Вивчення поєднань різних генотипів свиней в умовах свинокомплексу з метою одержання високопродуктивних товарних гібридів: Автореф. дис… канд. с.-г.

OE?A?INUeEA AEAAeAI?ss AA?A?IEO IAOE

?INOEOOO NAEIA?NOAA OAAI

OAeE 638.4.082.43.

Ii?ic Ieaa A?eai?iae/

Aea/aiiy ii?aeiaiue ??cieo aaiioei?a naeiae a oiiaao naeiieiiieaeno c
iaoith iaea?aeaiiy aenieii?iaeoeoeaieo

oiaa?ieo a?a?eae?a.

06.02.01 – ?icaaaeaiiy oa naeaeoe?y oaa?ei

Aaoi?aoa?ao

aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

Iieoaaa, 1999

Aeena?oaoe??th ? ?oeiien.

?iaioa aeeiiaia o a?aeae?e? ?icaaaeaiiy oa aaiaoeee

?inoeoooo naeia?noaa Oe?a?inueei? aeaaeai?? aa?a?ieo iaoe.

Iaoeiaee ea??aiee –aeieoi? n?euenueeiainiiaea?nueeeo iaoe,
/eai-ei?aniiiaeaio OAAI, i?ioani? Ieeiea Aeaaeaeiae/ Aa?acianueeee,
?inoeooo naeia?noaa OAAI, caa. a?aeae?eii ?icaaaeaiiy oa aaiaoeee.

Io?oe?ei? iiiiaioe: -aeieoi? n?euenueeiainiiaea?nueeeo iaoe, i?ioani?
Na?a?e Aioiiiae/ Aieiaeiae/, E?einueeee aea?aeaaiee aa?a?iee
oi?aa?neoao, caa?aeoaa/ eaoaae?e oaa?eiieoeoaa;

-eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe, ?aai/aiei Ieeiea ?aaiiae/,
aieiaa i?aae?iiy (AAO “Iieoaaaieaina?a?n”, Oe?ieaiia’?aeiaiiy
I?i?noa?noaa AIE Oe?a?ie).

I?ia?aeia onoaiiaa -Oa?niinueeee aea?aeaaiee aa?a?iee oi?aa?neoao ?i.
I.Ae. Oeth?thie, eaoaae?a ?icaaaeaiiy ? aaiaoeee, I?i?noa?noai
aa?ii?iieneiaiai eiiieaeno Oe?a?ie, i. Oa?nii.

Caoeno a?aeaoaeaoueny 10 aa?aniy 1999 ?ieo i 10 aiaeei? ia can?aeaii?

niaoe?ae?ciaaii? a/aii? ?aaee E 44.351.01 i?e ?inoeooo? naeia?noaa OAAI
ca aae?anith: 314006, Iieoaaa, Oaaaenueea Iiaeea.

C aeena?oaoe??th iiaeia iciaeiieoeny o a?ae?ioaoe? ?inoeoooo naeia?noaa
OAAI, 314006, Iieoaaa, Oaaaenueea Iiaeea.

Aaoi?aoa?ao ?ic?neaiee “9 “ na?iiy 1999 ?.

A/aiee nae?aoa? niaoe?ae?ciaaii? a/aii? ?aaee,

eaiaeeaeao a?ieia?/ieo iaoe I.?.
I?aeoa?aaa

Caaaeueia oa?aeoa?enoeea ?iaioe

Aeooaeuei?noue oaie aeine?aeaeaiue. A naeia?noa? i?enei?aiiy
iaoeiai-oaoi?/iiai i?ia?ano, ye ? a ?ioeo aaeocyo, aecia/a?oueny oni?oii
a iaeano? naeaeoe?? ii oaeineiiaeaiith ?nioth/eo oa noai?aiith iiaeo
ii??ae, oei?a oa ?o e?in?a aeey iaea?aeaiiy a?a?eaeieo naeiae.

Oe?a?ia ia? aeinoaoi?e aaiaoe/iee iioaioe?ae ii??ae naeiae, yeeo iiaeia
aoaeoeaii aeei?enoiaoaaoe a nenoai? a?a?eaeecaoe?? aeey iaea?aeaiiy
iieiaeiyeo ia a?aeaiae?aeth. Iaeiae ? aei oaia??oiueiai /ano o aaaaoueio
ainiiaea?noaao, a oiio /ene? ? niaoe?ae?ciaaieo, iaei i?noea aacnenoaiia
aeei?enoaiiy aaiioei?a, ui iaaaoeaii a?aeaeeiny ia oi?ioaaii? noaae, oa
onoaaei iiaeeea?noue aoaeoeaii ai?iaaaeaeoaaoe o ae?iaieoeoai iia?
naeaeoe?ei? aeinyaiaiiy.

Inoaii?i /anii oe?ieiai ?iciianthaeaeaiiy iaaoea i?aeii?iaeia
a?a?eaeecaoe?y, yea ia?aaeaa/a? ?icaaaeaiiy naeiae ca niaoe?ae?ciaaieie
e?i?yie, oeiaie ? ii?iaeaie oa ?o e?inaie aeey cia/ii aoaeoeai?oiai a?ae
i?iieneiaiai no?auoaaiiy aeei?enoaiiy aaoa?iceno. Oiio aaaeeeaei
oaeoi?ii o ca?eueoaii? i?iyao aaoa?iceno i?e no?auoaaii?, a iniaeeai
a?a?eaeecaoe?? ? aea/aiiy aieeao ii??ae, e?i?e ia ?o eiia?iaoe?eio
caeaoi?noue.

Iaoeiai-iaa?oioiaaia aeei?enoaiiy aaiioiiaeo Oe?a?ie ?c canoinoaaiiyi
no/anieo aeinyaiaiue aaiaoeee (AeIE-oaoiieia?e) aeanoue iiaeeea?noue
aeaaaeaiiy aenieii?iaeoeoeaieo e?in?a naeiae aeey iaea?aeaiiy oiaa?ieo
a?a?eae?a. ? a iaeaooiueiio ae?iaieoeoai a?a?eae?a neeaaeaoeia iniiaio
/anoeio oiaa?ii? i?iaeoeoe??, oiaoi 11-12 iei. iii?nieo naeiae, c yeeo
6-8 iei. a?a?eae?a (?eaaeei A.I. 1991.).

Ioaea, i?aenoaaith aeey aeeiiaiiy aeine?aeaeaiue ii oai? aeena?oaoe??
aoea iaiao?aei?noue aea/aiiy aoaeoeaiino? /enoiii?iaeiiai ?icaaaeaiiy oa
i?aeii?iaeiiai no?auoaaiiy ia aac? oanoe aaiioei?a naeiae oe?a?inueei?
oa aiae?enueei? naeaeoe??. I?e oeueiio a?aoiaoaaeenue oiiae
“Eaeeoyinueeiai” naeiieiiieaeno, ye? ca nai?ie oaoiieia?/ieie
ia?aiao?aie a?aeaiae?ae? oa oo?eiaiiy oaa?ei i?aeoe/ii ia
a?ae??ciythoueny a?ae aaaaoueio i?iieneiaeo ainiiaea?noa Oe?a?ie, ui ?
aecia/a? aeooaeuei?noue aea?aii? oaie.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. ?iaioa aoea
neeaaeiaith /anoeiith iaoeiai-aeine?aeieo ?ia?o ?inoeoooo naeia?noaa
OAAI (iiia? aea?aeaaii? ?a?no?aoe?? 01.86.0.042.309).

Iaoa ? caaea/? aeine?aeaeaiue. Aecia/eoe aaciina?aaeiuei a oiiaao
?aaeainio-eiia?iaoo “Eaeeoyinueeee” iaea?eueo aoaeoeai? aa??aioe
ii?aeiaiue aeo?aeieo oi?i oe?a?inueeeo oa aiae?enueeeo aaiioei?a naeiae
aeey iaea?aeaiiy oiaa?ieo a?a?eae?a, a oaeiae ?ic?iaeoe i?iiiceoe?? ii
?o aeei?enoaiith a i?iieneiaiio naeia?noa?.

Caaaeaiiyi aeine?aeaeaiue aoei aea/eoe:

— a?aeoai?thaaeuei? yeino? naeiiiaoie aioo??ii?iaeiiai oeio aaeeei?
a?ei? ii?iaee (OAA-1) o ii?aeiaii? c eio?aie oe?a?inueei? oa
aiae?enueei? naeaeoe??;

— ??no oa ?icaeoie i?aeaeine?aeiiai iieiaeiyeo ia a?aeaiae?ae? aei
aeeai? iane 100 ? 120 ea;

— a?aeaiae?aaeuei?, caa?ei? oa i’yni? yeino? /enoiii?iaeiiai oa
iii?niiai iieiaeiyeo i?e a?aeaiae?ae? aei 100 ? 120 ea;

— ye?noue i’yna ? naea a caeaaeiino? a?ae aaiioeio i?aeaeine?aeieo
oaa?ei;

— iieaeoey?ii-aaiaoe/i? iniaeeaino? /enoiii?iaeieo ? a?a?eaeieo naeiae
i?e aeei?enoaii? AeIE-oaoiieia?e;

— aeae?eeoe iaea?eueo aeiiii?/ii aoaeoeai? ii?aeiaiiy aeey ae?iaieoeoaa
oiaa?ieo a?a?eae?a;

— ?ic?iaeoe i?iiiceoe?? ii ?aoe?iiaeueiiio aeei?enoaiith iaoa?einueeeo
? aaouee?anueeeo oi?i a nenoai? a?a?eaeecaoe?? naeiieiiieaeno.

Iaoeiaa iiaecia. Aia?oa aea/aii:

— aoaeoeai?noue aeei?enoaiiy aioo??ii?iaeiiai oeio o aaeee?e a?e?e
ii?iae? (OAA-1) c eio?aie oe?a?inueei? oa aiae?enueei? naeaeoe?? a
oiiaao i?iieneiai? oaoiieia??;

— i?iaaaeaii ii??aiyeuei? aeine?aeaeaiiy uiaei aeyaeaiiy
a?aeaiae?aaeueieo ? i’yni-naeueieo yeinoae /enoiii?iaeiiai ? iii?niiai
iieiaeiyeo i?e aeinyaiaii? ??cieo aaaiaeo eiiaeeoe?e (100 ? 120 ea);

— iieaeoey?ii-aaiaoe/i? iniaeeaino? /enoiii?iaeieo ? iii?nieo naeiae.

C o?aooaaiiyi iaea?aeaieo ?acoeueoao?a aeine?aeaeaiue, iiaeaeueoee
?icaeoie ae?noaei iieiaeaiiy i?i aeioe?euei?noue a?aeaiae?ae? naeiae
io?eiaieo aaiioei?a aei a?eueo aaaeeeo aaaiaeo eiiaeeoe?e(115…120 ea).

I?aeoe/ia oe?ii?noue iaea?aeaieo ?acoeueoao?a. A ?acoeueoao?
aeine?aeaeaiue aecia/ai? oa ?aeiiaiaeiaai? ae?iaieoeoao iaea?eueo
aoaeoeai? ii?aeiaiiy aaiioei?a naeiae oe?a?inueei? oa aiae?enueei?
naeaeoe?? a oiiaao “Eaeeoyinueeiai” naeiieiiieaeno, a oaeiae ?ioeo
ainiiaea?noa Oe?a?ie c i?iieneiaith oaoiieia??th. Iaea?aeai? aeai?
aeathoue iiaeeea?noue i?aeaeueoe aaeeeiie?aei?noue iaoie
aioo??ii?iaeiiai oeio o aaeee?e a?e?e ii?iae? i?e no?auoaaii? c eio?aie
ii??ae aeth?ie, eaiae?an oa aaeeei? a?ei? ii?iaee aiae?enueei?
naeaeoe??. Aeei?enoaoe aoaeo aaoa?iceno aeey iie?auaiiy
a?aeaiae?aaeueieo oa i’ynieo yeinoae oiaa?ieo a?a?eae?a.

?aae?caoe?y ?acoeueoao?a aeine?aeaeaiue. E?au? ii?aeiaiiy aaiioei?a
?aeiiaiaeiaai? oa ai?iaaaeaeai? ia naeiieiiieaen? “Eaeeoyinueeee”
Ee?anueei? iaeano?.

Iniaenoee aianie caeiaoaa/a. I?aeoe/ia /anoeia ?iaioe aeeiiaia iniaenoi
aaoi?ii, a ?ic?iaea oai?aoe/ii? /anoeie neeaaea? aeecueei 90%. Ooi/iaiiy
oai?aoe/ieo iieiaeaiue i?iaaaeaii iaoeiaei ea??aieeii aeieoi?ii
n?euenueeiainiiaea?nueeeo iaoe, /eai-ei?aniiiaeaioii OAAI i?ioani?ii
I.Ae.Aa?acianueeei.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iniiai? iieiaeaiiy aeena?oaoe??
aeiiia?aeaeenue ? iaaiai?thaaeenue ia i?aeia?iaei?e iaoeiai-ae?iaie/?e
eiioa?aioe?? “Eioaineoeeaoeey i?iecaiaenoaa i?iaeoeoeee aeeaioiiaiaenoaa
a ?anioaeeea Aaea?onue” (i. AEiae?ii, 1998), ?? i?aeia?iaei?e
eiioa?aioe?? “Aeei?enoaiiy no/anieo iieaeoey?ii-aaiaoe/ieo ?
a?ioaoiieia?/ieo ?ic?iaie o aaiaoeei-naeaeoe?eieo aeine?aeaeaiiyo”
(i.Iaeana,1998), iaoeiai-ae?iaie/?e ?aae? “Eaeeoyinueeiai”
naeiieiiieaeno (o aeiaoi? 1996), a/ai?e ?aae? ?inoeoooo naeia?noaa OAAI
(o ethoiio 1997). A iiaiiio ia’?i? ?acoeueoaoe aeena?oaoe?eieo
aeine?aeaeaiue aeiiia?aeaeenue ? iaaiai?thaaeenue ia ?icoe?aiiio
can?aeaii? a?aeae?eo ?icaaaeaiiy oa aaiaoeee ?inoeoooo naeia?noaa OAAI
(a ethoiio 1999) ? io?eiaee iiceoeaio ioe?ieo.

Ioae?eaoe??. Ca oaiith aeena?oaoe?? iioae?eiaaii 5 noaoae.

Ianya ? no?oeoo?a ?iaioe. Aeena?oaoe?y neeaaea?oueny ?c anooio, iniiaii?
/anoeie c o?ueio ?icae?e?a, aeniiae?a, i?iiiceoe?e ae?iaieoeoao oa
nieneo aeei?enoaieo aeaea?ae. ?iaioa aeeeaaeaia ia 153 noi??ieao
iaoeiiieniiai oaenoo ? ia? 30 oaaeeoeue oa 5 ?enoie?a. Nienie
aeei?enoaieo aeaea?ae iae?/o? 297 iaeiaioaaiue, c ieo 93 ia ?iicaiieo
iiaao.

Iaoa??ae, iaoiaee aeine?aeaeaiue.

Iaoeiai-ae?iaie/? aeine?aeaeaiiy i?iaaaeai? i?ioyaii 1995-1997 ?ie?a a
oiiaao “Eaeeoyinueeiai” naeiieiiieaeno oa a eaai?aoi??yo ?inoeoooo
naeia?noaa OAAI. Iaoa??aeii aeine?aeaeaiue aoee naei? oe?a?inueei? oa
aiae?enueei? naeaeoe?? ii??ae: aioo??ii?iaeiee oei o aaeee?e a?e?e
ii?iae? (OAA-1), oe?a?inueea oa iieoaanueea i’yni? ii?iaee (OI oa II),
aeth?ie (Ae), eaiae?an(E) oa aaeeea a?ea ii?iaea aiae?enueei? naeaeoe??
(AAA) ? aeaioe?i?ei? eio?e iaea?aeai? a?ae ii?aeiaiiy aaiioei?a
oe?a?inueei? oa aiae?enueei? naeaeoe??(OAA-1 o AAA).

Aeine?aeaeaiiy i?iaaaeai? ca a?aeiia?aeiith iaoiaee/iith noaiith
(oaae.1). A?oie iaoie aoee noi?iiaai? iaoiaeii aiaeia?a ca
iioiaeaeaiiyi, a?eii oa i?iaeoeoeai?noth. ?o ooo/ii in?iaiyee,
aeei?enoiaoth/e aeey oeueiai ii 3 eio?e o eiaei?e a?oi?. An? caae?yi? a
aeine?ae? oaa?eie aoee oeiiaeie aeey aeaieo aaiioei?a ? a?aeiia?aeaee
aeiiaai ia ieae/a ia?oiai eeano.

Oaaeeoey 1.

Noaia aeine?aeaeaiue

A?oie

oaa?eiI?ecia/ai-

iy a?oiIi?iaea ? ii?iaei?noueE?euee?noue,
aieIaoieEio??aIaoieEio??a?Eiio?ieueiaOAA-1OAA-113-153??Aeine?aeiaOAA-1OI
13-153???Aeine?aeiaOAA-1II13-153IVAeine?aeiaOAA-1Ae13-153VAeine?aeiaOAA-
1E13-153VIAeine?aeiaOAA-1AAA13-153VIIAeine?aeiaOAA-1OAA-1oAAA13-153?ai?i
aeoeoeai? yeino? i?aeaeine?aeieo naeiiiaoie ioe?ithaaee ca
aaaaoiie?aei?noth, aaeeeiie?aei?noth, iiei/i?noth, ianith ii?inyo i?e
a?aeeo/aii? ? caa?aaeaii?noth i?eieiaeo aei a?aeeo/aiiy (a 26 aeaiiiio
a?oe?).

A?ae?a?ai? aeey iiaeaeueoiai aea/aiiy oaa?eie (1 naeiea ? 1 eano?ao c
eiaeiiai ai?caea, oa?aeoa?i? ca ianith) aoee iinoaaeai? ia aei?iuoaaiiy,
a i?e aeinyaiaii? aeeai? iane 35-38 ea ia a?aeaiae?aeth.

??no oa ?icaeoie eiio?iethaaee ca ci?iith aeeai? iane iaoiaeii
?iaeea?aeoaeueiiai caaaeoaaiiy i?e ia?iaeaeaii?, a 26-, 106-, 222-,
256-aeaiiiio a?oe? ? i?e aeinyaiaii? aeeai? iane 100 ? 120ea.

Aeey aea/aiiy a?aeaiae?aaeueieo ? i’ynieo yeinoae a 106-aeaiiiio a?oe?
i?aenaeiee aoee iinoaaeai? ia a?aeaiae?aeth a oiiaao i?iieneiaiai
eiiieaeno aei aeeai? iane 100 ? 120 ea.

A?aeaiae?aey iieiaeiyeo naeiae i?iaiaeeeanue ca oaoiieia??th
?oae?enueei? o??ie “Aeae?-a-aeae?”. O i?aeaeine?aeieo oaa?ei aecia/aee
ne?aeoth/? iieacieee: a?e aeinyaiaiiy aeeai? iane 100 ? 120ea;
na?aaeiueiaeiaiaee i?e??no i?e a?aeaiae?ae? aei 100 ? 120 ea; aeo?aoe
ei?i?a o ei?iiaeo iaeeieoeyo ca oeae aea ia??iae.

I?ney cae?i/aiiy a?aeaiae?ae? caa?e oa iaaaeoaaiiy ooo aeine?aeieo
oaa?ei i?iaaaeaii a oiiaao naeiieiiieaeno (ii 5 aie?a c eiaeii? a?oie, a
oaeiae a eiaaaniiio oeaoo ii 5 aie?a c eiaeii? a?oie (aac iaaaeoaaiiy
ooo), ca?aeii ?c caaaeueiii?eeiyoeie iaoiaeeeaie.

Caa?ei? oa i’yni-naeuei? yeino? aecia/aee ca oaeei iieacieeaie: iana
ia?ii? ? ioieiaeaeaii? ooo?; ao?aoe iane ooo? i?e ioieiaeaeaii?; iana
aieiae, i?aeie, ia/?iee, eaaaiue ? na?oey; caa?eiee aeo?ae; aeiaaeeia
ooo?; oiaueia naea iaae 6…7 a?oaeieie o?aaoeyie; ieiua “i’yciaiai
a?/ea”; iana caaeiuei? o?aoeie i?aooo?; ii?oieia?/iee neeaae ooo?.

Anueiai ca ia??iae aeine?aeaeaiue aoei caaeoi 70 i?aenaeie?a, a 35
i?aooo i?aeaeaii iaaaeoaaiith.

I?e aea/aii? o?ceei-o?i?/ieo aeanoeainoae i’yna aecia/aee: aeoeaio
eeneioi?noue – ?I (/a?ac 48 aiaeei i?ney caaith); aieiaioo?eioth/o
caeaoi?noue; i?aei?noue i’yna; ?ioaineai?noue caaa?aeaiiy.

O?i?/iee neeaae i’yna ioe?ithaaee ca ai?noii ii/aoeiai?, a?a?ineii?/ii?
? caaaeueii? aieiae, aee?o, “ne?i?” ciee, i?ioa?io.

A na?aeiaeieaaeaiiio nae? aecia/aee: caaaeueio aieiao, oaiia?aoo?o
ieaaeaiiy (ii/aoeiao ? e?ioeaao) oa eiao?oe??io ?ao?aeoe??.

Ioe?ieo no?an/ooeeaino? naeiae cae?enithaaee ca aeiiiiiaith i?eeaaeo
o??ie “Boring Medical Engineering LTD” (Aiae?y), ca ?acoeueoaoaie
oanooaaiiy aaeioaiiaei iaoiaeii o iiaeeo?eaoe?? ?inoeoooo naeia?noaa
OAAI.

Iieaeoey?ii-aaiaoe/io oa?aeoa?enoeeo eio??a, naeiiiaoie ? anueiai
iieiaeiyeo ia a?aeaiae?ae? i?iaiaeeee oeyoii (aeei?enoaiiy
iieaeoey?ii-aaiaoe/ieo ia?ea??a) aea/aiiy aaiieo /anoio, ??ai?a
aiiiceaioiino? oa aaiaoe/ii? iiae?aiino? naeiae ??cieo aaiioei?a.

Oaa?eie an?o i?aeaeine?aeieo a?oi i?ioyaii aeine?aeaeaiue ciaoiaeeeenue
a aiaeia?/ieo oiiaao aiae?ae? oa aeiaeyaeo, ca?aeii c oaoiieia??th
?oae?enueei? o??ie “Aeae?-a-aeae?”, i?eeiyoith ia naeiieiiieaen?
“Eaeeoyinueeee”. Aiae?aey oaa?ei cae?enithaaeanue aea?/? ia aeiao,
iiaii?aoe?iiieie eiia?ei?iaie, aeaioiaeaieie ia Eaeeoyinueeiio
eiia?ei?iiaiio caaiae? o ni?aa?aeiioaii? c aiaeith 1:3.

?acoeueoaoe aeine?aeaeaiue ia?iaeai? iaoiaeaie aa??aoe?eii? noaoenoeee
ii I.A. Ieioeinueeiio., A.E. Ia?eo?’?a?e., A.I. Oaaei-Aa?acianueeiio.,
A.O. Eae?io. A?iiao?e/ia ia?iaea iniiaii? /anoeie iaoa??aeo
cae?enithaaeanue ia i?e?ieaeueeoeyoi?? IE-61 oa ia ia?niiaeueiiio
eiii’thoa?? ?AI AO – 486, ca aeiiiiiaith iaeaoo i?eeeaaeieo i?ia?ai.

?acoeueoaoe aeine?aeaeaiue.

I?iaeoeoeai? yeino? naeiiiaoie i?e /enoiii?iaeiiio ?icaaaeaii? oa
no?auoaaii?. Aiae?c aenia?eiaioaeueieo aeaieo i?iaiaeeee oeyoii
ii??aiyiiy eiaeiiai aa??aioo no?auoaaiiy c /enoiii?iaeiei ?icaaaeaiiyi
(? a?oia). Iaeaeua aaaaoiie?aei?noue iaea?aeaia i?e /enoiii?iaeiiio
?icaaaeaii? 11,2 ii?inyoe, ui ia 1,5; 1,0; 0,4 oa 0,7 ii?inyoe a?eueoa,
i?ae o naeiiiaoie ??, ???, ?V oa V a?oi i?e no?auoaaii? (oaae. 2).
Noaoenoe/ii a??ia?aeia ??cieoey o?eueee c ?? a?oiith (?0,05). Ca
aaeeeiie?aei?noth ? ianith iaei??? aieiae i?e a?aeeo/aii? o IV oa V
a?oiao aoa iaea?aeaiee aoaeo no?auoaaiiy, ii?inyoa a oeeo a?oiao
ia?iaeaeoaaeeny ia 14,3 oa 16,1 % aaae/? i?e (?0,001) ? iaee a?eueoo
aeeao iano a 26-aeaiiiio a?oe? ia 6,5 oa 4,5 % a?aeiia?aeii.

Oaaeeoey 2.

?ai?iaeoeoeai? yeino? i?aeaeine?aeieo naeiiiaoie (Mm)

A?oienAaaaoiAaeeeiIiei/A 26 aei?aCaa?aaeaie?aei?noue, aie?aie?aei?noue,
eai?noue, eaE?eue-e?noue ii-?inyo, aieIana ai?caea, eaIana 1 aieiae,
eai?noue, %?1311,2
0,461,120,0249,71,109,80,3057,00,975,810,1088,33,20??159,70,531,220,0344
,51,088,40,3249,21,245,900,1387,82,71???1510,20,431,200,0247,01,048,90,2
952,41,105,950,1287,32,47IV1510,80,391,280,0252,31,09,30,2757,51,286,190
,1287,01,97V1410,50,391,300,0350,90,858,90,2754,11,576,070,1185,42,98VI1
310,70,461,00,0142,81,508,50,3948,11,555,680,1381,34,52VII1310,80,591,11
0,0145,01,059,10,3351,71,295,730,1385,63,93A na-?aaeiue-

iio ii
II-V5910,30,221,250,0148,60,648,90,1553,30,746,030,0686,91,24Aoaeo
aaoa-?iceno, %

-8,03

+11,61

-2,21

-9,18

-6,49

+3,79

-1,58Caa?aaeai?noue ii?inyo i?e a?aeeo/aii? e?auith aoea o
eiio?ieuei?e a?oi? ? neeaea 88,3 %. ?io? a?oie iaee aeaui ieae/ee
??aaiue caa?aaeaiiino?, a i?i?iaeueiee iieaciee iaeaaeeoue oaa?eiai
ii?aeiaiiy (OAA-? o AAA) – 81,3 %. Iaeiae, ??cieoey na?aaei?o iieaciee?a
i?ae on?ia a?oiaie noaoenoe/ii ia a??ia?aeia.

A?eia? ci?ie aeeai? iane i?aeaeine?aeiiai iieiaeiyea. I?iaeoii
ia?aaeaa/aia a?aeaiae?aey naeiae a oiiaao i?iieneiai? oaoiieia?? i?e
aacaeaoeueiiio oo?eiaii?, eiioeao?aoiiio oeio aiae?ae? oa ?aaoeueiaaiiio
i?e?iee?iao?. A oeeo oiiaao iieiaeiye ??cieo aaiioei?a ??n oa ?icaeaaany
ia iaeiaeiai.

Aiae?c aeaieo (oaae. 3) na?ae/eoue, ui iieiaeiye i?e ii?aeiaii?
aioo??ii?iaeiiai oeio o aaeee?e a?e?e ii?iae? (OAA-?) c eio?aie ii??ae
aeth?ie oa eaiae?an iaa e?au? iieacieee ?inoo oa ?icaeoeo, i?ae
/enoiii?iaei? aiaeiae a on? a?eia? ia??iaee. Oae i?e ia?iaeaeaii? ?
a?aeeo/aii? a 26-aeaiiiio a?oe? e?au? iieacieee aeeai? iane iaee
oaa?eie ii?aeiaiiy (OAA-1 o Ae) oa (OAA-1 o E), ui oe?eeii caeiiii??ii,
ine?eueee aiie iaee iaioo na?aaeith e?euee?noue ii?inyo i?e a?aeeo/aii?
o oeeo a?oiao. A?aeiii, ui oeae iieaciee ia? iaaaoeaio ei?aeyoe?th c
na?aaeiueith ianith ii?inyo ia ai?caei.

Oaaeeoey 3.

AEeaa iana i?aeaeine?aeiiai iieiaeiyea a caeaaeiino? a?ae a?eo (Mm), ea

A?oieE?euee?noueA?e, aei?aoaa?eiaie?a, nI?e ia?iaeaeaii?O 26-aeaiiiio

a?oe?O 106 -aeaiiiio

a?oe?O 222-aeaiiiio

a?oe?O 256 -aeaiiiio
a?oe??351,120,026,011,1236,20,30101,60,91120,91,34??351,220,035,92
0,0836,00,20103,41,09119,71,78???351,200,026,04
0,1036,60,35102,91,16119,82,13IV351,28 0,02
6,230,0537,70,39107,31,04126,41,74V351,300,036,130,1038,30,42
108,61,41128,32,04VI351,0
0,015,831,1135,90,47106,01,38125,92,33VII351,110,025,920,1236,50,48104,1
1,26121,62,10I?ney 106-aeaiiiai a?eo, iieiaeiye aoa iinoaaeaiee ia
a?aeaiae?aeth ? oai?aoe/ii an? a?oie iaee iaeiaeia? iiaeeeaino? o ?ino?
oa ?icaeoeo i?e ?aeaioe/ieo oiiaao aiae?ae?, oo?eiaiiy oa aaaiaeo
iieacieeao.

Ca aeaieo oiia, a caeeth/iiio oaoiieia?/iiio ia??iae? c 106- aei
222-aeaiiiai a?eo, a?a?eaei? oaa?eie (?? – V a?oi) ia?aaa?ooaaee
/enoiii?iaeieo ?iaaniee?a o 222-aeaiiiio a?oe? ia 1,3…7,0 ea aai 1,8 …
6,89 % a?aeiia?aeii. Aea i?e a?aeaiae?ae? aei a?eueo aaaeeeo eiiaeeoe?e
(256 aeaiiiai a?eo) /enoiii?iaei? i?aenaeiee iinooiaeeny eeoa IV,V oa VI
a?oiai, a ii?aeiaiiy (OAA-? o OI) oa (OAA-? o II) ia?aaa?ooaaee ia 1,2
…1,1 ea, aai 0,99 … 0,91 %.

Ia aeeaey/enue ia iaeiaioo aeeao iano ii?inyo i?e ia?iaeaeaii? oa
a?aeeo/aii? oaa?ei VI a?oie, ii?aeiaiiy (OAA-1oAAA) iieacaei aeineoue
aenieee ??aaiue aia?a?? ?inoo o aeinyaiaii? aeeai? iane 100 oa 120 ea,
ui aeaco? ia iiaeeea?noue iaea?aeaiiy aoaeoo aaoa?iceno i?e
aioo??ii?iaei?e a?a?eaeecaoe?? ??cieo iiioeyoe?e aaeeei? a?ei? ii?iaee.

Oaiie a?aeiinieo i?e?ino?a, ye? ?ice?eaathoue iai?oaeai?noue oa
oaeaee?noue ?inoo, ii an?o a?oiao cieaeoaaeenue ca a?eii. Iaeneiaeueii?
aaee/eie aiie aeinyaaee a ia??iae aei?iuoaaiiy c 26- aei 106-aeaiiiai
a?eo ? noaiiaeee 502,99…524,96 %.

A?aeaiae?aaeuei? yeino? /enoiii?iaeieo ? a?a?eaeieo i?aenaeie?a.

Aenia?eiaioaeuei? aeai? na?ae/aoue i?i oa, ui naei?, iaea?aeai? i?e
aioo??ii?iaeieo oa i?aeii?iaeieo ii?aeiaiiyo, iaee ia?aaaao iaae
/enoiii?iaeieie ?iaanieeaie (? a?oia). I?aenaeiee VI oa VII a?oi i?e
a?aeaiae?ae? aei aeeai? iane 100 ea, a na?aaeiueiio ia 6,6…3,8 aeiy
?ai?oa aeinyaaee eiiaeeoe??, iaee ia 41…18 a a?eueo? na?aeiueiaeiaia?
i?e?inoe oa ia 0,3…0,13 ei?iiai? iaeeieoe? iaioa aeo?a/aee ia 1 ea
i?e?inoo a ii??aiyii? c eiio?ieueiith a?oiith (oaae. 4).

Oaaeeoey 4.

A?aeaiae?aaeuei? yeino? iieiaeiyea i?e aeinyaiaii? aeeai? iane 100 ? 120
ea (Im)

A?oieE?euee?noue

oaa?ei

aie?aO?eaae?noue a?aeaiae?ae?,

aei?aAaeiaee

i?e??no,eaA?e i?e cae?i/aii?

a?aeaiae?ae?, aei?aNa?aaeiueiaeiaiaee i?e??no,aAeo?aoe ei?io ia

iaeei ea i?e?inooAEeaa iana 100 ea.?31113,2 1,7465,2
0,62219,21,745625,364,500,04??30111,0
1,8967,40,92217,01,895817,964,370,06???29111,6
1,9966,20,84217,61,995717,294,440,06IY30104,5
1,7269,50,71210,51,725996,174,220,05Y31102,8
2,2270,21,05208,82,226059,064,200,06YI28106,6
2,2569,90,98212,62,256038,504,200,06YII29109,4 2,1267,50,83215,4
2,125807,254,370,05A na?aaeiueiio ii II-VII
a?.107,61,4668,40,67213,61,465905,824,300,04Aaoa?iceniee aoaeo
%-4,95+4,91-2,55+4,98-4,44AEeaa iana 120
ea.?21148,32,3284,51,01254,32,325646,774,990,06??20155,84,3083,51,55259,
04,0455610,475,070,10???18149,64,3283,11,68255,64,3255411,245,100,11IY20
137,03,3388,11,31243,03,335888,794,790,08Y20126,11,5889,81,61232,11,5859
910,744,720,09YI18138,24,2589,31,75244,24,2559511,674,740,10YII19148,24,
6684,41,50254,14,665639,994,870,13A na?aaeiueiio ii II-VII
a?.142,54,3986,41,24248,04,115768,344,880,07Aaoa?iceniee aoaeo
%-3,91+2,25-2,48+2,13-2,20Iaeneiaeuei? iieacieee aoee cao?eniaai? i?e
a?aeaiae?ae? i?aenaee?a IV oa V i?aeaeine?aeieo a?oi. *enoiii?iaeieo
?iaaniee?a aiie ia?aaa?ooaaee, a?aeiia?aeii: ca nei?inoeae?noth ia
8,7…10,4 aeiy; na?aaeiueiaeiaiaei i?e?inoii 37…43 a; cao?aoao ei?i?a ia
0,28 ei?iiai? iaeeieoe?, i?e noaoenoe/i?e a??ia?aeiino? (?0,001).

Aeaia oaiaeaioe?y a iniiaiiio caa??aaeany ? i?e a?aeaiae?ae? aei 120 ea
aeeai? iane , e??i ?? oa ??? a?oi. Oaa?eie oeeo a?oi iinooiaeeny
/enoiii?iaeiei ?iaanieeai a?eii aeinyaiaiiy 120ea ia 1,8…0,5 %,
na?aaeiueiaeiaiaei i?e?inoii – 1,4…1,8 % oa cao?aoao ei?i?a ia
iaeeieoeth i?iaeoeoe?? – 1:…2,2 %, aea a??ia?aeia ??cieoey ia
niinoa??aaeanue.

Iaee?au? iieacieee a?aeaiae?aaeueieo yeinoae iaea?aeai? o V a?oi?
ii?aeiaiiy aaiioei?a (OAA-1oE), ye i?e a?aeaiae?ae? aei 100 oae ? 120
ea.

Caa?ei? ? i’yni-naeuei? yeino? iieiaeiyeo. Aiae?c eiio?ieueiiai caaith ?
iaaaeoaaiiy o?o naeiae, caaeoeo i?e aeinyaiaii? ??cieo aaaiaeo
eiiaeeoe?e iieacaa, ui i?e aiaeia?/i?e aiae?ae? i?aeaeine?aeiee
iieiaeiye II–VII a?oi ca aeiaaeeiith i?aooo?, ieiu? “i’yciaiai a?/ea” oa
ian? caaeiuei? o?aoeie i?aooo? ia?aaa?ooaaa ?iaaniee?a ? a?oie (oaae.
5).

Oaaeeoey 5.

Caa?ei? yeino? naeiae ??cieo aaiioei?a i?e aeea?e ian? 100 ? 120 ea (I
i)

A?oieCaa?eiee

aeo?ae a %Aeiaaeeia i?aooo?, niOiaueia

oieeo, iiIeiua

“i’yciaiai

a?/ea” ni2Iana

ieinoo, eaAEeaa iana 100
ea?60,710,8693,471,2532,401,2527,840,439,950,11??61,071,0693,611,4231,44
1,2529,040,7610,610,18???61,211,0294,611,6731,381,4229,360,6510,750,18IY
62,650,7195,932,4729,491,5231,370,5910,780,17Y62,110,8096,242,2829,761,3
930,740,3810,820,22YI62,530,9296,442,5526,841,2232,810,1010,900,21YII60,
860,9694,332,2831,311,0729,2304910,020,14A na?aaeiueiio ii II-V
a?.61,730,3595,190,8230,040,5330,420,3210,640,08Aaoa?i-ceniee aoaeo,
%+1,68+1,84- 7,28+9,26+6,93AEeaa iana 120
ea?61,611,0696,701,6637,821,1429,730,5911,270,25??61,860,9796,941,7236,5
31,7230,080,4211,730,27???62,070,9797,181,3834,491,3530,480,7311,860,20I
Y63,140,9698,451,8132,421,6032,340,7812,690,21Y62,590,7898,032,2133,341,
6831,610,5812,130,23YI63,110,8699,562,7530,721,2033,421,1312,350,19YII62
,270,3597,382,7536,631,3030,740,4211,580,15A na?aae-iueiio ii II-V??
a?.62,510,3497,920,7734,020,6131,440,3212,050,09Aaoa?i-ceniee aoaeo,
%+1,46+1,26-10,05+5,75+6,92Caa?e i?aenaeie?a aeeaith ianith 100 ea
aeyaea, ui aioo??ii?iaei? ii?aeiaiiy (OAA-1oAAA) oa [OAA-1o(OAA-1oAAA)]
ia?aaa?ooaaee eiio?ieueia ii?aeiaiiy (OAA-1oOAA-1) ii caa?eiiio aeoiaeo
ia 0,15…1,82 %, a i?aeii?iaeia a?a?eaeecaoe?y II – V a?oi ia 0,36…1,94%.
Iaea?eueoith aeiaaeeiith i?aooo? oa?aeoa?ecoaaany iieiaeiye IV,V ? VI
i?aeaeine?aeieo a?oi. Ei?iooeie aoee ooo? /enoiii?iaeieo i?aenaeie?a
(93,47 1,25) ? a?a?eae?a a?ae ii?aeiaiiy iaoie OAA-1 c eio?aie
oe?a?inueei? i’ynii? ii?iaee (93,61 1,42).

I?i?iaeueio oiaueio naea iaae 6-7 a?oaeieie o?aaoeyie iaee i?aenaeiee VI
a?oie, iaea?aeai? i?e ii?aeiaii? naeiiiaoie OAA-1 c ie?aeieeaie aaeeei?
a?ei? ii?iaee aiae?enueei? naeaeoe?? – 26,841,22, ui ia 17,20% iaioa,
i?ae o iieiaeiyea eiio?ieueii? a?oie. Naieie a???aiyieie ca oeei
iieacieeii aoee a?a?eaee (OAA-1 o OI), (Nv=5,81%).

Oaaeeoey 6.

Ii?oieia?/iee neeaae ooo i?aeaeine?aeiiai iieiaeiyea i?e aeea?e ian?
100 ? 120 ea (Mm)

A?oieNa?aaeiy iana i?aooo, eaAi?no o i?aooo?,%Ni?aa?aeiioaiiy
i’yni:naeiI’ynaNaeaE?noieAEeaa iana 100
ea?30,450,5656,110,2832,580,6311,300,701:0,58??31,160,6556,490,9731,750,
8211,750,261:0,56???31,400,4456,880,4131,780,3911,330,111:0,56IY32,840,5
659,610,3629,220,8312,080,181:0,49Y32,180,7457,570,6730,350,6312,080,181
:0,53YI32,060,6360,060,5726,730,7713,210,351:0,44YII30,940,4856,320,3731
,260,5412,420,501:0,55A na?aaeiueiio ii II-V a?.

31,760,25

57,82035

30,180,42

11,980,18

1:0,52Aaoa?iceniee

aoaeo %+4,3+3,05-7,37+6,02-10,34AEeaa iana 120
ea?37,041,4453,650,8935,340,8911,010311:0,66??37,060,4754,820,7333.840,5
511,340,201:0,62???37,320,5655,170,6733,500,7911,330,331:0,61IV37,900,56
58,220,6031,310,6010,460,361:0,54V37,950,4556,770,4032,240,3310,990,361:
0,57VI37,900,5453,760,4629,060,6112,180,321:0,49VII37,530,3553,960,7234,
940,6211,100,161:0,65A na?aaeiueiio ii II-V??
a?.37,610,7756,280,4032,510,4311,230,141:0,58Aaoa?iceniee aoaeo,
%+1,54+4,90-8,01+2,00-12,12Ui ae noino?oueny ieiu? “i’yciaiai a?/ea”, oi
iaeneiaeueiee iieaciee iaei ii?aeiaiiy (OAA-1oAAA) – i?e noaoenoe/i?e
a??ia?aeiino? a 100 ea (?0,001) ? 120 ea (?0,01). I?aeii?iaei? a?a?eaee
IV oa V a?oi oaeiae iaee aeia?ee iieaciee a ii??aiyii? c eiio?ieueiith
a?oiith i?e noaoenoe/i?e a??ia?aeiino? a 100 ea (?0,01), a 120 ea
(?0,05).

Ca?aeii c ?acoeueoaoaie iaaaeoaaiiy ooo naeiae (oaae. 6), caaeoeo i?e
aeinyaiaii? aeeai? iane 100 ea, iaeaeuei aeoiaeii i’yna ? iaeieae/ei
naea – oa?aeoa?ecoaaeeny oaa?eie IV,VI oa V i?aeaeine?aeieo a?oi, ye?
a??ia?aeii ia?aaa?ooaaee ? a?oio ia 3,50…3,95 % (?0,001) oa 1,46 %
(?0,05).

Iaeiaioa cia/aiiy aeoiaeo i’yna i?e caai? a 120 ea niinoa??aaeenue o
oooao /enoiii?iaeieo oaa?ei ? a?oie (53,65 0,89). E?aueie aoee
ii?aeiaiiy naeiiiaoie c ie?aeieeaie aeth?ie oa aaeeea a?ea ii?iaea
aiae?enueei? naeaeoe?? (58,22…58,76 %), (?0,01…0,001)

O?ceei-o?i?/ia oa?aeoa?enoeea i’yciai? oa aee?iai? oeaieie. Aeoeaia
eeneioi?noue i’yciai? oeaieie aea/aieo ii?aeiaiue aaiioei?a aoea a
iaaeao 5,50…6,09 (oaae. 7), a??ia?aeio ??cieoeth c ? a?oiith iaee IV –
(?0,01) oa VI – (?0,001) i?aeaeine?aei? a?oie.

Oaaeeoey 7.

O?ceei-o?i?/i? aeanoeaino? i’yciai? oeaieie i?aeaeine?aeieo oaa?ei (Mm)

A?oien

?I, iaeeieoeue eeneioiino?Aieiaioo?e-ioth/a caeaoi?noue, %I?aei?noue,
nae?ioaineai?noue caaa?aeaiiy

( A o 1000)Ia?aaecaa?eia iana 100
ea?105,850,0451,910,2412,270,1252,40,27??105,770,0651,890,4812,350,1750,
20,57???105,830,0750,06-,1112,020,0552,70,52IV106,090,0648,80,2612,060,1
050,20,55V105,760,0347,930,1612,570,2047,80,39VI105,500,0547,580,3513,73
0,2152,00,42VII105,800,0452,660,1512,650,1448,90,67Ia?aaecaa?eia iana
120
ea?105,850,0453,220,9812,130,1452,70,71??106,060,1452,370,2412,640,1551,
90,72???105,760,0351,640,8111,540,1850,70,37IV105,910,0451,470,3812,340,
3456,00,39V105,830,0550,490,3712,200,1350,80,39VI105,650,0750,890,6312,8
00,2950,00,37VII105,960,0550,130,1912,870,2649,10,48Ca ?ioaineai?noth
caaa?aeaiiy oa i?aei?noth i’yna e?au? iieacieee iaee /enoiii?iaei?
i?aenaeiee i?e ??ci?e noaoenoe/i?e a??ia?aeiino?.

An? ii?aeiaiiy oa?aeoa?ecoaaeeny aeaui cieaeaiith aieiaioo?eioth/ith
caeaoi?noth, yea ciaoiaeeeanue a iaaeao 47,58… 52,22%, i?e/eiith /iai
iiaea aooe i?iieneiaa oaoiieia?y naeiieiieaeno.

I?e aea/aii? o?ceei-o?i?/ieo aeanoeainoae naea nooo?ai? ??cieoe? i?ae
a?oiaie ca eiaeiei /eneii oa eiao?oe??ioii ?ao?aeoe?? ia anoaiiaeaii.
I?e i?aeaeuaii? aeeai? iane oaa?ei a?ae 100 aei 120 ea a?aei?/a?oueny
cieaeaiiy caaaeueii? aieiaino? naea ia 0,14…1,41%.

I?ia?aeueiee neeaae e?noie i?aeaeine?aeieo oaa?ei. O?i?/iee neeaae
noaaiiaeo e?noie naeiae ??cieo aaiioei?a noaoenoe/ii a??ia?aeii?
??cieoe? ia iaa, ine?eueee oiiae aiae?ae? oa oo?eiaiiy aoee ?aeaioe/i?.

A na?aaeiueiio, o e?noeia?e oeaiei? i?noeeiny 33,9 % ciee. Iaeneiaeueiee
?? a?aenioie cao?eniaaiee o ii?aeiaii? aaiioei?a oe?a?inueei? oa
aiae?enueei? naeaeoe?? [OAA-1o(OAA-1oAAA)], ye? ia?aaa?ooaaee
eiio?ieueio a?oio ia 2,3 % ? i?i?iaeueiee iieaciee iaea ??? a?oia, aea
aaouee?anueeith oi?iith aoee eio?e iieoaanueei? i’ynii? ii?iaee.

I?e aeine?aeaeaii? i?ia?aeueiiai neeaaeo e?noie ca ai?noii ? oa Na a
cie? iaei ii?aeiaiiy (OAA-1oII) ia 0,84 % oa 0,13% a?aeiia?aeii iaioa
eiio?ieueii? a?oie. A?aeiioaiiy oinoi?o aei eaeueoe?th a na?aaeiueiio ii
I-VII a?oiao neeaaea? 0,68, ui ciaoiaeeoueny a iaaeao iieacieea
o?eoinoaoa eaeueoe?th, yeee ? iniiaith e?noie.

Iaea?aeai? aeai? o?i?/iiai neeaaeo e?noie i?aeaeine?aeiiai iieiaeiyeo
aeacothoue ia caeaaei?noue aea/aieo iieaciee?a a?ae aaiioeio naeiae c
o?aooaaiiyi a?eo oaa?ei, aiae?ae? oa ?o oo?eiaiiy.

No?an/ooeea?noue /enoiii?iaeiiai ? a?ae?eaeiiai iieiaeiyeo. Oanooaaiiy
ii?inyo i?iaaaeaii aaeaoaiiaei iaoiaeii o iiaeeo?eaoe?? ?inoeoooo
naeia?noaa OAAI. Ca iniiaiei e?eoa???i – ?ea?aei?noth i’yc?a, o aeiaei?e
?c aea/aieo a?oi, ni?eyoeeaeo aei no?an?a oaa?ei ia aeyaeaii. Iaeiae, ca
oaeeie e?eoa??yie ioe?iee, ye /an i?eoiaeo a ii?iaeueiee o?c?ieia?/iee
noai oa o?aii?, /anoea noeeueieo aei no?an?a oaa?ei noaiiaeea,
a?aeiia?aeii: 7,69…30 % oa 16,67…40 %. Oae, iiceoeaio ?aaeoe?th
noeeueieo aei no?an?a ca /anii i?eoiaeo oaa?ei o ii?iaeueiee
o?c?ieia?/iee noai cao?eniaaii o 18,84 %, a ca o?aii?ii – 32,26 %.
Iaea?eueoee a?aenioie noeeueieo oaa?ei aei no?ano iaea?aeaiee o VI a?oi?
(42,85 % oa 66,67 % a?aeiia?aeii) ii?aeiaiiy (OAA-1oAAA). Iaeiae,
a??ia?aeii? ??cieoe? i?ae a?oiaie anoaiiaeaii ia aoei. Oe? oanoe iaeaea
a iaeiaeia?e i??? oa?aeoa?ecothoue ??aaiue no?an/ooeeaino? naeiae
aea/aieo aaiioei?a, oi/a ca o?aii?ii a?i aeyaeany aeuei ia 8,98…10 %.

Iieaeoey?ii-aaiaoe/i? aeine?aeaeaiiy naeiae ??cieo aaiioei?a. Iacia/ia
aeei?enoaiiy RAPO-PCR o iniiaieo aeae?a n?euenueeiainiiaea?nueeeo oaa?ei
iiaeia iiynieoe ne?aeoth/eie oaeoi?aie: ?yaeii iaiaaeaiue, iia’ycaieo
ianaiia?aae c a?aeoai?thaaeuei?noth ?acoeueoao?a, aiii?ii?noth ae?eyiie
AeIE, caeo/aieo a aiie?o?eaoe?th, a c ae?oaiai aieo aaeeeith e?euee?noth
eieon-niaoeeo?/ieo ia?ea??a. Iaeiae a aeayeeo aeiaaeeao canoinoaaiiy
iaoiaeo RAPP-PCR ii?aaaeo? naaa [4,5].

Aoee i?iaiae?ciaai? aiie?eiie AeIE naeiae aaeeei? a?ei? ii?iaee oa ??
a?a?eae?a c eio?aie ??cieo aaiioei?a. Ia iniia? i?iaaaeaieo eaai?aoi?ieo
aeine?aeaeaiue, ye? ia?aoiaai? ca aeiiiiiaith eiii’thoa?ii? i?ia?aie
“GELSTAR”, iaea?aeai? oae? eiao?oe??ioe aaiaoe/ii? iiae?aiino? (oaae.
8).

Aiae?coth/e iaea?aeai? aeai? , ca aeiiiiiaith eiii’thoa?ii? i?ia?aie
“TREES”, aea/ai? iaie ia’?eoe aeyaeee i?ae niaith iiae?ai?noue aai
a?aei?ii?noue, ui ? aaaeeeaei e?eoa???i aeey ?iciiae?eo ?o ia eeane.
Aieiaia iaoa eeanoa?iiai aiae?co – ciaoiaeaeaiiy iiae?aieo a?oi ia’?eo?a
o aea??oe? aeaieo. Iaea?aeaia aeaiae?ia?aia a?aeia?aaea? oa?aeoa?
o?eiaaiaoe/ieo ca’yce?a a?aeiieo c ?noi??? ?o noai?aiiy, ui ? a?aei?/a?
?yae aaoi??a [1,2,3] [?en1].

Oaaeeoey 8.

Eiao?oe??io aaiaoe/ii? iiae?aiino? ??cieo a?oi naeiae

Ii?aeiaiiyOAA-1OIIIAeEAAAOAA-?-OI0,627-II0,5540,475-Ae0,5070,4230,479-E0
,6110,6280,5530,393AAA0,4840,5120,4240,4520,462-?en.1.Aeaiae?ia?aia
aaiaoe/ieo aca?iia?aeiinei i?ae ??cieie aaiioeiaie naeiae, ?ic?aoiaaia
ia iniia? RAPD–PCR aiae?c.

Aeiiii?/ia aoaeoeai?noue a?aeaiae?ae? naeiae ??cieo aaiioei?a.
?ic?aooiie aeiiii?/ii? aoaeoeaiino? iieacaa nooo?ao ??cieoeth a
nia?aa?oino? oa ??ai? ?aioaaaeueiino? a?aeaiae?ae? a?a?eaeieo ?
/enoiii?iaeieo naeiae. Iaea?eueoeie na?aaeiueiaeiaiaeie i?e?inoaie ?
i?i?iaeueieie aeo?aoaie ei?i?a ia 1 ea i?e?inoo oa?aeoa?ecoaaeenue
a?a?eaei? oaa?eie IV,V oa VI a?oi. Nia?aa?o?noue iaeiiai oeaioia?a
aeeai? iane (o oe?iao 1995 ?ieo) o aeaieo a?oiao aeyaeeany ieae/ith,
ii??aiyii c eiio?ieueiith a?oiith ia 1,12, 1,12 oa 1,20 iei. e?a, aai
6,22, 6,22 oa 6,67 % a?aeiia?aeii.

Iaeei c iniiaieo aeiiii?/ieo iieaciee?a aoaeoeaiino? a?aeaiae?ae? ?
??aaiue ?aioaaaeueiino? ae?iaieoeoaa. A ia?o?e (eiio?ieuei?e) a?oi? oeae
iieaciee neeaa 3,9 %. O aeine?aeieo a?oiao a?i eieeaaany o iaaeao 5,3 –
11,3 %. Iaeaeua ?aioaaaeuei?noue cao?eniaaia a VI a?oi? (ii?aeiaiiy
OAA-1oAAA)-11,3%. Aeia?a naaa ca?aeiiaiaeoaaee ? i?aeii?iaei? a?a?eaee
IV oa V a?oi, aea ??aaiue ?aioaaaeueiino? neeaa 10,8 %, ui ia 6,9 %
aeua o ii??aiyii? c /enoiii?iaeieie ?iaanieeaie.

Na?aaei?e a?aenioie ?aioaaaeueiino? ii?iaeii-e?i?eieo oa i?aeii?iaeieo
ii?aeiaiue neeaa 8,68 %, ui ia?aaeuo? eiio?ieueio a?oio ia 4,78 %.

Aeniiaee

Iaea?aeai? ?acoeueoaoe a aeine?aeao aeathoue i?aenoaao c?iaeoe
ne?aeoth/? aeniiaee :

1. I?iya aoaeoo aaoa?iceno i?e aioo??ii?iaeiiio ?icaaaeaii? oa
i?aeii?iaeiiio no?auoaaii? naeiae aecia/a?oueny aaiaoe/iith
a?aenaeaeoe?iiiaai?noth ? eiia?iaoi?iith caeaoi?noth aaouee?anueeeo oa
iaoa?einueeeo oi?i (ii??ae, oei?a ? e?i?e), ye? aeei?enoiaooue.

2. A?aeoai?i? yeino? naeiiiaoie caeaaeaoue a?ae iaoiae?a ?icaaaeaiiy oa
ii?aeiaiiy aaiioei?a. Iaeaeu? iieacieee aaaaoiie?aeiino? aoee iaea?aeai?
i?e /enoiii?iaeiiio ?icaaaeaii? (11,2 ii?inyoe ia iii?in) ? aeaui ieae/?
?acoeueoaoe ca oe??th iciaeith (9,7…10,8) aoee i?e i?aeii?iaeiiio
no?auoaaii? c eio?aie ii??ae oe?a?inueeith oa iieoaanueeith i’ynieie,
aeth?ie ? eaiae?an.

3. Ii?aeiaiiy iaoie aioo??ii?iaeiiai oeio OAA-1 c eio?aie aeth?ie oa
eaiae?an ni?eyei ca?eueoaiith aaeeeiie?aeiino? ia 14,3…16,1 %,
iiei/iino? 2,4…5,2 %, iane 1-ai ii?inyoe i?e a?aeeo/aii? ia 4,5…6,5 %.
Ca iieacieeaie aaaaoiie?aeiino? ? caa?aaeaiino? ii?inyo ia?aaaaa aoea ia
aioe? oaa?ei aaeeei? a?ei? ii?iaee i?e /enoiii?iaeiiio ?icaaaeaii?
(3,6…6,2 % ? 1,3…2,9 % a?aeiia?aeii).

4. A?aeaiae?aey /enoiii?iaeieo ? a?a?eaeieo i?aenaeie?a an?o aea/aieo
ii?aeiaiue aei aeeai? iane 100 ea ni?eyea iaea?aeaiith aenieiai ??aiy
na?aaeiueiaeiaiaeo i?e?ino?a 570…605 a. Oey oaiaeaioe?y caa??aaeanue ?
i?e a?aeaiae?ae? aei a?eueo aaaeeeo aaaiaeo eiiaeeoe?e –120 ea. Iaeaeu?
na?aaeiueiaeiaia? i?e?inoe i?e aeaio aa??aioao a?aeaiae?ae? iaa
iieiaeiye, iaea?aeaiee a?ae ii?aeiaiiy naeiiiaoie OAA-1 c eio?aie ii??ae
aeth?ie, eaiae?an, aaeeea a?ea ii?iaea aiae?enueei? naeaeoe??.

5. I?i?iaeueieie aeo?aoaie ei?i?a ia 1 ea i?e?inoo aeeai? iane
oa?aeoa?ecoaaeenue i?aeii?iaei? ii?aeiaiiy (OAA-1oAe) oa (OAA-1oE) –
4,22 e.iae. Iniaeeai aeae?eyeinue ii?aeiaiiy aaiioei?a oe?a?inueei? oa
aiae?enueei? naeaeoe?? (OAA-1o AAA), aeei?enoaiiy yeiai aeicaiey?
caiuaaeaeoaaoe aei 30 ea eiioeei?i?a ia eiaei?e aieia? ca ia??iae
a?aeaiae?ae? aei 100 ea, ii??aiyii c eiio?ieueiith a?oiith oaa?ei.

6. I’yni?noue ooo i?e caaiyo a 100 ? 120 ea aoea ??ciith ? caeaaeaea
a?ae aaiioeio oaa?ei. E?aueie ca oeei iieacieeii aoee ii?aeiaiiy aaeeei?
a?ei? OAA-1 c ie?aeieeii aeth?ie, eaiae?an oa aaeeei? a?ei? ii?iaee
aiae?enueei? naeaeoe??. Oaa?eie oeeo ii?aeiaiue iaee aeiao? i?aooo? ia
2,4…2,9 ni i?e aioo??ii?iaeieo ii?aeiaiiyo oa 1,3…2,9 ni –
i?aeii?iaeieo, oiaueio naea ia 2,6…5,6 ii ? 4,5…7,1 ii oa a?eueoo ia
0,9…1 ea iano iei?iea, a?aeiia?aeii a ii??aiyii? c /enoiii?iaeieie
i?aenaeieaie aaeeei? a?ei? ii?iaee.

7. Aecia/aii a?eueo aenieo i’yni?noue ooo o i?aenaeie?a aioo??ii?iaeiiai
ii?aeiaiiy (OAA-1oAAA). Aeaui iinooiaeeny ?i ca oe??th iciaeith a?a?eaee
a?ae ii?aeiaiiy: iaoee OAA-1 c eio?aie ii??ae aeth?ie ? eaiae?an.
Aeei?enoaiiy ie?aeiee?a i’ynieo ii??ae ca?oa?aeii? naeaeoe?? (Ae, E,
AAA) ni?eyei ca?eueoaiith ai?noo i’yna a oooao ia 1,46…3,95 % oa
ciaioaiith naea a ieo.

8. sse?noue i’yna /enoiii?iaeieo ? i?aeii?iaeieo i?aenaeie?a i?e caai?
aeeaith ianith ye a 100 oae ? 120 ea a?aeiia?aeaei aeiiaai i’yna
caaeia?eueii? yeino?. Eaai?aoi?ieie aeine?aeaeaiiyie anoaiiaeaii, ui
e?aueie iieacieeaie yeino? i’yna oa?aeoa?ecoaaany iieiaeiye a?ae
ii?aeiaiue (OAA-1oOAA-1) oa [OAA-1o(OAA-1oAAA)].

9. Oaa?eie aea/aieo aaiioei?a oa?aeoa?ecoaaeenue aenieith no?ee?noth aei
no?an?a, i?i ui na?ae/eoue a?aenooi?noue ??a?aeiino? i’yc?a, ye
iniiaiiai iieacieea no?an/ooeeaino? i?e oanooaaii? aaeioaiiaei iaoiaeii.

10. Aeei?enoaiiy iieaeoey?ii-aaiaoe/ieo aeine?aeaeaiue, a aea-iiio
aeiaaeeo oaoi?ee RAPD, ni?ey? ioe?ioe? iaoa?einueei? oa aaouee?anueeeo
oi?i ia aaiiiio ??ai?, ui a iaai?e i??? iiaeia aeei?enoiaoaaoe aeey
iaa?oiooaaiiy e?aueo ii?aeiaiue a nenoai? a?a?eaeecaoe??.

11. Iaea?eueo aeioe?eueieie aa??aioaie iaea?aeaiiy a?a?eaeiiai
iieiaeiyeo aeey a?aeaiae?ae?, ia iniia? i?iaaaeaieo aeine?aeaeaiue a
oiiaao ?aaeainio-eiia?iaoo “Eaeeoyinueeee” ? ii?aeiaiiy naeiiiaoie OAA-1
c eio?aie ii??ae aeth?ie, eaiae?an, aaeeei? a?ei? aiae?enueei?
naeaeoe??, ui aeicaiey? nooo?ai ca?eueoeoe ae?iaieoeoai naeieie aeniei?
yeino? ? a ?ioeo ainiiaea?noaao Oe?a?ie c i?iieneiaith oaoiieia??th.

12. ?ic?aooiie aeiiii?/ii? aoaeoeaiino? a?aeaiae?ae? i?aenaeie?a aei
aeeai? iane 100 ea iieacaa i?aeaeuaiiy ?aioaaaeueiino? ? ciaioaiiy
nia?aa?oino? 1 oe i?e?inoo i?e i?aeii?iaei?e a?a?eaeecaoe?? ia 6,9 % ?
1120 e?a. oa aioo??ii?iaeiiio ?icaaaeaii? ii?aeiaiiy (OAA-1oAAA) – 7,4 %
? 1200 e?a. (o oe?iao 1995 ?ieo) a ii??aiyii? c /enoiii?iaeieie
aiaeiaaie aioo??ii?iaeiiai oeio OAA-1.

I?iiiceoe?? ae?iaieoeoao.

1. A?aoiaoth/e oa, ui naeiiiaoee aioo??ii?iaeiiai oeio aaeeei? a?ei?
ii?iaee (OAA-1) iathoue caaeia?euei? aaeaioaoe?ei? oa
a?aeoai?thaaeuei? yeino?, a oiaa?i? a?a?eaee oa?aeoa?ecothoueny aeia?eie
a?aeaiae?aaeueieie ? i’ynieie iieacieeaie i?iaeoeoeaiino?, aeioe?eueii
aeei?enoiaoaaoe ?o ia i?iieneiaeo eiiieaenao, eio?? iathoue i?oeio
ei?iiao aaco, aeey ca?eueoaiiy ae?iaieoeoaa i?nii? naeieie.

2. ?aeiiaiaeoaaoe aeey ai?iaaaeaeaiiy o ae?iaieoeoai a oiiaao
Eaeeoyinueeiai naeiieiiieaeno, a oaeiae ?ioeo ainiiaea?noa Oe?a?ie c
i?iieneiaith oaoiieia??th ne?aeoth/? aa??aioe ii?aeiaiiy aaiioei?a
aioo??ii?iaeiiai iaoa?einueeiai oeio aaeeei? a?ei? ii?iaee (OAA-1) c
eio?aie ii??ae aeth?ie, eaiae?an, aaeeea a?ea aiae?enueei? naeaeoe??.

3. A?aeaiae?aeth a?a?eaeiiai iiaie?a’y ?aeiiaiaeiaaieo ii?aeiaiue
aaiioei?a cae?i/oaaoe i?e aeinyaiai? oaa?eiaie aeeai? iane 115…120 ea,
ui aeicaieeoue ca?eueoeoe ae?iaieoeoai naeieie aac cieaeaiiy yeino?
i?iaeoeoe??.

Iniiai? iieiaeaiiy aeena?oaoe?? aeeeaaeai? a ne?aeoth/eo ?iaioao:

1.Ii?ic I.A. Aieea eio??a ??cieo aaiioei?a ia ?ai?iaeoeoeai? yeino?
iaoie OAA-1 // Iaoeiai-ae?iaie/ee atheeaoaiue
“Naeaeoe?y”.-E.-1998.-*enei iyoa.-n.224…225.

2.Ii?ic I.A. A?aeaiae?aaeuei? yeino? naeiae ??cieo aaiioei?a a oiiaao
i?iieneiaiai eiiieaeno // A?niee aa?a?ii? iaoee.-1998.-?4.-n.73…74.

3.Ii?ic I.A. I?iaeoeoeai?noue naeiiiaoie aioo??ii?iaeiiai oeio OAA-1 o
ii?aeiaii? c eio?aie ??cieo aaiioei?a // Oaa?eiieoeoai
Oe?a?ie.-1998.-?5.-n.18…19.

4.Ii?ic I.A., Oaiiaaeiaa I.A., ssoieoeeay A.I., Ii/a?iyaa E.O.,
Aaeaoeeee A.I., Aa?acianeee I.Ae. Eniieueciaaiea RAPD-PCR aeey
ii?aaeaeaiey oeeiaaiaoe/aneeo naycae /enoiii?iaeiuo e aea?eaeiuo naeiae
// Aeei?enoaiiy no/anieo iieaeoey?ii-aaiaoe/ieo ? a?ioaoiieia?/ieo
?ic?iaie o aaiaoeei-naeaeoe?eieo aeine?aeaeaiiyo. Ca??iee iaoa??ae?a ??
I?aeia?iaeii? eiioa?aioe??. – Iaeana. – E. – 1998.

5. Aa?acianeee I.Ae., Ii?ic I.A. Aea?eaeecaoeey ia aioo?eii?iaeiie e
iaaeii?iaeiie iniiaa // Naeiiaiaenoai. – 1999. — ?2. – n.11…15.

Ii?ic I.A. Aea/aiiy ??cieo ii?aeiaiue aaiioei?a naeiae a oiiaao
naeiieiiieaeno c iaoith iaea?aeaiiy aenieii?i-aeoeoeaieo oiaa?ieo
a?a?eae?a. — ?oeiien.

Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa
n?euenueeiainiiaea?nueeeo iaoe ca niaoe?aeueiinoth 06.02.01 —
?icaaaeaiiy oa naeaeoe?y oaa?ei.- ?inoeooo naeia?noaa OAAI. Iieoaaa,
1999.

Aeena?oaoe?eia ?iaioa i?enay/aia aea/aiith iaea?eueo aoaeoeaieo
aa??aio?a ii?aeiaiue aaiioei?a naeiae oe?a?inueei? oa aiae?enueei?
naeaeoe?? aeey iaea?aeaiiy oiaa?ieo a?a?eae?a ia aioo??ii?iaei?e ?
i?aeii?iaei?e iniia? a oiiaao “Eaeeoyineiai” naeiieiiieaeno Ee?anueei?
iaeano?. Anoaiiaeaii, ui i?e aeei?enoaii? aioo??ii?iaeiiai oeio OAA-1,
ye iaoa?einueei? oi?ie, a eio??a ii??ae aeth?ie, eaiae?an oa aaeeei?
a?ei? aiae?enueei? naeaeoe??, aeinyaathoueny iaeaeu? iieacieee
a?aeoai?thaaeueieo, a?aeaiae?aaeueieo ? i’ynieo yeinoae. A?aeaiae?aeth
oiaa?ieo a?a?eae?a, iaea?aeaieo ca oaeith noaiith, a oiiaao Oe?a?ie
aeioe?eueii cae?i/oaaoe i?e aeinyaiaii? oaa?eiaie aeeai? iane 115…120
ea.

Eeth/ia? neiaa: ii?iaea, ii?aeiaiiy aaiioei?a, i?iieneiaa oaoiieia?y,
a?a?eaee, a?aeaiae?aey, aioo??ii?iaeia ? i?aeii?iaeia a?a?eaeecaoe?y.

Ii?ic I.A. Eco/aiea ?aciuo ni/aoaiee aaiioeiia naeiae a oneiaeyo
naeiieiiieaena n oeaeueth iieo/aiey aunieii?iaeoeoeaiuo oiaa?iuo
aea?eaeia. – ?oeiienue.

Aeenna?oaoeey ia nieneaiea iao/iie noaiaie eaiaeeaeaoa
naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe 06.02.01 – ?acaaaeaiea e
naeaeoeey aeeaioiuo.- Einoeooo naeiiaiaenoaa OAAI. Iieoaaa, 1999.

Aeenna?oaoeeiiiay ?aaioa iinayuaia eco/aieth iaeaieaa yooaeoeaiuo
aa?eaioia ni/aoaiee aaiioeiia naeiae oe?aeineie e aiaeeeneie naeaeoeee
aeey iieo/aiey oiaa?iuo aea?eaeia ia aioo?eii?iaeiie e iaaeii?iaeiie
iniiaa a oneiaeyo “Eaeeoyineiai” naeiieiiieaena Eeaaneie iaeanoe.
Onoaiiaeaii, /oi i?e eniieueciaaiee aioo?eii?iaeiiai oeia OEA-1, eae
iaoa?eineie oi?iu, a o?yeia ii?iae aeth?ie, eaiae?an e e?oiiie aaeie
aiaeeeneie naeaeoeee, aeinoeaathony iaeaieaa aunieea iieacaoaee ii
aini?iecaiaeeoaeueiui, ioei?ii/iui e iyniui ea/anoaai. Ioei?i oiaa?iuo
aea?eaeia, iieo/aiiuo ii oaeie noaia a oneiaeyo Oe?aeiu, oeaeaniia?acii
ieai/eaaoue i?e aeinoeaeaiee naeiueyie aeeaie iannu 115…120 ea.

Eeth/aaua neiaa: ii?iaea, ni/aoaiea aaiioeiia, i?iiuoeaiiay oaoiieiaey,
aea?eaeu, ioei?i, aioo?eii?iaeiay e iaaeii?iaeiay aea?eaeecaoeey.

O.G.Moros. Studying the various combinations of pig genotypes in the
complex production farm conditions with the purpose of receiving highly
– productive commercial hybrids. – Manuscript.

Dissertation for the science degree of the Candidate of Agricultural
Sciences in speciality 06.02.01 – Animal breeding and selection. – The
UAAS Pig-Breeding Institute, Poltava, 1999.

The dissertation work deals with studying the most effective variants of
genotypes combination in pigs of Ukrainian and English selection for
receiving commercial hybrids on the interbreed and cross-breed basis.
The experiments were conducted at the “Kalityansky” complex production
farm in the Kiev Region. It was determined, that at using the interbreed
type UWL-1 as the mother-form, and boars of duroc, landrace and English
selection Large White breeds, the highest indices for the reproductive,
fattening and meaty qualities are gained. Under the conditions of
Ukraine, fattening of the commercial hybrids, received by the described
way, should be finished at the age of pigs gaining 115-120 kg living
weight.

Key words: breed, genotypes combination, industrial technology, hybrids,
fattening, intrabreed and cross-breed hybridization.

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