?iaeoeoeaiinoe ?anoeoaeueiuo
niiauanoa ii iniiaiui a?aaeeaioai n?aaeu ia ioaaeao oaieueiuo oaoo
Aeiiaanna // I?iiuoeaiiay aioaieea: ninoiyiea e ia?niaeoeau ?acaeoey.
Oacenu aeieeaaeia iaaeaeoia?iaeiie iao/iie eiioa?aioeee (E?eaie ?ia, iae
1993 a.). — Aeiiaoee, 1993. — N. 182.

6. AEoeia N. I. I iaaeaeaeiaie nii?yaeaiiinoe ia oa??eeiieeao Aeiiaoeea
// Aeooaeueiua aii?inu aioaieee e yeieiaee. Iaoa?eaeu eiioa?aioeee
iieiaeuo o/?iuo ( Oa?ueeia, 1996 a.). — Oa?ueeia, 1996. — N. 49.

7. AEoeia N.I. Aei aeai?o iaoiaeeee aea/aiiy aeeiai?ee ?ineeiiino? a
oiiaao oa?eeii?a // I?iaeaie aioai?ee ? i?eieia?? ia ii?ic? o?aoueiai
oeny/ie?ooy. Iaoa??aee X c’?caeo oe?a?inueeiai aioai?/iiai oiaa?enoaa
(Iieoaaa, 22-23 o?aaiy 1997 ?.). — Ee?a, 1997. — N.197-198.

8. AEoeia N. I. Aieea i?noea?icoaooaaiiy ia i?ioean oi?ioaaiiy
?ineeiiino? a?aeaae?a // Aeooaeuei? ieoaiiy aioai?ee oa aeieia??.
Iaoa??aee eiioa?aioe?? iieiaeeo a/aieo (Oa?nii-Eaco?ia, 7-11 aa?aniy
1998 ?.) . — Oa?nii, 1998. — N. 118-119.

AEoeia N. I. Aio?iiiaaiiay noeoeanney ?anoeoaeueiinoe ioaaeia oaieueiuo
oaoo Aeiiaanna. — ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/?iie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.16 — yeieiaey. — Aeiai?iiao?ianeee
ainoaea?noaaiiue oieaa?neoao, Aeiai?iiao?iane, 1999.

A aeenna?oaoeee ?anniao?eaaaony i?ioeann oi?ie?iaaiey
?anoeoaeueiinoe eiie/aneeo ioaaeia oaieueiuo oaoo Aeiiaanna. Iaoiaeii
eiiiiceoeeiiiie i?aeeiaoeee, iiaeeoeoee?iaaiiui aeey oneiaee ioaaeia,
?aeiino?oe?iaaia iineaaeiaaoaeueiinoue i?ioaeaiey noeoeannee. A?
?aciinoi?iiiaa eco/aiea ia iniiaa eiee/anoaaiiuo iiaeoiaeia iicaieeei
auaeaeeoue e eco/eoue oei?enoe/aneea e oeaiioe/aneea iniaaiiinoe
noeoeanneiiiuo noaaeee, aeeiaieeo i?ino?ainoaaiiiai ?ani?aaeaeaiey
aeaeia e eo iauaaeeiaiey a yeieiae/anee noiaeiua a?oiiu, eciaiaiey
iaaeaeaeiauo acaeiiioiioaiee. Iieo/aiiue oeaiio?iiieeei i?aaenoaaeyao
niaie yiie?eei-noaoenoe/aneoth iiaeaeue noeoeannee; ia iai ii eciaiaieth
oei?enoe/aneiai ninoaaa e oeoioeaiioe/aneie ?iee aeaeia auaeaeythony o?e
noaaeee noeoeannee. I?iaiaeece?iaaia nenoaiaoe/aneay e
yeiaeiii?oieiae/aneay no?oeoo?a oei?u ii noaaeeyi noeoeannee n
eniieueciaaieai aaoiiaoece?iaaiiie aacu aeaiiuo ii oei?a ?aaeiia
(Ao?aea, Inoaiei, 1993). Onoaiiaeaii, /oi ?anoeoaeueiinoue ioaaeia
?acaeaaaony a iai?aaeaiee oi?ie?iaaiey no?oeoo?, oa?aeoa?iuo aeey
ciiaeueiie ?anoeoaeueiinoe i?e aio?iiiaaiiii aeeyiee ia oiiiaii aeey
?aaeiia o?iaia. ?ani?aaeaeaiea aeaeia ii ieiuaaee oeoioeaiicia ioaaeia
eiaao eiioaaeiciue oa?aeoa? i?e ianooaaa ?anniio?aiey, niecia?eiii n
iyoienoinoueth ?ani?aaeaeaiey eiio?iee?othueo oaeoi?ia n?aaeu. Ca/anooth
yoi niaiaaeaao n ?acia?aie o/anoeia, iaeiioeiiuo ii oa?aeoa?o ?aeueaoa.
Aioo?e aea yoeo o/anoeia oa?aeoa? ?ani?aaeaeaiey ii?aaeaeyaony
noaiaiueth aaeaioaoeee aeaea e eo oneiaeyi, oa?aeoa?ii ?aciiiaeaiey e
a?aiaiiie oacie nouanoaiaaiey aeaiiiai aeaea. I?e i?eaeeaeaiee oneiaee
ianoiiaeoaiey e iioeiaeueiui aeey aeaea ?ani?aaeaeaiea no?aieony e
neo/aeiiio eee ?aaoey?iiio. Aiiiaaieoao niiauanoa iiauoaaony e eiioeo
noeoeannee. I?aaeeiaeaia iaoiaeeea oeoieiaeeeaoeee aeey ii?aaeaeaiey
ianoiyuaai ninoiyiey, noeoeanneiiiuo oaiaeaioeee eiie?aoiuo niiauanoa e
aeeythueo ia ieo oaeoi?ia ia iniiaa eniieueciaaiey noeoeanneiiiuo
noaoonia aeaeia niiauanoaa. ?acoeueoaou eeanneoeeaoeee e aeenia?neiiiiai
aiaeeca naeaeaoaeuenoaoao, /oi oeaiio?iiieeei ?anoeoaeueiinoe ioaaeia
oaieueiuo oaoo i?aaenoaaeyao aaeeiue noeoeanneiiiue ?yae, ?anoeoaeueiua
niiauanoaa naycaiu iaaeaeo niaie iinoaiaiiuie ia?aoiaeaie. Onoaiiaeaiu
aeaa iiaua annioeeaoeee: Persicario — Obernetum behenii e Poo compressae
— Hieracietum virosae, naeaeaoaeuenoaothuea i naiaia?acee
?anoeoaeueiinoe ioaaeia. Auaeaeaiu yeieiae/anee noiaeiua a?oiiu aeaeia.
Yoa aeeooa?aioeeaoeey ia iniiaa aiaeeca ei??aeyoeeiiiuo iao?eoe
io?aaeaao nouanoaothuaa ?ani?aaeaeaiea aeaeia, ai iiiaii iaoneiaeaiiia
aeeyieai ae?oaeo aeaeia. I?eiaiaiea /anoiuo ei??aeyoeee auyaeyao
noiaenoai yeieiaee aeaeia i?e ono?aiaiiii aeeyiee iaaeaeaeiauo
acaeiiioiioaiee. I?e noeoeanneiiiuo eciaiaieyo ?anoeoaeueiinoe iaiyaony
e noiaenoai aeaeia ii yeieiaee. Ia ?aciuo noaaeeyo iie aoiaeyo a a?oiiu
n ?aciuie aeaeaie. Anee aia/aea iie ia?acotho ?aciua eeanoa?u n aeaeaie
i?aaeuaeouae noaaeee, oi iiyaeaiea niaiythuae eo a?oiiu aeaeia
ioiineoaeueii naeeaeaao inoaaoeany aeaeu i?aaeoanoaothueo noaaeee.
I?aaeeiaeai iaoiae eco/aiey acaeiiioiioaiee aeaeia a iiiaiaeaeiauo
niiauanoaao, eioi?ue iicaieee iieo/eoue eiee/anoaaiioth ioeaieo
ioiineoaeueiie eiieo?aioiniiniaiinoe aeaeia.I?e yoii ii?aaeaeyaony eae
aeeyiea aeaea ia inoaeueiua aeaeu a niiauanoaa, oae e aai
/oanoaeoaeueiinoue e aicaeaenoaeth, ieacuaaaiiio ia iaai ni noi?iiu
ae?oaeo aeaeia. Yoe aeaa iieacaoaey ii?aaeaeaiu aeey iaeaieaa cia/eiuo
aeaeia ia anao noaaeeyo noeoeannee. I?e aiaae?aiee aeaeu iaeaaeatho
aunieie noaiaiueth aeeyiey ia ae?oaea aeaeu i?e aunieie
/oanoaeoaeueiinoe e ia?aoiiio aeeyieth. Yoi iaoneiaeeaaaony, aa?iyoii,
aieueoei cia/aieai eiieo?aioeee a noaaeee i?i?anoaiey e anoiaeia. Caoai
/oanoaeoaeueiinoue nieaeaaony, a iinea i?ioiaeaeaiey oeoioeaiioe/aneiai
iioeioia ia/eiaao iaaeaoue e ieacuaaaiia aeeyiea. Aiineaaenoaee iiaeao
niiaa oiaiueoeoueny /oanoaeoaeueiinoue aeaea, eiaaea ii auoaniyaony a
ia?ooaaiua o/anoee oeaiicia. Iai?yae?iiinoue iaaeaeaeiauo
acaeiiioiioaiee aic?anoaao a i?ioeanna noeoeannee. Ia ia?aie e aoi?ie
noaaeee iie iaoneiaeeaatho 25-35 % aa?uee?iaaiey ?anoeoaeueiinoe.
Ii?aaeaeythuoth ?ieue a no?oeoo?a niiauanoa iaaeaeaeiaua acaeiiioiioaiey
ia/eiatho ea?aoue oieueei ia o?aoueae noaaeee noeoeannee (a n?aaeiai
Ksen > 0.5). Oaeei ia?acii, noeoeanney ?anoeoaeueiinoe ioaaeia oaieueiuo
oaoo eiio?iee?oaony aeeiaie/aneei eiiieaenii ec aaeioe/aneeo oaeoi?ia e
iaaeaeaeiauo acaeiiaeaenoaee a nouanoaothueo ?anoeoaeueiuo niiauanoaao,
/oi i?eaiaeeo e iiyaeaieth eaneaaeiiai yooaeoa a oi?ie?iaaiee
?anoeoaeueiinoe, ?aaeueiui i?iyaeaieai eioi?iai yaeythony noeoeanneiiiua
noaaeee. ?ac?aaioaiiua a oneiaeyo ioaaeia iaoiaeu aiaeeca aiiiaaieoaoa e
iaaeaeaeiauo acaeiiioiioaiee iiaoo i?eiaiyoueny e i?e eco/aiee
?anoeoaeueiinoe ae?oaiai aaiacena. Auyaeaiiua a oiaea enneaaeiaaiee
caeiiiia?iinoe ?acaeoey ?anoeoaeueiiai iie?iaa ia ioaaeao oaieueiuo oaoo
e iaeiieaiiay aaca aeaiiuo i?aaenoaaeytho iniiao aeey oi?ie?iaaiey
onoie/eauo eneonoaaiiuo oeoioeaiicia ia ioaaeao oaieueiuo oaoo Aeiiaanna
e oi?aaeaiey eo noeoeanneiiiuie eciaiaieyie.

Eeth/aaua neiaa: noeoeanney, i?aeeiaoeey, ?ani?aaeaeaiea, ei??aeyoeey,
eiieo?aioiniiniaiinoue, noeoeanneiiiue noaoon, oeoieiaeeeaoeey, ioaaeu
oaieueiuo oaoo Aeiiaanna.

AEoeia N. I. Aio?iiiaaiia noeoean?y ?ineeiiino? a?aeaae?a aoa?eueieo
oaoo Aeiiaano. — ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
ca niaoe?aeuei?noth 03.00.16 — aeieia?y. — Aei?i?iiao?ianueeee
aea?aeaaiee oi?aa?neoao, Aei?i?iiao?ianuee – 1999 .

A aeena?oaoe?? ?icaeyaea?oueny i?ioean oi?ioaaiiy ?ineeiiino? eii?/ieo
a?aeaae?a aoa?eueieo oaoo Aeiiaano. Iaoiaeii eiiiiceoe?eii? i?aeeiaoe??,
iiaeeo?eiaaiei aei oiia a?aeaae?a, ?aeiino?oeiaaia iine?aeiai?noue
noeoeann??. ?? ??ciia?/ia aea/aiiy ia iniia? e?euee?nieo i?aeoiae?a
aeicaieeei aeae?eeoe oa aea/eoe oei?enoe/i? oa oeaiioe/i? oa?aeoa?enoeee
noeoean?eieo noaae?e, aeeiai?eo i?inoi?iaiai ?iciiae?eo aeae?a, ?o
iacaaeiaiiy a aeieia?/ii iiae?ai? a?oie, i?aeaeaeia? aca?iia?aeiioaiiy.
Anoaiiaeaii, ui ?ineeii?noue a?aeaae?a ?acaeaa?oueny o iai?yi?
oi?ioaaiiy no?oeoo?, oa?aeoa?ieo aeey ciiaeueii? ?ineeiiino?. Aecia/aii
noeoean?eiee noaoon aeae?a, ca yeei aeyaeythoueny oaiaeaioe?? ?icaeoeo
oa?oiiaaiue. ?ic?iaeaii iaoiae aea/aiiy, ca yeei aecia/aia a?aeiinia
eiieo?aioicaeaoi?noue aeae?a o aaaaoiaeaeiaeo o?oioeaiicao. Noai?aia
aaca aeaieo aeey canoinoaaiiy io?eiaieo ?acoeueoao?a i?e ?aeoeueoeaaoe??
a?aeaae?a.

Eeth/ia? neiaa: noeoean?y, i?aeeiaoe?y, ?iciiae?e, ei?aeyoe?y,
eiieo?aioicaeaoi?noue, noeoean?eiee noaoon, o?oi?iaeeeaoe?y, a?aeaaee
aoa?eueieo oaoo Aeiiaano.

Zhukov S. P. Anthropogenous succession of vegetation on Dondass’s coal
dumps.- Manuscript

Thesis for conferringon a degree of a candidate of biological sciences,
specification 03.00.16 — ecology. — Dnepropetrovsk State University.-
Dnepropetrovsk, 1999.

The process of vegetation formation on conic dumps of Donbass’s coal
mines is considered in the thesis. The sequence of the succession course
has been reconstructed by a modified method of compositional ordination
for conditions of dumps. Its versatile study on the basis of
quantitative approaches enabled to define and study floristic and
cenotic peculiarities of successional stages, dynamics of species
spatial distribution, of their unification into ecologically similar
groups, of interspecific interrelations. It has been revealed that the
vegetation of dumps develops in the direction of structures formation,
characteristic of zonal vegetation. The succession species status, by
which tendencies of communities development are elucidated, has been
determined. The method of relative competitiveness definition in
multispecific communities has been suggested. The database has been
created for employing obtained results in dump restoration.

Key words: succession, ordination, distribution, correlation,
competitiveness, successional status, phytoindication, Dondass’s coal
dumps.

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