Синантропна флора верхньої течії басейну Дністра (Україна): Автореф. дис… канд. біол. наук / Ю.А. Скиба, НАН України. Центр. ботан. сад ім. М.М.Гриш

IAOe?IIAEUeIA AEAAeAI?ss IAOE OE?A?IE

Oeaio?aeueiee aioai?/iee naae ?iai? I.I. A?eoea

NEEAA TH??E AIAe??EIAE*

OAeE 581.9. 632.51 (477)

NEIAIO?IIIA OEI?A AA?OIUeI? OA*?? AANAEIO AeI?NO?A (OE?A?IA)

03.00.05 – aioai?ea

Aaoi?aoa?ao aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy
eaiaeeaeaoa a?ieia?/ieo iaoe

Ee?a — 1999

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia a Iaoe?iiaeueiiio iaaeaaia?/iiio oi?aa?neoao? ?i. I.I.
Ae?aaiiaiiaa, I?i?noa?noai ina?oe Oe?a?ie

Iaoeiaee ea??aiee: eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe, i?ioani?,

A?eoei — Aiaiaiei Ai?en Einoyioeiiae/

Iaoe?iiaeueiee iaaeaaia?/iee oi?aa?neoao ?i. I.I. Ae?aaiiaiiaa,
aeaeai i?e?iaeie/i-aaia?ao?/iiai oaeoeueoaoo

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani? *iiee

Aieiaeeie? ?aaiiae/, Iaoe?iiaeueiee
oi?aa?neoao

?i. Oa?ana Oaa/aiea, i?ioani? eaoaae?e

aioai?ee

eaiaeeaeao a?ieia?/ieo iaoe Iinye?i Na?a?e

Eaii?aeiae/, ?inoeoooo aioai?ee

?i. I.A. Oieiaeiiai IAI Oe?a?ie,
caa?aeoth/ee

a?aeae?eii nenoaiaoeee aeueo ?ineei

I?ia?aeia onoaiiaa: Euea?anueeee aea?aeaaiee oi?aa?neoao ?i. ?aaia

O?aiea, I?i?noa?noai ina?oe Oe?a?ie,
eaoaae?a

aioai?ee, i. Euea?a

Caoeno a?aeaoaeaoueny «_19__» _aa?aciy___ 1999 ?. i 10_ aiaeei? ia
can?aeaii? Niaoe?ae?ciaaii? a/aii? ?aaee E. 26.215.01 Oeaio?aeueiiai
aioai?/iiai naaeo ?i. I.I.A?eoea IAI Oe?a?ie ca aae?anith: 252014, i.
Ee?a, aoe. O?i??yc?anueea, 1.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? Oeaio?aeueiiai
aioai?/iiai naaeo ?i. I.I. A?eoea IAI Oe?a?ie ca aae?anith: 252014, i.
Ee?a, aoe. O?i??yc?anueea, 1.

Aaoi?aoa?ao ?ic?neaiee ________18 ethoiai 1999 ?.____

A/aiee nae?aoa?

Niaoe?ae?ciaaii? a/aii? ?aaee ________ Ioce/oe A.I.

Anooi

Aeooaeuei?noue oaie. Cia/aiiy ciaiue i?i oei?o iineeththoueny a aniaeo?
iieiaeaiue I?aeia?iaeii? eiiaaioe?? i?i caa?aaeaiiy a?i??ciiiai?oiino?
(i. ??i–aea–AEaiae?i, 1992) oa ?aciethoe?? I?aeia?iaeii? eiioa?aioe??
Planta Europa (i. ??? (O?aioe?y), 1995). Ci?ie a a?inoa?? i?ecaiaeyoue
aei ciaioaiiy e?eueeino? aeae?a ?ineei oa ao?aoe oe?iiiai aaiioiiaeo
oei?e. Oaeaee? oaiie ?icaeoeo noni?euenoaa ? iineeaiiy aieeao ethaeeie
ia i?e?iaeiee ?ineeiiee iie?ea ni?e/eiththoue neiaio?ii?caoe?th
aai?eaaiii? oei?e oa ?ineeiiino?. Oiio, iaeiei c aieiaieo iaoeiaeo
caaaeaiue ? aea/aiiy no/aniiai noaio oei?e oa eiai ci?i aiane?aeie
aio?ii?/iiai aieeao.

Ii/aoie anaa?/iiio aea/aiith neiaio?iiii? oei?e Oe?a?ie iieeaaeaii
i?aoeyie I.?. Eioiaa, A.A. I?ioiiiiiai? oa ?i. Aeioaia? i?iaaaeaii
?aa?iiaeuei? aeine?aeaeaiiy neiaio?iiii? oei?e Ee?anueeiai Iie?nny (N.E.
Iinye?i), I?aaeiy Oe?a?ie (?.?. Ao?aea, A.I. Oiooa?ue, N.A.Iao?ee, O.A.
Aaneeue?aa–Iaia?oeaeiaa), Caea?iaooy (A.A. I?ioeue). Ie aeine?aeaeoaaee
i?ai?/ii–cao?aeio /anoeio I?eea?iaonueeiai oei?enoe/iiai ie?oao.

Oei?o aeine?aeaeoaaii? oa?eoi??? ii/aee aea/aoe c O?O no. oai?nuee?,
iieuenuee? oa oe?a?inuee? aeine?aeieee: O. Aa?a?o, A. Atheeaeue, Aeae.
Eiai, A. Oo?/einueeee, N. Eoeue/einueeee, A. Iaaeianueeee, I.?.Eioia,
A.?. *iiee, TH.?. Oaeya-Niniiei, N.I. Noieei, A.O.Caeai/oe, A.I. Oea/ee
oa ?i. I?ioa niaoe?aeuei? aeine?aeaeaiiy neiaio?iiieo aeae?a ?ineei
oeueiai ?aa?iio ia i?iaiaeeeeny, aoee a?coaeueieie ? o?aaiaioa?ieie.

?aa?iiaeuei? aeine?aeaeaiiy neiaio?iiii? oei?e ni?eyoeiooue ooi/iaiith
?? aeaeiaiai neeaaeo, a anaa?/iee aiae?c – aeyaeaiith caeiiii??iinoae
?? oi?ioaaiiy, ui ? oe?iiei iaoa??aeii aeey ?ic?iaee i?iaiico ?icaeoeo
oei?e i?ae aieeaii aio?ii?/iiai /eiieea, ye ia eieaeueiiio, oae ? ia
ieaiaoa?iiio ??aiyo, ui ia no/aniiio aoai? ? iaoiiaeth? ?ic?iaeo oaie
aeena?oaoe?eii? ?iaioe: «Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio
Aei?no?a».

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie.
Aeena?oaoe?eia ?iaioa aeeiiaia ia eaoaae?? aioai?ee IIO ?i.
I.I.Ae?aaiiaiiaa o 1995-1998 ??. ? iia’ycaia c ieaiii
iaoeiai–aeine?aeii? ?iaioe eaoaae?e ca oaiith «Ci?no ina?oe: oi?ie,
iaoiaee ? caniae i?aeaioiaee a/eoae?a».

Iaoa ? caaaeaiiy aeine?aeaeaiiy. Iiaeaoe iiaiee eiiniaeo
neiaio?iiii? oei?e ?aa?iio, aecia/eoe no/aniee noai oei?e oa i?iaanoe
?? aiae?c. I?inoaaeeoe ?noi??th oi?ioaaiiy neiaio?iiii? oei?e ? aeyaeoe
iiaeeea? oaiaeaioe?? ?? o?ainoi?iaoe?? i?ae ae??th aa?ioe/ieo, a?ioe/ieo
oa ?noi?e/ieo oaeoi??a.

Ia?oi/a?aia? caaaeaiiy aoee oaeeie:

anoaiiaeoe aeaeiaee neeaae neiaio?iiii? oei?e aeine?aeaeoaaii?
oa?eoi???;

i?iaanoe nenoaiaoe/iee, aaia?ao?/iee, a?iaeieia?/iee aiae?c;

anoaiiaeoe /an, niiniae i?a?aoe?? oa nooi?iue iaoo?ae?caoe??
aaeaaioeaieo ?ineei, ui c?inoathoue a ?aeii? aeine?aeaeaiiy;

aeyaeoe caeiiii??iino? aeioiiieia?/ii? i?eo?i/aiino? neiaio?iiieo
?ineei aei iaaieo o?ainoi?iiaaieo aeioii?a;

neeanoe eiiniaeo neiaio?iiii? oei?e ia iniia? ?acoeueoao?a aeanieo
aeine?aeaeaiue, aa?aa?ieo iaoa??ae?a oa e?oa?aoo?ieo a?aeiiinoae;

?ic?iaeoe i?iaiic iiaeeeaeo ci?i ?ineeiiiai iie?eao i?ae aio?iiiaaiiei
aieeaii.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Aia?oa anoaiiaeaii aeaeiaee
neeaae neiaio?iiii? oei?e aeine?aeaeoaaii? oa?eoi??? oa neeaaeaii ??
e?eoe/iee eiiniaeo. Iiaeaii ?icai?iooo oa?aeoa?enoeeo eiaeiiai aeaeo.

Aeyaeai? iia? aeey oei?e cao?aeieo iaeanoae Oe?a?ie i?noeaciaoiaeaeaiiy
iecee aaeaaioeaieo aeae?a. Cie?aia, aia?oa a ?aeii? aeine?aeaeaiiy
ciaeaeaii ea?aioeii? aeaee Ambrosia artemisiifolia L., Solanum cornutum
Lam., Centaurea diffusa Lam.

Aia?oa i?iaaaeaii anaa?/iee aiae?c neiaio?iiii? oei?e ?aeiio
aeine?aeaeaiue oa aeyaeaii ?? no?oeoo?i? iniaeeaino?.

Aena?oeaii ?noi??th iiyae ? ?icnaeaiiy iecee aaeaaioeaieo ?ineei ia
oa?eoi??? Oe?a?ie oa neeaaeaii ea?oe ?o a?aae?a.

Aeyaeaii aeioiiieia?/io i?eo?i/ai?noue neiaio?iiieo ?ineei aei
o?ainoi?iiaaieo aeioii?a.

Neeaaeaii i?iaiic iiaeeeaeo ci?i oei?e i?ae aieeaii ainiiaea?nueei?
ae?yeueiino?.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. I?aeoe/ia cia/aiiy iathoue
iaoa??aee c aeaeiaiai neeaaeo neiaio?iiii? oei?e, ye? iiaeia aoaea
aeei?enoaoe i?e i?aeaioiaoe? ??cieo oei?enoe/ieo, oi?ieia?/ieo aai
ocaaaeueithth/eo i?aoeue, i?enay/aieo neiaio?iii?e oei?? Oe?a?ie aai
?a?iie.

Iaoa??aee aeena?oaoe?? ? iaeiei c aeaea?ae anoaiiaeaiiy no/anieo
oaiaeaioe?e ?icaeoeo oei?e ia oa?eoi??? Oe?a?ie i?ae aieeaii aio?ii?/ieo
oaeoi??a.

Aeyaeaii a?aeiinii no?ee? aei ??cieo oi?i ? o?eaaeino? aieeao
aio?ii?/ieo i?an–oaeoi??a aeaee, ye? iiaeooue aooe a?i?iaeeeaoi?aie
caaaeueieo ci?i aeiae?eey, i?eaeaoi? aeey aeei?enoaiiy i?e i?iaaaeaii?
aeieia?/ii? aenia?oece ? iii?oi?eiao caaaeueiiai noaio ?ineeiiiai
iie?eao.

Iaoa??aee aeena?oaoe?? aeei?enoiaothoueny a iaa/aeueiiio i?ioean?: i?ae
/an i?iaaaeaiiy eaai?aoi?ieo caiyoue, iieueiaeo i?aeoee, aeeiiaiiy
eo?niaeo ?ia?o.

Iniaenoee aianie caeiaoaa/a. Ia an?o aoaiao ?iaioa aeeiioaaeany
aeena?oaioii naiino?eii ?c canoinoaaiiyi a?aeiia?aeieo iaoiaeee.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. ?acoeueoaoe aeine?aeaeaiue
aeeeaaeaii ia iaoeiaeo eiioa?aioe?yo «Oai??y ? i?aeoeea aea/aiiy
i?e?iaeie/eo aeenoeeie?i o aeu?e oa na?aaei?e oeie?» (Oiaiue, 1997);
«I?aeia?iaei? aniaeoe aea/aiiy oa ioi?iie a?i??ciiiai?ooy Ea?iao»
(?ao?a, 1997); «Ieoaiiy a?i?iaeeeaoe?? oa aeieia??» (Caii??aeaey, 1998);
«I?iieneiaa aioai?ea» (Aeiiaoeuee, 1998); «Iaaaoeinuee? /eoaiiy» (Ee?a,
1998) oa iaoeiai-i?aeoe/ieo eiioa?aioe?yo aeeeaaea/?a IIO ?i. I.I.
Ae?aaiiaiiaa. E??i oiai, ?acoeueoaoe aeena?oaoe?eiiai aeine?aeaeaiiy
aeei?enoaii i?ae /an i?iaaaeaiiy caiyoue oa niaoeeo?n?a c? nooaeaioaie
i?e?iaeie/i–aaia?ao?/iiai oaeoeueoaoo IIO ?i. I.I.Ae?aaiiaiiaa.

Ioae?eaoe??. Iniiai? ?acoeueoaoe aeena?oaoe?? a?aeia?aaeai? o 8
ae?oeiaaieo iaoeiaeo i?aoeyo.

No?oeoo?a oa ianya ?iaioe. Aeena?oaoe?y aeeeaaeaia ia 201 noi??ieao
iaoeiiieniiai oaenoo, i?noeoue 6 oaaeeoeue, 33 ?enoiee. Neeaaea?oueny c?
anooio, 7 ?icae?e?a, aeniiae?a, nieneo e?oa?aoo?e. A?ae?ia?ao?y aeeth/a?
187 i?aoeue, cie?aia 43 – ?ioiiiai? aeaea?aea.

CI?NO ?IAIOE

?ICAe?E ?

O?ceei–aaia?ao?/ia oa?aeoa?enoeea

?aeiio aeine?aeaeaiiy

?aeiiii aeine?aeaeaiiy aoea i?aaiaa?aaeia /anoeia aa?oiuei? oa/??
aanaeio ?. Aei?noa?. Eiai ieiua noaiiaeoue aeecueei 1600 ei2.
I?ai?/ii–cao?aeia iaaea ?aa?iio ca?aa?oueny c ?oneii aa?o?a’y Aei?no?a,
i?aaeaiia – c ieaeiueith nioaith a??nueeiai e?niaiai iiyno, a no?aeia –
i?ioiaeeoue ii ?oneo ?. No?ee.

Ia’?eoii aeine?aeaeaiiy aoea neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio
Aei?no?a. Oei?o aeine?aeaeoaaee o 1994-1998 ??., i?ae /an
ia?o?ooii-oei?enoe/iiai ianoaaeaiiy oa?eoi???. A iniiao iieeaaeaii
ii??aiyeueiee ii?oieiai-aeieiai-aaia?ao?/iee iaoiae aeine?aeaeaiiy.
Aeey ii??aiyiiy aeei?enoaii aa?aa?i? iaoa??aee c aa?aa???a
Euea?anueeiai oi?aa?neoaoo ?i. ?.O?aiea (LW), ?inoeoooo aeieia?? Ea?iao
IAI Oe?a?ie (LWE) oa ?inoeoooo aioai?ee ?i. I.A.Oieiaeiiai IAI Oe?a?ie
(KW), Ae?iaiaeoeueeiai oa No?eenueeiai e?a?ciaa/eo ioca?a, a oaeiae
iiene aeae?a, iioae?eiaai? o 1866–1996 ??. Eaoeinuee? iacae oaenii?a
ooi/iaii ca N.E. *a?aiaiiaei.

Ia i?aenoaa? aiae?co o?ceei-aaia?ao?/ieo oiia ?aeiio aeine?aeaeaiiy
anoaiiaeaii, ui aiie ? aaoa?iaaiieie oa cia/ii ci?iaieie i?ae aieeaii
ethaeeie.

?ICAe?E 2

Ia?en oei?iaaiaco oa no/aniiai noaio

?ineeiiiai iie?eao

Ieoaiiyi oei?iaaiaco ia oa?eoi??? cao?aeieo iaeanoae Oe?a?ie caeiaeany
iecea aeine?aeiee?a (Uae?ia, ?eue?inueea [134, 43]; Nya?ye, Uae?ia [93];
Oaa?aaa [125]; N??aiei, Oo?ei [83]; Oaoa? [174], A?othoaiei oa ?i. [7];
Aaconueei, Ee?iaiia [8]). Anoaiiaeaii, ui a?ae aeaaiueiai aieioeaio aei
iaoiai /ano ci?ithaaany ee?iao, ui, a naith /a?ao, ni?e/eithaaei aei
ci?ie aeaeiaiai neeaaeo oei?e ? ?ineeiiino?. Aiane?aeie oeeo
aaoa?io?iiieo i?ioean?a o oei?? ?aa?iio aeyaeaii i?aaenoaaiee?a eiaeii?
c aiio.

A oe?eiio i?e?iaeiee ?ineeiiee iie?ea ?aeiio aeine?aeaeaiue aeineoue
iaiaeii??aeiee, ooo aeae?ey?oueny e?eueea oei?a i?e?iaeii? ?ineeiiino?,
a naia: oe?ieieenoyi? oa i?oai? e?ne, eoee ? aieioa, ui iathoue nai?
oa?aeoa?i? oei?enoe/i? iniaeeaino?. I?ioean oi?ioaaiiy neiaio?iiii?
oei?e o?nii iia’ycaiee c? ci?iaie i?e?iaeii? oei?e, ai cia/ia
e?euee?noue aaeaaioeaieo aeae?a aaeaioo?oueny aei o?ainoi?iiaaieo
aeioii?a.

A?aeiia?aeii aei oei?enoe/iiai ?aeiioaaiiy Oe?a?inueeeo Ea?iao, [120],
yei? ie aeio?eio?iiny, ?aeii aeine?aeaeaiiy ?icoaoiaaiee a iaaeao
No?aeiiea?iaonueei? i?aei?ia?ioe?? I?eea?iaonueeiai oei?enoe/iiai
?aeiio.

?ICAe?E 3

Ia?en ?noi??? aea/aiiy oei?e ?aeiio aeine?aeaeaiue

A ?noi??? aea/aiiy oei?e ?aeiio aeine?aeaeaiue caeaaeii a?ae ?noi?e/ieo
iiae?e, ie aeae?ey?ii o?e ia??iaee, a ye? aeine?aeieoeueea ?iaioa aoea
iaea?eueo aeoeaiith. O ia?oee ia??iae (O?O no.) oei?o aeine?aeaeoaaee
ia?aaaaeii aano?i-oai?nuee? aeine?aeieee. O ae?oaiio ia??iae? (ii/aoie
OO no.) aaeeeo ?iaioo a oeueiio iai?yieo i?iaiaeeee iieuenuee? a/ai?, ai
aei 1939 ?. oey oa?eoi??y aoea aaei?i?no?aoeaii i?aeii?yaeeiaaia
Iieueu?. Inoaii?e ia??iae ii/eia?oueny o i?neyai?ii? 40-ia? ?iee OO
no. ? o?eaa? aei oeueiai /ano.

O ?iaio? i?iaaaeaii aiae?c iaeiioe?ai?oeo eeaneo?eaoe?e neiaio?iiieo
aeae?a ?ineei, ?ic?iaeaieo A. Oaeeoiaii [176, 177], I.N.Eaieoaaei [46],
E. Aieoaii, A. ??anaeii [149], ss. Ei?ianai [156, 157], O. O?aaea?ii
[169], ?.A. Aei?iainoaenueeith [36, 37], A.N. ssa?iaa–Eieaeianueeith
[138], I.A. A’theiaith [24], A.A. *?/aaei [116], A.A.I?ioiiiiiaith [77,
79] oa ?ioeie iaoeiaoeyie. I?ioa ia aeaiee iiiaio iaia?
caaaeueiii?eeiyoi? eeaneo?eaoe??, yea a a?aeiia?aeaea iao? eiiieaenii?
ioe?iee oei?e.

Caaaeoth/e ia aenieee nooi?iue aio?iiiaai?caoe?? eaiaeoaooo, aeey
aiio?oii? o?aeoe?? ie i?iiiio?ii aeei?enoaoe eeaneo?eaoe?th
A.A.I?ioiiiiiai? [79], a iniiao yei? iieeaaeaii iiaeeea?noue oa nooi?iue
aaeaioaoe?? ?ineei aei o?ainoi?iiaaieo aeioii?a, a aeey aaeaaioeaii? –
nenoaio O. O?aaea?a [169], aeaui iiaeeo?eiaaio a?aeiia?aeii aei
caaaeaiue aeine?aeaeaiiy. Oiio ie i?iiiio?ii naith, aeaui iiaeeo?eiaaio
eeaneo?eaoe?th. I?e oeueiio, ii iiaeeeaino?, ie aeei?enoaee aaea
?nioth/? oa?i?ie.

Neiaio?iie

Aaeaaioe Aiio?oe

A. Ca /anii ?i?a?aoe??: A. Nooi?iue aaeaioaoe??

1) a?oaio?oe 1) aaaiio?oe

2) eaiio?oe 2) aai?aiio?oe

A. Ca niiniaaie ?i?a?aoe??: 3) iano?ee? aiio?oe

1) enaiio?oe

2) a?aac?io?oe

A.Ca nooiaiai iaoo?ae?caoe??:

1) aa??io?oe

2) aiaeio?oe

3) aoaia?io?oe

4) eieiiio?oe

?en. 3.1. Nenoaia eeaneo?eaoe?? neiaio?iiii? oei?e

?icae?e 4

Eiiniaeo neiaio?iiii? oei?e

Eiiniaeo neiaio?iiii? oei?e aeine?aeaeoaaiiai ?aeiio neeaaeaiee ca
?acoeueoaoaie aeanieo o?e??/ieo iieueiaeo aeine?aeaeaiue oa ia iniia?
ii?aoethaaiiy aa?aa?ieo eieaeoe?e Ae?iaiaeoeueeiai e?a?ciaa/iai iocath,
Euea?anueeiai oi?aa?neoaoo ?i. ?. O?aiea (LW), ?inoeoooo aeieia?? Ea?iao
IAI Oe?a?ie (LWE), ?inoeoooo aioai?ee ?i. I. A. Oieiaeiiai IAI Oe?a?ie
(KW) ? e?oa?aoo?ieo aeaiieo.

Aei «Eiiniaeoo…» aeeth/aii 365 aeae?a c 54 ?iaeei oa 213 ?iae?a,
aeyaeaieo iaie ia oa?eoi??? aeine?aeaeaiiy. Nthaee aeeth/aia ? cia/ia
e?euee?noue eoeueoo?ieo ?ineei, ye? aee/aa?thoue. Iiaeaii ?icai?iooo
oa?aeoa?enoeeo eiaeiiai aeaeo.

I?ae /an iieueiaeo aeine?aeaeaiue iaie aoee aeyaeai?
i?noeaciaoiaeaeaiiy oanoe aeae?a aaeaaioeaieo ?ineei, iiaeo aeey
cao?aeieo iaeanoae Oe?a?ie, cie?aia: Heracleum mantegazzianum S. et L.,
Lactuca tatarica (L.) C.A.Mey., Commelina communis L., Solanum cornutum
Lam., Centaurea diffusa Lam., Physalis ixocarpa Brot. ex Hornem.

E??i oiai, aeyaeaii iia? i?noeaciaoiaeaeaiiy oaeeo aaeaaioeaieo aeae?a,
ye Veronica arvensis L., Rudbeckia laciniata L., Ipomaea purpurea (L.)
Roth., Echinocystis lobata (Michx.) Torr. ex Gray, Tragopogon dubium
Scop., Verbena officinalis L., Bryonia dioica Jacq., Ambrosia
artemisiifolia L.oa ?i.

?ICAe?E 5

AIAE?C NEIAIO?IIII? OEI?E

Ca ?acoeueoaoaie aeine?aeaeaiue i?iaaaeaii aiae?c nenoaiaoe/ii?,
aaia?ao?/ii?, aeieia?/ii?, a?iii?oieia?/ii? no?oeoo?e oa aeeoo?aeo oi?i
aeae?a, iaaaaeaieo o «Eiiniaeo?…».

Aeyaeaii oae? iniaeeaino? nenoaiaoe/ii? no?oeoo?e neiaio?iiii? oei?e
?aa?iio aeine?aeaeaiue. Caaaeueiee aeaeiaee neeaae noaiiaeoue 365
aeae?a c 54 ?iaeei oa 213 ?iae?a. Aeanyoue i?ia?aeieo ?iaeei
aeeth/athoue 240 aeae?a (66%) oei?e ?aeiio. Iaea?eueoo e?euee?noue
aeae?a aeeth/athoue ?iaeeie Asteraceae (63 aeaee, 17,2%), Poaceae (32,
8,49%), Fabaceae (27, 7,39%), Brassicaceae (23, 6,30%), Scrophulariaceae
(19, 5,20%), Caryophyllaceae (18, 4,93%), Lamiaceae (17, 4,65%),
Rosaceae (16, 4,65%), Chenopodiaceae (13, 3,56%), Polygonaceae (12
aeae?a, 3,28%). Aei ianooiieo aeanyoe ?iaeei iaeaaeeoue 70 aeae?a
(19%), ui oo?e/? iaioa a?ae iiia?aaeiueiai iieacieea. A na?aaeiueiio ia
iaeio ?iaeeio i?eiaaea? n?i aeae?a, a?eueoa iieiaeie ?iaeei (34 aai
62%) i?aaenoaaeai? iaeiei – o?ueiia aeaeaie. Iecueea aeaeiaa
iane/ai?noth, na?ae/eoue i?ioa, ui oei?a oa?eoi??e ?icaeaa?oueny a
aeno?aiaeueieo oiiaao.

O ?iaeiaiio niaeo?? neiaio?iiii? oei?e ia?aaaaeathoue oae? ?iaee:
Veronica – 9 aeae?a, Vicia oa Polygonum ii 6, Malva, Chenopodium,
Geranium, Euphorbia oa Potentilla ii 5, Poa, Trifolium oa Atriplex ii 4.
Aeniea iane/ai?noue aeaeaie ?iaeo Veronica ? iciaeith ena?io?eueiino?,
oa?aeoa?ii? aeey na?aaecaiiiii?nueei? oei?e.

Iniiai? e?euee?ni? iieacieee nenoaiaoe/iiai niaeo?o na?ae/aoue i?i oa,
ui i?ae aieeaii ae?yeueiino? ethaeeie a?aeaoaa?oueny o?ainoi?iaoe?y
oei?e oa ca?eueoaiiy ?ie? ?iaeei ? ?iae?a, oa?aeoa?ieo aeey ena?e/ieo
iaeanoae Na?aaecaiiiii?’y.

Aaia?ao?/iee aiae?c aeaii? oei?e na?ae/eoue i?i oa, ui ca oeiii a?aaeo
neiaio?iii? aeaee ?ineei ?aeiio aeine?aeaeaiue iaeaaeaoue aei 26 a?oi.
Iaeoe?oa i?aaenoaaeaieie a?aaeia?/ieie a?oiaie ? aiea?eoe/ia – 107
aeae?a, ?a?iac?enueea – 64, a?oie einiiiie?o?a – 50 oa aai?einiiiie?o?a
– 35, ?a?iiaenueei–na?aaecaiiiii?nueea – 27,
?a?iiaenueei–na?aaecaiiiii?nueei–no?aeiiac?aonueea – 15,
?a?iiaenueei–i?ai?/iiaia?eeainueea – 9 aeae?a. ?io? a?oie i?aaenoaaeai?
1-3 aeaeaie.

*ioe?e oeie a?aae?a (aiea?eoe/iee, ?a?iac?enueeee, einiiiie?oe oa
aai?einiiiie?oe), ia?aaaaeathoue o no?oeoo?? neiaio?iiii? oei?e
aa?oiuei? oa/?? aanaeio Aei?no?a. Cia/ia i?noea na?aae aiio?o?a
caeiathoue oe?ieia?aaeuei? aiea?eoe/i? oa ?a?iac?enuee? aeaee ?c
iacio?eueieo oa?oiiaaiue. Na?aae aaeaaioeaieo aeae?a ia?aaaaeathoue
einiiiie?oe oa aai?einiiiie?oe, ye? ? i?aaenoaaieeaie ena?io?eueieo
oa?oiiaaiue.

O neiaio?iii?e oei?? ?aa?iio aeine?aeaeaiiy ie aeyaeee 178 aeae?a
aaeaaioeaieo ?ineei, ye? iaeaaeaoue aei 127 ?iae?a oa 42 ?iaeei. Ca
?acoeueoaoaie oei?iaaiaoe/iiai aiae?co aeo?aeieo a?aae?a, iniiaieie
eiiiiiaioaie neiaio?iiii? oei?e aeine?aeaeoaaiiai ?aa?iio ?
i?aaenoaaieee Aeaaiueiai Na?aaecaiiiii?’y – 117 aeae?a (32%), cia/iith
? e?euee?noue ?ineeie, ui iathoue i?ai?/iiaia?eeainueea iioiaeaeaiiy
(25, 7,1%). *eneaiio a?oio – 15 aeae?a (4,1%) – o oei?? ?aeiio
neeaaeathoue aeaee ?a?iiaenueeiai iioiaeaeaiiy. C Ac?? oa I?aaeaiii?
Aia?eee iioiaeeoue ii 6 aeae?a (1,6%). ?io? aeo?aei? a?aaee o oei??
?aeiio i?aaenoaaeai? 1-3 aeaeaie. I?ioa /a?ac iaaeinoaoi?noue
iaiao?aeieo iaoa??ae?a iai ia aaeaeiny anoaiiaeoe iioiaeaeaiiy 8 aeae?a.

Aeey aiae?co aaeaaioeaii? o?aeoe?? ie aeei?enoaee o?e iacaeaaei? iaeei
a?ae iaeiiai iieacieee: /an ?i?a?aoe??, niin?a ?i?a?aoe?? oa nooi?iue
iaoo?ae?caoe??, ye? cai?iiiiiaai? O. O?aaea?ii [169] oa
aeei?enoiaothoueny aaaaoueia aeine?aeieeaie. Aei a?oie a?oaio?o?a ie
a?aeianee 92 aeaee (25,2%), eaiio?o?a – 86 aeae?a (23,5%) ?ineei.
Aaeeeo ?ieue o iioe?aii? aaeaaioeaieo ?ineei a?ae?a?a? niin?a
?i?a?aoe??. Caeaaeii a?ae niiniao ?i?a?aoe?? ie aecia/eee aea? a?oie:
ia?aaaaeio a?eueo?noue (37,2%) o ?aeii? noaiiaeyoue enaiio?oe, ia aeieth
a?aac?io?o?a i?eiaaea? 11,8%. Caeaaeii a?ae nooiaiy iaoo?ae?caoe??
aaeaaioeai? ?ineeie, aeyaeai? o ?aa?ii?, iiae?eaii ia /ioe?e a?oie:
aa??io?oe – 11 aeae?a (3%), aiaeio?oe – 145 (39,7%), aoaia?io?oe – 15
(4,1%) oa eieiiio?oe – 7 aeae?a (1,9%).

A aeieia?/iiio niaeo?? neiaio?iiii? oei?e ca a?aeiioaiiyi aei aieiae ia
ia?oiio i?noe? ? a?oia ena?iiacio?o?a – 178 aeae?a (48,7%), ia ae?oaiio
– iacio?o?a 143 aeaee (39,4%), o?ao? i?noea caeiathoue iaciena?io?o?a
– 23 aeaee (6,3%). Iaeiaio /eneaiieie ? a?oie a?a?io?o?a, ena?io?o?a oa
a?ae?io?o?a – a?aeiia?aeii, 9 (2,5%), 8 (2,1%) oa 4 aeaee (1%). Aeniea
/anoea ena?iiacio?o?a a aeieia?/iiio niaeo?? na?ae/eoue i?i
a?eaeecaoe?th oa?eoi??? oa ena?io?oecaoe?th ?ineeiiiai iie?eao i?ae
aieeaii aio?ii?/iiai oaeoi?a.

Ca nooiaiai i?enoinoaaiiy aei ?ioaineaiino? ina?oeaiiy iae/eneaii?oith ?
a?oia aae?io?o?a – 240 aeae?a (65,8%), ae?oaa i?noea caeiathoue
noe?iaae?io?oe – 118 aeae?a (32,3%), o?ao? – noe?io?oe – 7 aeae?a
(1,9%).

Ia iniia? aiae?co aeeoo?aeo oi?i anoaiiaeaii, ui aei oa?io?o?a o ?aeii?
aeine?aeaeaiiy iaeaaeaoue 185 aeae?a (50,8%). Ia ae?oaiio i?noe?
ciaoiaeyoueny aai?e?eioio?oe – 157 aeae?a (43%), a?oia oaia?io?o?a
iae?/o? 12 aeae?a (3,3%), oaiao?o?a – 7 aeae?a (1,9%) ? 4 aeaee (1%)
iaeaaeaoue aei a?oie e?eioio?o?a. Oaeei /eiii, aeii?ioth/ith aeeoo?aith
oi?iith o neiaio?iii?e oei?? ?aeiio ? oa?io?oe, ui iiynith?oueny ?o
aenieith aaeaioaoe??th aei aio?iiiaaiieo aeioii?a.

I?ae /an aeine?aeaeaiue a?iii?oieia?/ii? no?oeoo?e neiaio?iiieo aeae?a,
aeyaeaii, ui ia?aaaaeathoue o?aa’yieno? ?ineeie – 346 aeae?a, aai 94,8%
a?ae caaaeueii? ?o e?eueeino?. Aea?aaa oa eou? ia/eneaii? – 6 oa 13
aeae?a a?aeiia?aeii, aai 1,6 oa 3,6%, oea ia?aaaaeii caee/aa?e?
eoeueoeaiaai? aeaee ?ineei.

Aiae?c niinia?a iioe?aiiy neiaio?iiieo ?ineei na?ae/eoue i?i oa, ui 156
aeae?a iaeaaeaoue aei aiaiioi?iiai oeio. Aaeeeo ?ieue o ia?aianaii?
ae?anii? a?ae?a?athoue ? oaa?eie, ai o aeine?aeaeai?e oei?? aeyaeaii 150
ciioi??a. Aio?iiioi??a iae?/o?oueny 80 aeae?a. O 89 aeae?a a?aecia/aia
aeniea caeaoi?noue aei iie?oi???. Cia/ia e?euee?noue – 66 aeae?a, ca
oa?aeoa?ii iioe?aiiy ? aa?ioi?aie. Iaeiaio i?aaenoaaeai? o neiaio?iii?e
oei?? aaoioi?e – 10 aeae?a. Niin?a iioe?aiiy a?eueoino? ?ineei
aecia/a?oueny iayai?noth (a?aenooi?noth) ia?aoeiae o eaiaeoaoo? oa
aieeaii ethaeeie.

Ia iniia? aeanieo niinoa?aaeaiue e?euee?niiai (ia’?iiiai) iioe?aiiy
ie?aieo aeae?a a iaaeao ?aeiio o ?ineeiiiio iie?ea? aeyaeaii oae?
caeiiii??iino?: cae/aeii (ianiai) iioe?aii ii an?e oa?eoi??? 72 aeaee
(19,7%), /anoi – 145 (40%), nii?aaee/ii – 74 (20,2%), c??aeea — 43
(11,7), aeoaea ??aeei — 31 (8,4%).

?ICAe?E 6

?noi??y oi?ioaaiiy oa aeaa?aaeaoe?? ?ineeiiiai iie?eao i?ae ae??th
aio?ii?/iiai oaeoi?A

Aeey a?aeoai?aiiy ?noi??? iiyae oa ?icnaeaiiy aaeaaioeaieo ?ineei ia
oa?eoi??? ?aa?iio aeine?aeaeaiue ie i?iaaee ii??aiyeueiee aiae?c
oei?enoe/iiai neeaaeo ?aa?iio ca inoaii? 150 ?ie?a.

Ia?o? a?oioiai? oei?enoe/i? aeine?aeaeaiiy aa?oiuei? oa/?? aanaeio
Aei?no?a i?iaiaeeeeny o 1850-1870 ??. oai?nueeeie oa iieuenueeeie
aeine?aeieeaie. Iaeiiai?o? a?aeiiino? i?i oei?o ?aa?iio ciaoiaeeii o
i?aoe? A. Atheeaey [150], yeee aeey ?aa?iio aeine?aeaeaiue iaaiaeeoue
988 aeae?a, cie?aia – 97 aaeaaioeaieo.

1898 ?ieo i?io. I. Eioethaith [155] iioae?eiaaiee aeiaeaoie
aei oei?e Ae?iaiae/a ca cai?aie A. Oo?/einueeiai, ui i?noeoue 24 iiaeo
aaeaaioeaieo aeaee.

Ia ii/aoeo OO no. o oei?? I?eea?iaooy iia? aaeaaioeai? aeaee
aoee aeyaeai? iieuenueeeie aeine?aeieeaie.

O OO no. oe?a?inuee? aeine?aeieee ia oa?eoi??? I?eea?iaooy ciaeoee ua
ieceo iiaeo aaeaaioeaieo aeae?a ?ineei [30, 39, 52, 89, 99, 120, 126,
130 oa ?i.].

Aiae?c e?oa?aoo?ieo aeaea?ae, ui i?noyoue ?acoeueoaoe ?iaaioa?ecaoe??
oei?e, oa aeani? aeine?aeaeaiiy aeicaieeee iai a?aeoai?eoe ?noi??th
iiyae oa ?icnaeaiiy ?yaeo aaeaaioeaieo ?ineei, ui o?aieythoueny a
?aa?ii? aeine?aeaeaiue cae/aeii (ianiai) (Amaranthus retroflexus L.,
Galinsoga parviflora Cav., G. ciliata (Rafin.) Blake, Chamomilla
suaveolens (Putsh) Rydb., Erigeron canadensis L., Juncus tenuis Willd.,
Impatiens parviflora DC.) aai c??aeea (Ambrosia artemisiifolia, Carduus
nutans L., Commelina communis, Centaurea diffusa, Cyclachaena
xanthifolia (Nutt.) Fresen., Echinocystis lobata, Impatiens glandulifera
Royle, Solanum cornutum, Xanthoxalis dillenii (Jacq.) Holub oa ?i.).

Oae, ?acii c ian?iiei iaoa??aeii 1886 ?ieo aei ?a?iie c I?ai?/ii?
Aia?eee aeiaaeeiai iio?aiea Solanum cornutum Lam. A Oe?a?i? aeae
aeyaeaii 1928 ?ieo Ae.E. Ea??iiiaei o iin?aao ca?iiaeo eoeueoo? ia
Ee?auei?. O i?neyai?ii? ?iee nii?aaee/ii cono??/aany a e?ninoaiiaeo oa
noaiiaeo ?aeiiao Oe?a?ie [110]. C?inoa? ia?aaaaeii ia aaooaiao, na?aae
ethoea?ie, o naaeao, ia aeiia?aaeieeao, a?ey aoae?aaeue. Ca 40 ?ie?a
cia/i? eieii?? noi?ioaaeeny o E??iaia?aaenuee?e, Aei?i?iiao?ianuee?e,
Caii??cuee?e, Iaeanuee?e, Ieeiea?anuee?e, Oa?niinuee?e, Eoaainuee?e oa
Aeiiaoeuee?e iaeanoyo. Nii?aaee/ii c?inoa? o noaiia?e cii? E?eio oa
A?iieoeuee?e iaeano? [17]. Iei? aeae i?iaeiaaeo? ?icnaeyoueny o
i?ai?/ii-cao?aei? ?aeiie E?ninoaio. Ie aia?oa aeyaeee ia cae?cie/i?e
eie?? o i.No?ee (Euea?anueea iae.).

Solanum cornutum – eaiio?o i?ai?/iiaia?eeainueeiai iioiaeaeaiiy c
?a?iiaenueei–na?aaecaiiiii?nueei–i?ai?/iiaia?eeainue-eei a?aaeii. Ia
oa?eoi??? Oe?a?ie i?ai?/ia iaaea eiai iioe?aiiy i?ioiaeeoue /a?ac
noaiiao ciio oa E?ei. E??i oiai, o No?aei?e ?a?ii? iioe?aiee o
Ieaeiueiaiecueeiio ?aeii?.

Aeae c?inoa? a iin?aao, naaeao, ia ai?iaeao oa acaeiaae cae?cieoeue.

Caaaeueia iioe?aiiy: Iaeneea ? i?aaeaiii-cao?aeia /anoeia NOA, a oaeiae
Na?aaeiy ?a?iia oa Na?aaecaiiiii?(y [109].

sse aa/eii, aio?iiiaai?caoe?y oa?eoi??? ni?ey? iioe?aiith aaaaoueio
aaeaaioeaieo aeae?a ?ineei, ianaiia?aae oeo, ui iathoue aenieo
aeieia?/io ieanoe/i?noue oa aeeoo?a?noue. A aonoa ia?aaea oeyo?a
niieo/aiiy a ?aeii? aeine?aeaeaiiy ni?ey? ooai?aiith i?a?aoe?eieo
ei?eaei??a aeey aaeaaio?a oa ?o iioe?aiith a iia? ?aa?iie.

Neeaae ? no?oeoo?a niaoeeo?/ieo oei?a i?noeac?inoaiue aecia/athoueny
oiiaaie iaaiiai aeioiio ? a cia/i?e i??? a?ae??ciythoueny iioiaeaeaiiyi
? iai?yieii ?icaeoeo i?ae aieeaii aio?ii?/ieo oaeoi??a. Oeiiaeie oi?iaie
aio?iiiaai?ciaaiiai eaiaeoaooo ? aa?ioeaiice, aaoiiia?euei? oa
cae?cie/i? oeyoe, ni?oieee, aoeeoe?, aaciie, eeoiae, aoae?aaeuei?
iaeaeai/eee oa naeeoaai? oa?eoi??? – oiaoi ae?eyiee, ia yeeo i?e?iaeiee
iie?ea ?noioii ii?ooaiee aai iiai?noth cieuaiee. ?o aeaeiaee neeaae i?e
an?e ??ciiiai?oiino? ia? aaaaoi ni?eueiiai. Oea iiynith?oueny oe?ieith
aeieia?/iith ieanoe/i?noth a?eueoino? ao?’yi?a, ianaiia?aae
aaeaaioeaieo, ? oei, ui a o?aai?ciaaiiio na?aaeiaeu? niaeo? aeieia?/ieo
oaeoi??a cia/ii aoae/ee, i?ae o i?e?iaeieo oiiaao. A i?ioean?
i?e?iaaiethoe?? ia o?ainoi?iiaaieo aeioiiao, iiae?aieo ca aaeao?/ieie
oiiaaie oa aeaeii aio?ii?/iiai aieeao, oi?iothoueny aeoaea aeecuee? ca
neeaaeii ?ineeii? oa?oiiaaiiy, ye? iiaeooue aooe a?i?iaeeeaoi?aie ci?i o
aeiae?ee?.

?ICAe?E 7

I?iaiic iiaeeeaeo ci?i ?ineeiiiai iie?eao i?ae aieeaii aio?iiiaaiieo
oaeoi??a

Ia iniia? i?iaaaeaieo aiae?c?a oa aeyaeaieo iniaeeainoae
neiaio?iiii? oei?e ie ?ic?iaeee i?iaiic iiaeeeaeo ci?i ?ineeiiiai
iie?eao i?ae ae??th aio?ii?/ieo oaeoi??a.

Aiane?aeie neiaio?ii?caoe?? ?aa?iiaeueia oei?a a iiaeaeueoiio
caaaaoeoueny ia aaeaaioeai? aeaee, ui iaoiiaeoue ci?io nenoaiaoe/ii?,
aaia?ao?/ii?, aeieia?/ii? no?oeoo?e oa niaeo?a aeeoo?aeo oi?i.
?icoe?youeny oei?iaaiaoe/i? ca’ycee oei?e ?aa?iio aeine?aeaeaiiy c
oei?ith ena?e/ieo iaeanoae ?a?ac?? oa ?ioeo eiioeiaio?a.

Ia iniia? aiae?co niaeo?a nenoaiaoe/ii? no?oeoo?e iiaeia ia?aaeaa/aoe,
ui e?euee?noue neiaio?iiieo aeae?a ia?aaaaeii aoaea ca?eueooaaoeny ca
?aooiie i?aaenoaaiee?a ?iaeei Asteraceae, Poaceae, Brassicaceae,
Chenopodiaceae, ai naia oe? ?iaeeie iaea?eueoa i?aaenoaaeai? a
?aa?iiaeuei?e oei??. Iineeaiiy ?ie? neiaio?iiieo aeae?a o iiaeaeueo?e
o?ainoi?iaoe?? oei?e ni?e/eith? nooo?a? ci?ie ia an?o ??aiyo –
iiioeyoe?eiiio, aeaeiaiio, oeaiioe/iiio oa aeinenoaiiiio.

Aaeaaioeaia o?aeoe?y iiaea iiiiaieoeny aeaeaie ?c ?iae?a Chenopodium,
Amaranthus, Malva, Veronica, Geranium, Euphorbia, ui aaea i?aaenoaaeai?
cia/iith e?euee?noth aeae?a o oei?? ?aa?iio. ?c aiio?oii? o?aeoe?? aei
neiaio?iiii? oei?e oa?eaeooue ?ineeie ?c ?iae?a Potentilla, Silene,
Polygonum, Rumex, Cirsium, Poa, Verbascum, ui iathoue aenieo aeieia?/io
ieanoe/i?noue ? eaaei aaeaioothoueny aei iiaeo oiia ?nioaaiiy.

A iaeaooiueiio niinoa??aaoeiaoueny oaiaeaioe?y aei ca?eueoaiiy o
oei?enoe/iiio niaeo?? ?aa?iio aae?io?o?a, ena?iiacio?o?a, oa?io?o?a.
Oae? ?ineeie iathoue a?eueoo aeieia?/io ieanoe/i?noue oa oaeaeoa
aaeaioothoueny aei ci?iaieo oiia.

AENIIAEE

Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio Aei?no?a iae?/o? 365
aeae?a, ui iaeaaeaoue aei 213 ?iae?a oa 54 ?iaeei. Aeey caoiaeo Oe?a?ie
aeyaeaii 6 iiaeo aeae?a.

A ?acoeueoao? aiae?co nenoaiaoe/ii? no?oeoo?e neiaio?iiii? oei?e
c’yniaai? ?? oa?aeoa?i? ?ene, ye? iieyaathoue a iiae?aiino? niaeo?a
i?ia?aeieo ?iaeei c aiaeia?/ieie oei?aie ?ioeo oa?eoi??e. Anoaiiaeaii,
ui a ?iaeiaiio niaeo?? ia?aaaaeathoue ?iaee c iaeith aeaeiaith
iane/ai?noth, ui oa?aeoa?ii aeey oei? oa?eoi??e c aeno?aiaeueieie
oiiaaie ?icaeoeo ?ineeiiiai iie?eao. A?aei?ii?noue a?ae niaeo?a
i?ia?aeieo ?iaeei neiaio?iiii? oei?e Oe?a?ie a?aeia?aaea? aeeeth/ii
aeniea iieiaeaiiy ?iaeeie Fabaceae – oeiiai? aeey Na?aaecaiiiii?nueei?
iaeano?, cia/io ?ieue ?iaeeie Chenopodiaceae, oa?aeoa?ii? aeey
ionoaeueieo oei?, a oaeiae aoiaeaeaiiy a ia?oo aeanyoeo ?iaeei Rosaceae
oa Polygonaceae, i?aaenoaaieee yeeo iioe?ai? a i?ai?/i?e iii??i?e
cii?.

Aiae?c a?iii?oieia?/ii? oa aeieia?/ii? no?oeoo?e neiaio?iiii? oei?e
na?ae/eoue i?i na?aaecaiiiii?nueeee oa?aeoa? oei?e ?aeiio
aeine?aeaeaiiy. Na?aae aeia?iii?o iaea?eueoa aae?io?o?a ?
iaciena?io?o?a, ui oa?aeoa?ii aeey ciie noai?a. Ia a?aei?io a?ae
?aa?iiaeueii? oei?e, o neiaio?iii?e oei?? ?aa?iio ca aaa?oonii oa
o?eaae?noth aeeooy ia?aaaaeathoue iiiiea?i?ee.

Iiaeaii eeaneo?eaoe?th neiaio?iiieo aeae?a ?ineei, ?ic?iaeaio ca
oi?i–oei?i–?noi?e/ieie e?eoa??yie ? iciaeaie iaoo?ae?caoe?? oa
aaeaioaoe??. Cai?iiiiiaaia nenoaia caaacia/o? aoaeoeai?oa i?iaaaeaiiy
aiae?co neiaio?iiii? oei?e.

Aeae?eaii aiio?oio oa aaeaaioeaio o?aeoe??. Ni?aa?aeiioaiiy i?ae ieie
noaiiaeoue 1:1, ui na?ae/eoue i?i cia/io ?ieue aaeaaioeaieo aeae?a o
oi?ioaaii? neiaio?iiii? oei?e, i?e/iio ca /anii i?iieeiaiiy
ia?aaaaeathoue a?oaio?oe, ca niiniaii ?i?a?aoe?? – enaiio?oe, a ca
nooiaiai iaoo?ae?caoe?? – aiaeio?oe. Iaoo?ae?caoe?y aaeaaioeaieo
aeae?a a?aeaoaa?oueny ia?aaaaeii a ?ineeiieo oa?oiiaaiiyo
o?ainoi?iiaaieo aeioii?a, a oie aea /an o i?e?iaeiiio ?ineeiiiio iie?ea?
/anoioa ?o iiyae ciaioo?oueny.

Aaia?ao?/ia no?oeoo?a aeine?aeaeoaaii? oei?e na?ae/eoue i?i oa, ui o
neeaae? neiaio?iiii? oei?e aa?oiuei? oa/?? aanaeio Aei?no?a
ia?aaaaeathoue /ioe?e oeie a?aae?a (aiea?eoe/iee, ?a?iac?aenueeee,
einiiiie?oe oa aai?einiiiie?oe). Cia/ia i?noea na?aae aiio?o?a
caeiathoue oe?ieia?aaeuei? aiea?eoe/i? oa ?a?iac?enuee? aeaee ?c
iacio?eueieo oa?oiiaaiue. Na?aae aaeaaioeaieo aeae?a ia?aaaaeathoue
einiiiie?oe oa aai?einiiiie?oe, ye? ? i?aaenoaaieeaie ena?io?eueieo
oa?oiiaaiue. Aiane?aeie neiaio?ii?caoe?? oei?a ?aa?iio caaaaoeeany
oei?enoe/ieie aeaiaioaie, aaia?ao?/ii iia’ycaieie c ena?e/ieie
oa?eoi??yie caiii? eoe?.

Oei?iaaiaoe/iee aiae?c aeo?aeieo a?aae?a na?ae/eoue i?i aenieo
iane/ai?noue neiaio?iiii? oei?e ?aa?iio aeine?aeaeaiiy aaeaaioeaieie
aeaeaie na?aaecaiiiii?nueeiai, ac?aonueeiai oa i?ai?/iiaia?eeainueeiai
iioiaeaeaiiy o ca’yceo c iiaioaaiiyi ieie iiaeo aio?iiiaaiii
ia?aoai?aieo aeioii?a.

Aena?oeaii ?noi??th iiyae oa iioe?aiiy ?yaeo aaeaaioeaieo ?ineei ia
oa?eoi??? Oe?a?ie. Ia iniia? neeaaeaieo ea?o a?aae?a aeyaeaii caaaeuei?
caeiiii??iino? i?a?aoe?? ia oa?eoi??? Oe?a?ie na?aaecaiiiii?nueeeo oa
ac?aonueeeo aeae?a a i?ai?/ii-cao?aeiiio iai?yieo.

?icaeyiooi oeiia? o?ainoi?iiaai? aeioiie, ui a?ae??ciythoueny ca
nooiaiai oa oa?aeoa?ii aio?iiiaaiiiai aieeao. Aeyaeaii a?aeiinii no?ee?
aeaee aei ??cieo oi?i ? o?eaaeino? aieeao aio?ii?/ieo i?an-oaeoi??a, ye?
iiaeooue a?ae?a?aaaoe ?ieue a?i?iaeeeaoi??a caaaeueieo ci?i o aeiae?ee?.

Ia i?aenoaa? oei?ieia?/ii? ioe?iee iane?aee?a neiaio?ii?caoe?? oa ?o
i?iya?a ia i?aai?ciiaiio, iiioeyoe?eiiio e aeinenoaiiiio ??aiyo
anoaiiaeaii, ui aiia a?aeaoaa?oueny a iai?yieao ci?ie no?oeoo?ii?
i?aai?caoe?? oei?e, aaeaioaoe?? iniaei e iiioeyoe?e aei ae??
aio?iiiaaiieo oaeoi??a ? ia?inoaiiy ?? aeeiai?/ieo oaiaeaioe?e.

Nienie iaoeiaeo i?aoeue, iioae?eiaaieo ca oaiith aeena?oaoe??

Neeaa TH.A. Neiaio?iiia oei?a aa?o?a’y aanaeio Aei?no?a oa i?iaeaie ??
eeaneo?eaoe?? // Iaoeia? caienee Oa?iii. aea?ae. iaae. oi-oo ?i.
Aieiaeeie?a Aiaothea. – Oa?iii?eue. – 1998.– 2 (4).– N.19-22.

Neeaa TH.A. Aeioiiieia?/ia i?eo?i/ai?noue neiaio?iiieo ?ineei // Iaoeia?
caienee Oa?i. aea?ae. iaae. oi-oo ?i. Aieiaeeie?a Aiaothea. –
Oa?iii?eue. – 1998. – 3 (4). – N.5-10.

Neeaa TH.A. ?iaac?y oa iaoo?ae?caoe?y aaeaaioeaieo ?ineei o aa?oi?e
oa/?? aanaeio Aei?no?a// Iaoeia? caienee Oa?i. aea?ae. iaae. oi-oo ?i.
Aieiaeeie?a Aiaothea. – Oa?iii?eue. – 1999. – 1(4). – N.7-11.

Neeaa TH.A. Iaeyae nenoai eeaneo?eaoe?? neiaio?iiii? oei?e oa i?iaeaie
?? aea/aiiy // Iaoeia? caienee: Ca??iee iaoeiaeo noaoae Iaoe. iaae.
oi-oo ?i. I.I. Ae?aaiiaiiaa – Ee?a: IIO, 1998.– Aei.2 – N. 165-172.

Neeaa TH.A. Aaeaaioeai? ?ineeie aa?oiuei? oa/?? aanaeio Aei?no?a
(Euea?anueea iae.) // I?iieneiaa aioai?ea: noai oa ia?niaeoeae
?icaeoeo (Iaoa??aee O?aouei? i?aeia?iaeii? eiioa?aioe??). – Aeiiaoeuee,
Aaainoai «Ioeueoei?an», 1998. – N.119-125

Neeaa TH.A., Iaaeai?iee A.A. No/aniee noai ?ineeiieo anioe?aoe?e oa
oaa?eiieo oa?oiiaaiue ia cae?cie/ieo aei?iaao I?eea?iaooy //
I?aeia?iaei? aniaeoe aea/aiiy oa ioi?iie a?i??ciiiai?ooy Ea?iao. –
?ao?a, 1997. – N. 188-191.

Iaaeai?iee A.A., Neeaa TH.A. Aeieia?/ia iiaeaeue i?noa — iaeia c oi?i
iiaeeaeaiiai aea/aiiy o/iyie oeeeeo i?e?iaeie/i-aaia?ao?/ieo aeenoeeie?i
// Oai??y ? i?aeoeea aea/aiiy i?e?iaeie/eo aeenoeeie?i o aeu?e oa
na?aaei?e oeie?. – Oiaiue, 1997.– N. 117-119.

Neeaa TH.A. ?noi?eei–a?ae?ia?ao?/iee iaeyae aea/aiiy neiaio?iiii? oei?e
aanaeio Aei?no?a // Iaoa??aee iaoeiaeo /eoaiue, i?enay/aieo 100-??//th
a?aee?eooy iiaea?eiiai caie?aeiaiiy o iie?eoiian?iieo ?ineei i?ioani?ii
oi?aa?neoaoo Nayoiai Aieiaeeie?a N.A.Iaaaoeiei. – E.: O?oinioe?ioeaio?,
1998. – N. 202-204.

Neeaa TH.A. Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio Aei?no?a
(Oe?a?ia).– ?oeiien.

Aeena?oaoe?y ia caeiaoooy a/aiiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
c? niaoe?aeueiino? 03.00.05. – aioai?ea. Oeaio?aeueiee aioai?/iee naae
?iai? I.I. A?eoea IAI Oe?a?ie, Ee?a, 1999.

Aeena?oaoe?y i?enay/aia ieoaiiyi ?iaaioa?ecaoe?eiiai, oi?ieia?/iiai ?
no?oeoo?iiai aea/aiiy neiaio?iiii? oei?e aa?oiuei? oa/?? aanaeio
Aei?no?a. Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio Aei?no?a ia?aoiao?
365 aeae?a ?c 213 ?iae?a oa 54 ?iaeei. Anaa?/ii i?iaiae?ciaaii no/aniee
noai neiaio?iiii? oei?e aeaiiai ?aeiio. Aeae?eaii o?aeoe?? oei?e. Aeey
aeae?a ?c aaeaaioeaii? o?aeoe?? i?iaaaeaii aiae?c aeo?aeieo a?aae?a, a
oaeiae a?aeoai?aii ?noi??th ?i?a?aoe?? ? ?icnaeaiiy iecee aaeaaioeaieo
?ineei ia oa?eoi??? Oe?a?ie. Anoaiiaeaii aeieia?/io i?eo?i/ai?noue
neiaio?iiieo aeae?a aei aio?iiiaaiieo aeioii?a. Iaaaaeaii i?iaiic
iiaeeeaeo ci?i ?ineeiiiai iie?eao i?ae aieeaii ainiiaea?nueei?
ae?yeueiino?.

Eeth/ia? neiaa: Aei?noa?, neiaio?iiia oei?a, no?oeoo?a, eeaneo?eaoe?y,
aaeaaioe, aiio?oe, ?i?a?aoe?y, neiaio?ii?caoe?y.

Neeaa TH.A. Neiaio?iiiay oei?a aa?oiaai oa/aiey aannaeia Aeiano?a
(Oe?aeia).– ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.05 – aioaieea. Oeaio?aeueiue aioaie/aneee
naae ei. I.I. A?eoei IAI Oe?aeiu, Eeaa, 1999.

Aeenna?oaoeey iinayuaia aii?inai eiaaioa?ecaoeeiiiiai, oi?ieiae/aneiai
e no?oeoo?iiai eco/aiey neiaio?iiiie oei?u aa?oiaai oa/aiey aannaeia
Aeiano?a. Neiaio?iiiay oei?a aa?oiaai oa/aiey aannaeia Aeiano?a
ian/eouaaao 365 aeaeia ec 213 ?iaeia e 54 naiaenoa.

Ia iniiaa aiaeeca nenoaiaoe/aneie no?oeoo?u neiaio?iiiie oei?u
auyaeaiiua aa oa?aeoa?iua iniaaiiinoe, eioi?ua i?iyaeythoueny a
noiaenoaa niaeo?a ainiiaenoaothueo naiaenoa n aiaeiae/iuie oei?aie
ae?oaeo oa??eoi?ee. Onoaiiaeaii, /oi a ?iaeiaii niaeo?a i?aainoiaeyo
?iaeu n ieceie aeaeiaie ianuuaiiinoth, /oi oa?aoa?ii aeey oei? n
yeno?aiaeueiuie oneiaeyie ?acaeoey ?anoeoaeueiiai iie?iaa. Io niaeo?a
ainiiaenoaothueo naiaenoa neiaio?iiiie oei?u Oe?aeiu ioee/aaony aunieei
iieiaeaiea naiaenoaa Fabaceae – oeie/iie aeey N?aaeecaiiiii?neie
iaeanoe, cia/eoaeueiay ?ieue naiaenoaa Chenopodiaceae, oa?aoa?ioth aeey
ionouiiuo oei?, a oaeaea a ia?aoth aeanyoeo aioee naiaenoaa Rosaceae e
Polygonaceae, i?aaenoaaeoaee eioi?uo i?iec?inoatho a naaa?iie oia?aiiie
ciia.

Aiaeec aeiii?oieiae/aneie e yeieiae/aneie no?oeoo?u neiaio?iiiie oei?u
naeaeaoaeuenoaoao i n?aaeecaiiiii?neii oa?aeoa?a oei?u ?aeiia
enneaaeiaaiee. N?aaee yeiaeiii?o i?aiaeaaeatho aaeeioeou e
iaciena?ioeou, /oi oa?aeoa?ii aeey ciiu noaiae. A ioee/ea io
?aaeiiaeueiie oei?u a neiaio?iiiie oei?a ?aeiia enneaaeiaaiee ca
aaaeoonii e i?iaeieaeeoaeueiinoth aeecie i?aiaeaaeatho iiiiea?ieee.

Ananoi?iiia i?iaiaeece?iaaii nia?aiaiiia ninoiyiea neiaio?iiiie oei?u
aeaiiiai ?aaeiia. Auaeaeaiu o?aeoeee oei?u. Niioiioaiea iaaeaeo ieie
ninoaaeyao 1:1, /oi naeaeaoaeuenoaoao i cia/eoaeueiie ?iee aaeaaioeaiuo
aeaeia a oi?ie?iaaiee neiaio?iiiie oei?u, i?e /ai ca a?aiaiai
eiiea?aoeee i?aiaeaaeatho a?oaioeou, ca niiniaii eiiea?aoeee –
enaiioeou, a ca noaiaiueth iaoo?aeecaoeee –yiaeioeou.

Aaia?aoe/aneay no?oeoo?a enneaaeoaiie oei?u naeaeaoaeuenoaoao i oii, /oi
a ninoaaa neiaio?iiiie oei?u aa?oiaai oa/aiey aannaeia Aeiano?a
aeiieie?otho /aou?a oeia a?aaeiae/aneeo a?oii (aiea?eoe/aneay,
aa?aceaoneay, einiiiieeoiay e aaieeiniiiieeoiay). Cia/eoaeueiia ianoi
n?aaee aiioeoia caieiatho oe?ieia?aaeueiua aiea?eoe/iua e aa?aceeneea
aeaeu ec iacioeeueiuo niiauanoa. N?aaee aaeaaioeaiuo aeaeia
i?aiaeaaeatho einiiiieeou e aaieeiniiiieeou, eioi?ua anoue
i?aaenoaaeoaeyie ena?ioeeueiuo niiauanoa. A ?acoeueoaoa neiaio?iiecaoeee
ai oei?a ?aaeiia oaaee/eeinue eiee/anoai aeaeia aaia?aoe/anee
i?eo?i/aiiuie e oa??eoi?eyi n ena?e/aneeie oneiaeyie.

Aeey aeaeia ec aaeaaioeaiie o?aeoeee i?iaaaeai aiaeec enoiaeiuo a?aaeia.
Ii naeaeaoaeuenoaoao i?i aunieoth ianuuaiiinoue neiaio?iiiie oei?u
?aeiia enneaaeiaaiee aaeaaioeaiuie aeaeaie n?aaeecaiiiii?neiai,
aceaoneiai e naaa?iaia?eeaineiai i?ienoiaeaeaiey a nayce n iniaiaieai
eie iiauo aio?iiiaaiii i?aia?aciaaiiuo yeioiiia.

Ainnicaeaia enoi?ey eiiea?aoeee e ?annaeaiey ?yaea aaeaaioeaiuo
aeaeia ia oa??eoi?ee Oe?aeiu. Ia iniiaa ninoaaeaiiuo ea?o a?aaeia
onoaiiaeaiu iauea caeiiiia?iinoe iea?aoeee ia oa??eoi?ee Oe?aeiu
n?aaeecaiiiii?neeo e aceaoneeo aeaeia a naaa?i–caiaaeiii iai?aaeaiee.
Onoaiiaeaia yeieiae/aneay i?eo?i/aiiinoue neiaio?iiiuo aeaeia e
aio?iiiaaiiui yeioiiai.

Ia iniiaa oei?ieiae/aneie ioeaiee iineaaenoaee neiaio?iiecaoeee e eo
i?iyaeaiee ia i?aaieciaiiii, iiioeyoeeiiiii e yeinenoaiiii o?iaiyo
onoaiiaeaii, /oi iia i?ienoiaeeo a iai?aaeaiee ecieiaiey no?oeoo?iie
i?aaiecaoeee oei?u, aaeaioaoeee aeaeia e iiioeyoeee e aeaenoaeth
aio?iiiaaiiuo oaeoi?ia e ia?anoaiea aa aeeiaie/iuo oaiaeaioeee.

Eeth/aaua neiaa: Aeiano?, neiaio?iiiay oei?a, no?oeoo?a, eeanneoeeaoeey,
aaeaaiou, aiioeou, eiiea?aoeey, neiaio?iiecaoeey.

Skyba Yu.A. The synanthropic flora of the upper reaches of the Dnistro
(Dnister) River, Ukraine.– Manuscript.

Dissertation on pretence of candidate of biological scienses degree.
Fild of study 03.00.05 – botany. M.M. Grishko Central Botanical Garden
of National Academy of Sciences of Ukraina, Kiev, 1999.

Dissertation is devoted to inventarization, chorological and structural
study of synantrope Flora in the upper course of Dnister river.
Synanthropic flora of the region mentioned includes 365 species, 213
genuses and 54 families, its modern condition has been analysed.
Different flora of has been determined. Initialales of adventive
fraction’s species has been defined. Immigration and distribution time
has been found out for some adventive species. Synanthropic species has
been reffered to antropogenic ecotops. The forecast of probable
successive changes in flora under infuence of human activities is given.

Key words: Dnister river, synanthropic flora, stuctura, classification,
adventive plants, apophytos, immigration, synantropization.

IAOe?IIAEUeIA AEAAeAI?ss IAOE OE?A?IE

Oeaio?aeueiee aioai?/iee naae ?iai? I.I. A?eoea

NEEAA TH??E AIAe??EIAE*

OAeE 581.9. 632.51 (477)

NEIAIO?IIIA OEI?A AA?OIUeI? OA*?? AANAEIO AeI?NO?A (OE?A?IA)

03.00.05 – aioai?ea

Aaoi?aoa?ao aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy
eaiaeeaeaoa a?ieia?/ieo iaoe

Ee?a — 1999

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia a Iaoe?iiaeueiiio iaaeaaia?/iiio oi?aa?neoao? ?i. I.I.
Ae?aaiiaiiaa, I?i?noa?noai ina?oe Oe?a?ie

Iaoeiaee ea??aiee: eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe, i?ioani?,

A?eoei — Aiaiaiei Ai?en Einoyioeiiae/

Iaoe?iiaeueiee iaaeaaia?/iee oi?aa?neoao ?i. I.I. Ae?aaiiaiiaa,
aeaeai i?e?iaeie/i-aaia?ao?/iiai oaeoeueoaoo

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe, i?ioani? *iiee

Aieiaeeie? ?aaiiae/, Iaoe?iiaeueiee
oi?aa?neoao

?i. Oa?ana Oaa/aiea, i?ioani? eaoaae?e

aioai?ee

eaiaeeaeao a?ieia?/ieo iaoe Iinye?i Na?a?e

Eaii?aeiae/, ?inoeoooo aioai?ee

?i. I.A. Oieiaeiiai IAI Oe?a?ie,
caa?aeoth/ee

a?aeae?eii nenoaiaoeee aeueo ?ineei

I?ia?aeia onoaiiaa: Euea?anueeee aea?aeaaiee oi?aa?neoao ?i. ?aaia

O?aiea, I?i?noa?noai ina?oe Oe?a?ie,
eaoaae?a

aioai?ee, i. Euea?a

Caoeno a?aeaoaeaoueny «_19__» _aa?aciy___ 1999 ?. i 10_ aiaeei? ia
can?aeaii? Niaoe?ae?ciaaii? a/aii? ?aaee E. 26.215.01 Oeaio?aeueiiai
aioai?/iiai naaeo ?i. I.I.A?eoea IAI Oe?a?ie ca aae?anith: 252014, i.
Ee?a, aoe. O?i??yc?anueea, 1.

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? Oeaio?aeueiiai
aioai?/iiai naaeo ?i. I.I. A?eoea IAI Oe?a?ie ca aae?anith: 252014, i.
Ee?a, aoe. O?i??yc?anueea, 1.

Aaoi?aoa?ao ?ic?neaiee ________18 ethoiai 1999 ?.____

A/aiee nae?aoa?

Niaoe?ae?ciaaii? a/aii? ?aaee ________ Ioce/oe A.I.

Anooi

Aeooaeuei?noue oaie. Cia/aiiy ciaiue i?i oei?o iineeththoueny a aniaeo?
iieiaeaiue I?aeia?iaeii? eiiaaioe?? i?i caa?aaeaiiy a?i??ciiiai?oiino?
(i. ??i–aea–AEaiae?i, 1992) oa ?aciethoe?? I?aeia?iaeii? eiioa?aioe??
Planta Europa (i. ??? (O?aioe?y), 1995). Ci?ie a a?inoa?? i?ecaiaeyoue
aei ciaioaiiy e?eueeino? aeae?a ?ineei oa ao?aoe oe?iiiai aaiioiiaeo
oei?e. Oaeaee? oaiie ?icaeoeo noni?euenoaa ? iineeaiiy aieeao ethaeeie
ia i?e?iaeiee ?ineeiiee iie?ea ni?e/eiththoue neiaio?ii?caoe?th
aai?eaaiii? oei?e oa ?ineeiiino?. Oiio, iaeiei c aieiaieo iaoeiaeo
caaaeaiue ? aea/aiiy no/aniiai noaio oei?e oa eiai ci?i aiane?aeie
aio?ii?/iiai aieeao.

Ii/aoie anaa?/iiio aea/aiith neiaio?iiii? oei?e Oe?a?ie iieeaaeaii
i?aoeyie I.?. Eioiaa, A.A. I?ioiiiiiai? oa ?i. Aeioaia? i?iaaaeaii
?aa?iiaeuei? aeine?aeaeaiiy neiaio?iiii? oei?e Ee?anueeiai Iie?nny (N.E.
Iinye?i), I?aaeiy Oe?a?ie (?.?. Ao?aea, A.I. Oiooa?ue, N.A.Iao?ee, O.A.
Aaneeue?aa–Iaia?oeaeiaa), Caea?iaooy (A.A. I?ioeue). Ie aeine?aeaeoaaee
i?ai?/ii–cao?aeio /anoeio I?eea?iaonueeiai oei?enoe/iiai ie?oao.

Oei?o aeine?aeaeoaaii? oa?eoi??? ii/aee aea/aoe c O?O no. oai?nuee?,
iieuenuee? oa oe?a?inuee? aeine?aeieee: O. Aa?a?o, A. Atheeaeue, Aeae.
Eiai, A. Oo?/einueeee, N. Eoeue/einueeee, A. Iaaeianueeee, I.?.Eioia,
A.?. *iiee, TH.?. Oaeya-Niniiei, N.I. Noieei, A.O.Caeai/oe, A.I. Oea/ee
oa ?i. I?ioa niaoe?aeuei? aeine?aeaeaiiy neiaio?iiieo aeae?a ?ineei
oeueiai ?aa?iio ia i?iaiaeeeeny, aoee a?coaeueieie ? o?aaiaioa?ieie.

?aa?iiaeuei? aeine?aeaeaiiy neiaio?iiii? oei?e ni?eyoeiooue ooi/iaiith
?? aeaeiaiai neeaaeo, a anaa?/iee aiae?c – aeyaeaiith caeiiii??iinoae
?? oi?ioaaiiy, ui ? oe?iiei iaoa??aeii aeey ?ic?iaee i?iaiico ?icaeoeo
oei?e i?ae aieeaii aio?ii?/iiai /eiieea, ye ia eieaeueiiio, oae ? ia
ieaiaoa?iiio ??aiyo, ui ia no/aniiio aoai? ? iaoiiaeth? ?ic?iaeo oaie
aeena?oaoe?eii? ?iaioe: «Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio
Aei?no?a».

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie.
Aeena?oaoe?eia ?iaioa aeeiiaia ia eaoaae?? aioai?ee IIO ?i.
I.I.Ae?aaiiaiiaa o 1995-1998 ??. ? iia’ycaia c ieaiii
iaoeiai–aeine?aeii? ?iaioe eaoaae?e ca oaiith «Ci?no ina?oe: oi?ie,
iaoiaee ? caniae i?aeaioiaee a/eoae?a».

Iaoa ? caaaeaiiy aeine?aeaeaiiy. Iiaeaoe iiaiee eiiniaeo
neiaio?iiii? oei?e ?aa?iio, aecia/eoe no/aniee noai oei?e oa i?iaanoe
?? aiae?c. I?inoaaeeoe ?noi??th oi?ioaaiiy neiaio?iiii? oei?e ? aeyaeoe
iiaeeea? oaiaeaioe?? ?? o?ainoi?iaoe?? i?ae ae??th aa?ioe/ieo, a?ioe/ieo
oa ?noi?e/ieo oaeoi??a.

Ia?oi/a?aia? caaaeaiiy aoee oaeeie:

anoaiiaeoe aeaeiaee neeaae neiaio?iiii? oei?e aeine?aeaeoaaii?
oa?eoi???;

i?iaanoe nenoaiaoe/iee, aaia?ao?/iee, a?iaeieia?/iee aiae?c;

anoaiiaeoe /an, niiniae i?a?aoe?? oa nooi?iue iaoo?ae?caoe??
aaeaaioeaieo ?ineei, ui c?inoathoue a ?aeii? aeine?aeaeaiiy;

aeyaeoe caeiiii??iino? aeioiiieia?/ii? i?eo?i/aiino? neiaio?iiieo
?ineei aei iaaieo o?ainoi?iiaaieo aeioii?a;

neeanoe eiiniaeo neiaio?iiii? oei?e ia iniia? ?acoeueoao?a aeanieo
aeine?aeaeaiue, aa?aa?ieo iaoa??ae?a oa e?oa?aoo?ieo a?aeiiinoae;

?ic?iaeoe i?iaiic iiaeeeaeo ci?i ?ineeiiiai iie?eao i?ae aio?iiiaaiiei
aieeaii.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Aia?oa anoaiiaeaii aeaeiaee
neeaae neiaio?iiii? oei?e aeine?aeaeoaaii? oa?eoi??? oa neeaaeaii ??
e?eoe/iee eiiniaeo. Iiaeaii ?icai?iooo oa?aeoa?enoeeo eiaeiiai aeaeo.

Aeyaeai? iia? aeey oei?e cao?aeieo iaeanoae Oe?a?ie i?noeaciaoiaeaeaiiy
iecee aaeaaioeaieo aeae?a. Cie?aia, aia?oa a ?aeii? aeine?aeaeaiiy
ciaeaeaii ea?aioeii? aeaee Ambrosia artemisiifolia L., Solanum cornutum
Lam., Centaurea diffusa Lam.

Aia?oa i?iaaaeaii anaa?/iee aiae?c neiaio?iiii? oei?e ?aeiio
aeine?aeaeaiue oa aeyaeaii ?? no?oeoo?i? iniaeeaino?.

Aena?oeaii ?noi??th iiyae ? ?icnaeaiiy iecee aaeaaioeaieo ?ineei ia
oa?eoi??? Oe?a?ie oa neeaaeaii ea?oe ?o a?aae?a.

Aeyaeaii aeioiiieia?/io i?eo?i/ai?noue neiaio?iiieo ?ineei aei
o?ainoi?iiaaieo aeioii?a.

Neeaaeaii i?iaiic iiaeeeaeo ci?i oei?e i?ae aieeaii ainiiaea?nueei?
ae?yeueiino?.

I?aeoe/ia cia/aiiy iaea?aeaieo ?acoeueoao?a. I?aeoe/ia cia/aiiy iathoue
iaoa??aee c aeaeiaiai neeaaeo neiaio?iiii? oei?e, ye? iiaeia aoaea
aeei?enoaoe i?e i?aeaioiaoe? ??cieo oei?enoe/ieo, oi?ieia?/ieo aai
ocaaaeueithth/eo i?aoeue, i?enay/aieo neiaio?iii?e oei?? Oe?a?ie aai
?a?iie.

Iaoa??aee aeena?oaoe?? ? iaeiei c aeaea?ae anoaiiaeaiiy no/anieo
oaiaeaioe?e ?icaeoeo oei?e ia oa?eoi??? Oe?a?ie i?ae aieeaii aio?ii?/ieo
oaeoi??a.

Aeyaeaii a?aeiinii no?ee? aei ??cieo oi?i ? o?eaaeino? aieeao
aio?ii?/ieo i?an–oaeoi??a aeaee, ye? iiaeooue aooe a?i?iaeeeaoi?aie
caaaeueieo ci?i aeiae?eey, i?eaeaoi? aeey aeei?enoaiiy i?e i?iaaaeaii?
aeieia?/ii? aenia?oece ? iii?oi?eiao caaaeueiiai noaio ?ineeiiiai
iie?eao.

Iaoa??aee aeena?oaoe?? aeei?enoiaothoueny a iaa/aeueiiio i?ioean?: i?ae
/an i?iaaaeaiiy eaai?aoi?ieo caiyoue, iieueiaeo i?aeoee, aeeiiaiiy
eo?niaeo ?ia?o.

Iniaenoee aianie caeiaoaa/a. Ia an?o aoaiao ?iaioa aeeiioaaeany
aeena?oaioii naiino?eii ?c canoinoaaiiyi a?aeiia?aeieo iaoiaeee.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. ?acoeueoaoe aeine?aeaeaiue
aeeeaaeaii ia iaoeiaeo eiioa?aioe?yo «Oai??y ? i?aeoeea aea/aiiy
i?e?iaeie/eo aeenoeeie?i o aeu?e oa na?aaei?e oeie?» (Oiaiue, 1997);
«I?aeia?iaei? aniaeoe aea/aiiy oa ioi?iie a?i??ciiiai?ooy Ea?iao»
(?ao?a, 1997); «Ieoaiiy a?i?iaeeeaoe?? oa aeieia??» (Caii??aeaey, 1998);
«I?iieneiaa aioai?ea» (Aeiiaoeuee, 1998); «Iaaaoeinuee? /eoaiiy» (Ee?a,
1998) oa iaoeiai-i?aeoe/ieo eiioa?aioe?yo aeeeaaea/?a IIO ?i. I.I.
Ae?aaiiaiiaa. E??i oiai, ?acoeueoaoe aeena?oaoe?eiiai aeine?aeaeaiiy
aeei?enoaii i?ae /an i?iaaaeaiiy caiyoue oa niaoeeo?n?a c? nooaeaioaie
i?e?iaeie/i–aaia?ao?/iiai oaeoeueoaoo IIO ?i. I.I.Ae?aaiiaiiaa.

Ioae?eaoe??. Iniiai? ?acoeueoaoe aeena?oaoe?? a?aeia?aaeai? o 8
ae?oeiaaieo iaoeiaeo i?aoeyo.

No?oeoo?a oa ianya ?iaioe. Aeena?oaoe?y aeeeaaeaia ia 201 noi??ieao
iaoeiiieniiai oaenoo, i?noeoue 6 oaaeeoeue, 33 ?enoiee. Neeaaea?oueny c?
anooio, 7 ?icae?e?a, aeniiae?a, nieneo e?oa?aoo?e. A?ae?ia?ao?y aeeth/a?
187 i?aoeue, cie?aia 43 – ?ioiiiai? aeaea?aea.

CI?NO ?IAIOE

?ICAe?E ?

O?ceei–aaia?ao?/ia oa?aeoa?enoeea

?aeiio aeine?aeaeaiiy

?aeiiii aeine?aeaeaiiy aoea i?aaiaa?aaeia /anoeia aa?oiuei? oa/??
aanaeio ?. Aei?noa?. Eiai ieiua noaiiaeoue aeecueei 1600 ei2.
I?ai?/ii–cao?aeia iaaea ?aa?iio ca?aa?oueny c ?oneii aa?o?a’y Aei?no?a,
i?aaeaiia – c ieaeiueith nioaith a??nueeiai e?niaiai iiyno, a no?aeia –
i?ioiaeeoue ii ?oneo ?. No?ee.

Ia’?eoii aeine?aeaeaiiy aoea neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio
Aei?no?a. Oei?o aeine?aeaeoaaee o 1994-1998 ??., i?ae /an
ia?o?ooii-oei?enoe/iiai ianoaaeaiiy oa?eoi???. A iniiao iieeaaeaii
ii??aiyeueiee ii?oieiai-aeieiai-aaia?ao?/iee iaoiae aeine?aeaeaiiy.
Aeey ii??aiyiiy aeei?enoaii aa?aa?i? iaoa??aee c aa?aa???a
Euea?anueeiai oi?aa?neoaoo ?i. ?.O?aiea (LW), ?inoeoooo aeieia?? Ea?iao
IAI Oe?a?ie (LWE) oa ?inoeoooo aioai?ee ?i. I.A.Oieiaeiiai IAI Oe?a?ie
(KW), Ae?iaiaeoeueeiai oa No?eenueeiai e?a?ciaa/eo ioca?a, a oaeiae
iiene aeae?a, iioae?eiaai? o 1866–1996 ??. Eaoeinuee? iacae oaenii?a
ooi/iaii ca N.E. *a?aiaiiaei.

Ia i?aenoaa? aiae?co o?ceei-aaia?ao?/ieo oiia ?aeiio aeine?aeaeaiiy
anoaiiaeaii, ui aiie ? aaoa?iaaiieie oa cia/ii ci?iaieie i?ae aieeaii
ethaeeie.

?ICAe?E 2

Ia?en oei?iaaiaco oa no/aniiai noaio

?ineeiiiai iie?eao

Ieoaiiyi oei?iaaiaco ia oa?eoi??? cao?aeieo iaeanoae Oe?a?ie caeiaeany
iecea aeine?aeiee?a (Uae?ia, ?eue?inueea [134, 43]; Nya?ye, Uae?ia [93];
Oaa?aaa [125]; N??aiei, Oo?ei [83]; Oaoa? [174], A?othoaiei oa ?i. [7];
Aaconueei, Ee?iaiia [8]). Anoaiiaeaii, ui a?ae aeaaiueiai aieioeaio aei
iaoiai /ano ci?ithaaany ee?iao, ui, a naith /a?ao, ni?e/eithaaei aei
ci?ie aeaeiaiai neeaaeo oei?e ? ?ineeiiino?. Aiane?aeie oeeo
aaoa?io?iiieo i?ioean?a o oei?? ?aa?iio aeyaeaii i?aaenoaaiee?a eiaeii?
c aiio.

A oe?eiio i?e?iaeiee ?ineeiiee iie?ea ?aeiio aeine?aeaeaiue aeineoue
iaiaeii??aeiee, ooo aeae?ey?oueny e?eueea oei?a i?e?iaeii? ?ineeiiino?,
a naia: oe?ieieenoyi? oa i?oai? e?ne, eoee ? aieioa, ui iathoue nai?
oa?aeoa?i? oei?enoe/i? iniaeeaino?. I?ioean oi?ioaaiiy neiaio?iiii?
oei?e o?nii iia’ycaiee c? ci?iaie i?e?iaeii? oei?e, ai cia/ia
e?euee?noue aaeaaioeaieo aeae?a aaeaioo?oueny aei o?ainoi?iiaaieo
aeioii?a.

A?aeiia?aeii aei oei?enoe/iiai ?aeiioaaiiy Oe?a?inueeeo Ea?iao, [120],
yei? ie aeio?eio?iiny, ?aeii aeine?aeaeaiiy ?icoaoiaaiee a iaaeao
No?aeiiea?iaonueei? i?aei?ia?ioe?? I?eea?iaonueeiai oei?enoe/iiai
?aeiio.

?ICAe?E 3

Ia?en ?noi??? aea/aiiy oei?e ?aeiio aeine?aeaeaiue

A ?noi??? aea/aiiy oei?e ?aeiio aeine?aeaeaiue caeaaeii a?ae ?noi?e/ieo
iiae?e, ie aeae?ey?ii o?e ia??iaee, a ye? aeine?aeieoeueea ?iaioa aoea
iaea?eueo aeoeaiith. O ia?oee ia??iae (O?O no.) oei?o aeine?aeaeoaaee
ia?aaaaeii aano?i-oai?nuee? aeine?aeieee. O ae?oaiio ia??iae? (ii/aoie
OO no.) aaeeeo ?iaioo a oeueiio iai?yieo i?iaiaeeee iieuenuee? a/ai?, ai
aei 1939 ?. oey oa?eoi??y aoea aaei?i?no?aoeaii i?aeii?yaeeiaaia
Iieueu?. Inoaii?e ia??iae ii/eia?oueny o i?neyai?ii? 40-ia? ?iee OO
no. ? o?eaa? aei oeueiai /ano.

O ?iaio? i?iaaaeaii aiae?c iaeiioe?ai?oeo eeaneo?eaoe?e neiaio?iiieo
aeae?a ?ineei, ?ic?iaeaieo A. Oaeeoiaii [176, 177], I.N.Eaieoaaei [46],
E. Aieoaii, A. ??anaeii [149], ss. Ei?ianai [156, 157], O. O?aaea?ii
[169], ?.A. Aei?iainoaenueeith [36, 37], A.N. ssa?iaa–Eieaeianueeith
[138], I.A. A’theiaith [24], A.A. *?/aaei [116], A.A.I?ioiiiiiaith [77,
79] oa ?ioeie iaoeiaoeyie. I?ioa ia aeaiee iiiaio iaia?
caaaeueiii?eeiyoi? eeaneo?eaoe??, yea a a?aeiia?aeaea iao? eiiieaenii?
ioe?iee oei?e.

Caaaeoth/e ia aenieee nooi?iue aio?iiiaai?caoe?? eaiaeoaooo, aeey
aiio?oii? o?aeoe?? ie i?iiiio?ii aeei?enoaoe eeaneo?eaoe?th
A.A.I?ioiiiiiai? [79], a iniiao yei? iieeaaeaii iiaeeea?noue oa nooi?iue
aaeaioaoe?? ?ineei aei o?ainoi?iiaaieo aeioii?a, a aeey aaeaaioeaii? –
nenoaio O. O?aaea?a [169], aeaui iiaeeo?eiaaio a?aeiia?aeii aei
caaaeaiue aeine?aeaeaiiy. Oiio ie i?iiiio?ii naith, aeaui iiaeeo?eiaaio
eeaneo?eaoe?th. I?e oeueiio, ii iiaeeeaino?, ie aeei?enoaee aaea
?nioth/? oa?i?ie.

Neiaio?iie

Aaeaaioe Aiio?oe

A. Ca /anii ?i?a?aoe??: A. Nooi?iue aaeaioaoe??

1) a?oaio?oe 1) aaaiio?oe

2) eaiio?oe 2) aai?aiio?oe

A. Ca niiniaaie ?i?a?aoe??: 3) iano?ee? aiio?oe

1) enaiio?oe

2) a?aac?io?oe

A.Ca nooiaiai iaoo?ae?caoe??:

1) aa??io?oe

2) aiaeio?oe

3) aoaia?io?oe

4) eieiiio?oe

?en. 3.1. Nenoaia eeaneo?eaoe?? neiaio?iiii? oei?e

?icae?e 4

Eiiniaeo neiaio?iiii? oei?e

Eiiniaeo neiaio?iiii? oei?e aeine?aeaeoaaiiai ?aeiio neeaaeaiee ca
?acoeueoaoaie aeanieo o?e??/ieo iieueiaeo aeine?aeaeaiue oa ia iniia?
ii?aoethaaiiy aa?aa?ieo eieaeoe?e Ae?iaiaeoeueeiai e?a?ciaa/iai iocath,
Euea?anueeiai oi?aa?neoaoo ?i. ?. O?aiea (LW), ?inoeoooo aeieia?? Ea?iao
IAI Oe?a?ie (LWE), ?inoeoooo aioai?ee ?i. I. A. Oieiaeiiai IAI Oe?a?ie
(KW) ? e?oa?aoo?ieo aeaiieo.

Aei «Eiiniaeoo…» aeeth/aii 365 aeae?a c 54 ?iaeei oa 213 ?iae?a,
aeyaeaieo iaie ia oa?eoi??? aeine?aeaeaiiy. Nthaee aeeth/aia ? cia/ia
e?euee?noue eoeueoo?ieo ?ineei, ye? aee/aa?thoue. Iiaeaii ?icai?iooo
oa?aeoa?enoeeo eiaeiiai aeaeo.

I?ae /an iieueiaeo aeine?aeaeaiue iaie aoee aeyaeai?
i?noeaciaoiaeaeaiiy oanoe aeae?a aaeaaioeaieo ?ineei, iiaeo aeey
cao?aeieo iaeanoae Oe?a?ie, cie?aia: Heracleum mantegazzianum S. et L.,
Lactuca tatarica (L.) C.A.Mey., Commelina communis L., Solanum cornutum
Lam., Centaurea diffusa Lam., Physalis ixocarpa Brot. ex Hornem.

E??i oiai, aeyaeaii iia? i?noeaciaoiaeaeaiiy oaeeo aaeaaioeaieo aeae?a,
ye Veronica arvensis L., Rudbeckia laciniata L., Ipomaea purpurea (L.)
Roth., Echinocystis lobata (Michx.) Torr. ex Gray, Tragopogon dubium
Scop., Verbena officinalis L., Bryonia dioica Jacq., Ambrosia
artemisiifolia L.oa ?i.

?ICAe?E 5

AIAE?C NEIAIO?IIII? OEI?E

Ca ?acoeueoaoaie aeine?aeaeaiue i?iaaaeaii aiae?c nenoaiaoe/ii?,
aaia?ao?/ii?, aeieia?/ii?, a?iii?oieia?/ii? no?oeoo?e oa aeeoo?aeo oi?i
aeae?a, iaaaaeaieo o «Eiiniaeo?…».

Aeyaeaii oae? iniaeeaino? nenoaiaoe/ii? no?oeoo?e neiaio?iiii? oei?e
?aa?iio aeine?aeaeaiue. Caaaeueiee aeaeiaee neeaae noaiiaeoue 365
aeae?a c 54 ?iaeei oa 213 ?iae?a. Aeanyoue i?ia?aeieo ?iaeei
aeeth/athoue 240 aeae?a (66%) oei?e ?aeiio. Iaea?eueoo e?euee?noue
aeae?a aeeth/athoue ?iaeeie Asteraceae (63 aeaee, 17,2%), Poaceae (32,
8,49%), Fabaceae (27, 7,39%), Brassicaceae (23, 6,30%), Scrophulariaceae
(19, 5,20%), Caryophyllaceae (18, 4,93%), Lamiaceae (17, 4,65%),
Rosaceae (16, 4,65%), Chenopodiaceae (13, 3,56%), Polygonaceae (12
aeae?a, 3,28%). Aei ianooiieo aeanyoe ?iaeei iaeaaeeoue 70 aeae?a
(19%), ui oo?e/? iaioa a?ae iiia?aaeiueiai iieacieea. A na?aaeiueiio ia
iaeio ?iaeeio i?eiaaea? n?i aeae?a, a?eueoa iieiaeie ?iaeei (34 aai
62%) i?aaenoaaeai? iaeiei – o?ueiia aeaeaie. Iecueea aeaeiaa
iane/ai?noth, na?ae/eoue i?ioa, ui oei?a oa?eoi??e ?icaeaa?oueny a
aeno?aiaeueieo oiiaao.

O ?iaeiaiio niaeo?? neiaio?iiii? oei?e ia?aaaaeathoue oae? ?iaee:
Veronica – 9 aeae?a, Vicia oa Polygonum ii 6, Malva, Chenopodium,
Geranium, Euphorbia oa Potentilla ii 5, Poa, Trifolium oa Atriplex ii 4.
Aeniea iane/ai?noue aeaeaie ?iaeo Veronica ? iciaeith ena?io?eueiino?,
oa?aeoa?ii? aeey na?aaecaiiiii?nueei? oei?e.

Iniiai? e?euee?ni? iieacieee nenoaiaoe/iiai niaeo?o na?ae/aoue i?i oa,
ui i?ae aieeaii ae?yeueiino? ethaeeie a?aeaoaa?oueny o?ainoi?iaoe?y
oei?e oa ca?eueoaiiy ?ie? ?iaeei ? ?iae?a, oa?aeoa?ieo aeey ena?e/ieo
iaeanoae Na?aaecaiiiii?’y.

Aaia?ao?/iee aiae?c aeaii? oei?e na?ae/eoue i?i oa, ui ca oeiii a?aaeo
neiaio?iii? aeaee ?ineei ?aeiio aeine?aeaeaiue iaeaaeaoue aei 26 a?oi.
Iaeoe?oa i?aaenoaaeaieie a?aaeia?/ieie a?oiaie ? aiea?eoe/ia – 107
aeae?a, ?a?iac?enueea – 64, a?oie einiiiie?o?a – 50 oa aai?einiiiie?o?a
– 35, ?a?iiaenueei–na?aaecaiiiii?nueea – 27,
?a?iiaenueei–na?aaecaiiiii?nueei–no?aeiiac?aonueea – 15,
?a?iiaenueei–i?ai?/iiaia?eeainueea – 9 aeae?a. ?io? a?oie i?aaenoaaeai?
1-3 aeaeaie.

*ioe?e oeie a?aae?a (aiea?eoe/iee, ?a?iac?enueeee, einiiiie?oe oa
aai?einiiiie?oe), ia?aaaaeathoue o no?oeoo?? neiaio?iiii? oei?e
aa?oiuei? oa/?? aanaeio Aei?no?a. Cia/ia i?noea na?aae aiio?o?a
caeiathoue oe?ieia?aaeuei? aiea?eoe/i? oa ?a?iac?enuee? aeaee ?c
iacio?eueieo oa?oiiaaiue. Na?aae aaeaaioeaieo aeae?a ia?aaaaeathoue
einiiiie?oe oa aai?einiiiie?oe, ye? ? i?aaenoaaieeaie ena?io?eueieo
oa?oiiaaiue.

O neiaio?iii?e oei?? ?aa?iio aeine?aeaeaiiy ie aeyaeee 178 aeae?a
aaeaaioeaieo ?ineei, ye? iaeaaeaoue aei 127 ?iae?a oa 42 ?iaeei. Ca
?acoeueoaoaie oei?iaaiaoe/iiai aiae?co aeo?aeieo a?aae?a, iniiaieie
eiiiiiaioaie neiaio?iiii? oei?e aeine?aeaeoaaiiai ?aa?iio ?
i?aaenoaaieee Aeaaiueiai Na?aaecaiiiii?’y – 117 aeae?a (32%), cia/iith
? e?euee?noue ?ineeie, ui iathoue i?ai?/iiaia?eeainueea iioiaeaeaiiy
(25, 7,1%). *eneaiio a?oio – 15 aeae?a (4,1%) – o oei?? ?aeiio
neeaaeathoue aeaee ?a?iiaenueeiai iioiaeaeaiiy. C Ac?? oa I?aaeaiii?
Aia?eee iioiaeeoue ii 6 aeae?a (1,6%). ?io? aeo?aei? a?aaee o oei??
?aeiio i?aaenoaaeai? 1-3 aeaeaie. I?ioa /a?ac iaaeinoaoi?noue
iaiao?aeieo iaoa??ae?a iai ia aaeaeiny anoaiiaeoe iioiaeaeaiiy 8 aeae?a.

Aeey aiae?co aaeaaioeaii? o?aeoe?? ie aeei?enoaee o?e iacaeaaei? iaeei
a?ae iaeiiai iieacieee: /an ?i?a?aoe??, niin?a ?i?a?aoe?? oa nooi?iue
iaoo?ae?caoe??, ye? cai?iiiiiaai? O. O?aaea?ii [169] oa
aeei?enoiaothoueny aaaaoueia aeine?aeieeaie. Aei a?oie a?oaio?o?a ie
a?aeianee 92 aeaee (25,2%), eaiio?o?a – 86 aeae?a (23,5%) ?ineei.
Aaeeeo ?ieue o iioe?aii? aaeaaioeaieo ?ineei a?ae?a?a? niin?a
?i?a?aoe??. Caeaaeii a?ae niiniao ?i?a?aoe?? ie aecia/eee aea? a?oie:
ia?aaaaeio a?eueo?noue (37,2%) o ?aeii? noaiiaeyoue enaiio?oe, ia aeieth
a?aac?io?o?a i?eiaaea? 11,8%. Caeaaeii a?ae nooiaiy iaoo?ae?caoe??
aaeaaioeai? ?ineeie, aeyaeai? o ?aa?ii?, iiae?eaii ia /ioe?e a?oie:
aa??io?oe – 11 aeae?a (3%), aiaeio?oe – 145 (39,7%), aoaia?io?oe – 15
(4,1%) oa eieiiio?oe – 7 aeae?a (1,9%).

A aeieia?/iiio niaeo?? neiaio?iiii? oei?e ca a?aeiioaiiyi aei aieiae ia
ia?oiio i?noe? ? a?oia ena?iiacio?o?a – 178 aeae?a (48,7%), ia ae?oaiio
– iacio?o?a 143 aeaee (39,4%), o?ao? i?noea caeiathoue iaciena?io?o?a
– 23 aeaee (6,3%). Iaeiaio /eneaiieie ? a?oie a?a?io?o?a, ena?io?o?a oa
a?ae?io?o?a – a?aeiia?aeii, 9 (2,5%), 8 (2,1%) oa 4 aeaee (1%). Aeniea
/anoea ena?iiacio?o?a a aeieia?/iiio niaeo?? na?ae/eoue i?i
a?eaeecaoe?th oa?eoi??? oa ena?io?oecaoe?th ?ineeiiiai iie?eao i?ae
aieeaii aio?ii?/iiai oaeoi?a.

Ca nooiaiai i?enoinoaaiiy aei ?ioaineaiino? ina?oeaiiy iae/eneaii?oith ?
a?oia aae?io?o?a – 240 aeae?a (65,8%), ae?oaa i?noea caeiathoue
noe?iaae?io?oe – 118 aeae?a (32,3%), o?ao? – noe?io?oe – 7 aeae?a
(1,9%).

Ia iniia? aiae?co aeeoo?aeo oi?i anoaiiaeaii, ui aei oa?io?o?a o ?aeii?
aeine?aeaeaiiy iaeaaeaoue 185 aeae?a (50,8%). Ia ae?oaiio i?noe?
ciaoiaeyoueny aai?e?eioio?oe – 157 aeae?a (43%), a?oia oaia?io?o?a
iae?/o? 12 aeae?a (3,3%), oaiao?o?a – 7 aeae?a (1,9%) ? 4 aeaee (1%)
iaeaaeaoue aei a?oie e?eioio?o?a. Oaeei /eiii, aeii?ioth/ith aeeoo?aith
oi?iith o neiaio?iii?e oei?? ?aeiio ? oa?io?oe, ui iiynith?oueny ?o
aenieith aaeaioaoe??th aei aio?iiiaaiieo aeioii?a.

I?ae /an aeine?aeaeaiue a?iii?oieia?/ii? no?oeoo?e neiaio?iiieo aeae?a,
aeyaeaii, ui ia?aaaaeathoue o?aa’yieno? ?ineeie – 346 aeae?a, aai 94,8%
a?ae caaaeueii? ?o e?eueeino?. Aea?aaa oa eou? ia/eneaii? – 6 oa 13
aeae?a a?aeiia?aeii, aai 1,6 oa 3,6%, oea ia?aaaaeii caee/aa?e?
eoeueoeaiaai? aeaee ?ineei.

Aiae?c niinia?a iioe?aiiy neiaio?iiieo ?ineei na?ae/eoue i?i oa, ui 156
aeae?a iaeaaeaoue aei aiaiioi?iiai oeio. Aaeeeo ?ieue o ia?aianaii?
ae?anii? a?ae?a?athoue ? oaa?eie, ai o aeine?aeaeai?e oei?? aeyaeaii 150
ciioi??a. Aio?iiioi??a iae?/o?oueny 80 aeae?a. O 89 aeae?a a?aecia/aia
aeniea caeaoi?noue aei iie?oi???. Cia/ia e?euee?noue – 66 aeae?a, ca
oa?aeoa?ii iioe?aiiy ? aa?ioi?aie. Iaeiaio i?aaenoaaeai? o neiaio?iii?e
oei?? aaoioi?e – 10 aeae?a. Niin?a iioe?aiiy a?eueoino? ?ineei
aecia/a?oueny iayai?noth (a?aenooi?noth) ia?aoeiae o eaiaeoaoo? oa
aieeaii ethaeeie.

Ia iniia? aeanieo niinoa?aaeaiue e?euee?niiai (ia’?iiiai) iioe?aiiy
ie?aieo aeae?a a iaaeao ?aeiio o ?ineeiiiio iie?ea? aeyaeaii oae?
caeiiii??iino?: cae/aeii (ianiai) iioe?aii ii an?e oa?eoi??? 72 aeaee
(19,7%), /anoi – 145 (40%), nii?aaee/ii – 74 (20,2%), c??aeea — 43
(11,7), aeoaea ??aeei — 31 (8,4%).

?ICAe?E 6

?noi??y oi?ioaaiiy oa aeaa?aaeaoe?? ?ineeiiiai iie?eao i?ae ae??th
aio?ii?/iiai oaeoi?A

Aeey a?aeoai?aiiy ?noi??? iiyae oa ?icnaeaiiy aaeaaioeaieo ?ineei ia
oa?eoi??? ?aa?iio aeine?aeaeaiue ie i?iaaee ii??aiyeueiee aiae?c
oei?enoe/iiai neeaaeo ?aa?iio ca inoaii? 150 ?ie?a.

Ia?o? a?oioiai? oei?enoe/i? aeine?aeaeaiiy aa?oiuei? oa/?? aanaeio
Aei?no?a i?iaiaeeeeny o 1850-1870 ??. oai?nueeeie oa iieuenueeeie
aeine?aeieeaie. Iaeiiai?o? a?aeiiino? i?i oei?o ?aa?iio ciaoiaeeii o
i?aoe? A. Atheeaey [150], yeee aeey ?aa?iio aeine?aeaeaiue iaaiaeeoue
988 aeae?a, cie?aia – 97 aaeaaioeaieo.

1898 ?ieo i?io. I. Eioethaith [155] iioae?eiaaiee aeiaeaoie
aei oei?e Ae?iaiae/a ca cai?aie A. Oo?/einueeiai, ui i?noeoue 24 iiaeo
aaeaaioeaieo aeaee.

Ia ii/aoeo OO no. o oei?? I?eea?iaooy iia? aaeaaioeai? aeaee
aoee aeyaeai? iieuenueeeie aeine?aeieeaie.

O OO no. oe?a?inuee? aeine?aeieee ia oa?eoi??? I?eea?iaooy ciaeoee ua
ieceo iiaeo aaeaaioeaieo aeae?a ?ineei [30, 39, 52, 89, 99, 120, 126,
130 oa ?i.].

Aiae?c e?oa?aoo?ieo aeaea?ae, ui i?noyoue ?acoeueoaoe ?iaaioa?ecaoe??
oei?e, oa aeani? aeine?aeaeaiiy aeicaieeee iai a?aeoai?eoe ?noi??th
iiyae oa ?icnaeaiiy ?yaeo aaeaaioeaieo ?ineei, ui o?aieythoueny a
?aa?ii? aeine?aeaeaiue cae/aeii (ianiai) (Amaranthus retroflexus L.,
Galinsoga parviflora Cav., G. ciliata (Rafin.) Blake, Chamomilla
suaveolens (Putsh) Rydb., Erigeron canadensis L., Juncus tenuis Willd.,
Impatiens parviflora DC.) aai c??aeea (Ambrosia artemisiifolia, Carduus
nutans L., Commelina communis, Centaurea diffusa, Cyclachaena
xanthifolia (Nutt.) Fresen., Echinocystis lobata, Impatiens glandulifera
Royle, Solanum cornutum, Xanthoxalis dillenii (Jacq.) Holub oa ?i.).

Oae, ?acii c ian?iiei iaoa??aeii 1886 ?ieo aei ?a?iie c I?ai?/ii?
Aia?eee aeiaaeeiai iio?aiea Solanum cornutum Lam. A Oe?a?i? aeae
aeyaeaii 1928 ?ieo Ae.E. Ea??iiiaei o iin?aao ca?iiaeo eoeueoo? ia
Ee?auei?. O i?neyai?ii? ?iee nii?aaee/ii cono??/aany a e?ninoaiiaeo oa
noaiiaeo ?aeiiao Oe?a?ie [110]. C?inoa? ia?aaaaeii ia aaooaiao, na?aae
ethoea?ie, o naaeao, ia aeiia?aaeieeao, a?ey aoae?aaeue. Ca 40 ?ie?a
cia/i? eieii?? noi?ioaaeeny o E??iaia?aaenuee?e, Aei?i?iiao?ianuee?e,
Caii??cuee?e, Iaeanuee?e, Ieeiea?anuee?e, Oa?niinuee?e, Eoaainuee?e oa
Aeiiaoeuee?e iaeanoyo. Nii?aaee/ii c?inoa? o noaiia?e cii? E?eio oa
A?iieoeuee?e iaeano? [17]. Iei? aeae i?iaeiaaeo? ?icnaeyoueny o
i?ai?/ii-cao?aei? ?aeiie E?ninoaio. Ie aia?oa aeyaeee ia cae?cie/i?e
eie?? o i.No?ee (Euea?anueea iae.).

Solanum cornutum – eaiio?o i?ai?/iiaia?eeainueeiai iioiaeaeaiiy c
?a?iiaenueei–na?aaecaiiiii?nueei–i?ai?/iiaia?eeainue-eei a?aaeii. Ia
oa?eoi??? Oe?a?ie i?ai?/ia iaaea eiai iioe?aiiy i?ioiaeeoue /a?ac
noaiiao ciio oa E?ei. E??i oiai, o No?aei?e ?a?ii? iioe?aiee o
Ieaeiueiaiecueeiio ?aeii?.

Aeae c?inoa? a iin?aao, naaeao, ia ai?iaeao oa acaeiaae cae?cieoeue.

Caaaeueia iioe?aiiy: Iaeneea ? i?aaeaiii-cao?aeia /anoeia NOA, a oaeiae
Na?aaeiy ?a?iia oa Na?aaecaiiiii?(y [109].

sse aa/eii, aio?iiiaai?caoe?y oa?eoi??? ni?ey? iioe?aiith aaaaoueio
aaeaaioeaieo aeae?a ?ineei, ianaiia?aae oeo, ui iathoue aenieo
aeieia?/io ieanoe/i?noue oa aeeoo?a?noue. A aonoa ia?aaea oeyo?a
niieo/aiiy a ?aeii? aeine?aeaeaiiy ni?ey? ooai?aiith i?a?aoe?eieo
ei?eaei??a aeey aaeaaio?a oa ?o iioe?aiith a iia? ?aa?iie.

Neeaae ? no?oeoo?a niaoeeo?/ieo oei?a i?noeac?inoaiue aecia/athoueny
oiiaaie iaaiiai aeioiio ? a cia/i?e i??? a?ae??ciythoueny iioiaeaeaiiyi
? iai?yieii ?icaeoeo i?ae aieeaii aio?ii?/ieo oaeoi??a. Oeiiaeie oi?iaie
aio?iiiaai?ciaaiiai eaiaeoaooo ? aa?ioeaiice, aaoiiia?euei? oa
cae?cie/i? oeyoe, ni?oieee, aoeeoe?, aaciie, eeoiae, aoae?aaeuei?
iaeaeai/eee oa naeeoaai? oa?eoi??? – oiaoi ae?eyiee, ia yeeo i?e?iaeiee
iie?ea ?noioii ii?ooaiee aai iiai?noth cieuaiee. ?o aeaeiaee neeaae i?e
an?e ??ciiiai?oiino? ia? aaaaoi ni?eueiiai. Oea iiynith?oueny oe?ieith
aeieia?/iith ieanoe/i?noth a?eueoino? ao?’yi?a, ianaiia?aae
aaeaaioeaieo, ? oei, ui a o?aai?ciaaiiio na?aaeiaeu? niaeo? aeieia?/ieo
oaeoi??a cia/ii aoae/ee, i?ae o i?e?iaeieo oiiaao. A i?ioean?
i?e?iaaiethoe?? ia o?ainoi?iiaaieo aeioiiao, iiae?aieo ca aaeao?/ieie
oiiaaie oa aeaeii aio?ii?/iiai aieeao, oi?iothoueny aeoaea aeecuee? ca
neeaaeii ?ineeii? oa?oiiaaiiy, ye? iiaeooue aooe a?i?iaeeeaoi?aie ci?i o
aeiae?ee?.

?ICAe?E 7

I?iaiic iiaeeeaeo ci?i ?ineeiiiai iie?eao i?ae aieeaii aio?iiiaaiieo
oaeoi??a

Ia iniia? i?iaaaeaieo aiae?c?a oa aeyaeaieo iniaeeainoae
neiaio?iiii? oei?e ie ?ic?iaeee i?iaiic iiaeeeaeo ci?i ?ineeiiiai
iie?eao i?ae ae??th aio?ii?/ieo oaeoi??a.

Aiane?aeie neiaio?ii?caoe?? ?aa?iiaeueia oei?a a iiaeaeueoiio
caaaaoeoueny ia aaeaaioeai? aeaee, ui iaoiiaeoue ci?io nenoaiaoe/ii?,
aaia?ao?/ii?, aeieia?/ii? no?oeoo?e oa niaeo?a aeeoo?aeo oi?i.
?icoe?youeny oei?iaaiaoe/i? ca’ycee oei?e ?aa?iio aeine?aeaeaiiy c
oei?ith ena?e/ieo iaeanoae ?a?ac?? oa ?ioeo eiioeiaio?a.

Ia iniia? aiae?co niaeo?a nenoaiaoe/ii? no?oeoo?e iiaeia ia?aaeaa/aoe,
ui e?euee?noue neiaio?iiieo aeae?a ia?aaaaeii aoaea ca?eueooaaoeny ca
?aooiie i?aaenoaaiee?a ?iaeei Asteraceae, Poaceae, Brassicaceae,
Chenopodiaceae, ai naia oe? ?iaeeie iaea?eueoa i?aaenoaaeai? a
?aa?iiaeuei?e oei??. Iineeaiiy ?ie? neiaio?iiieo aeae?a o iiaeaeueo?e
o?ainoi?iaoe?? oei?e ni?e/eith? nooo?a? ci?ie ia an?o ??aiyo –
iiioeyoe?eiiio, aeaeiaiio, oeaiioe/iiio oa aeinenoaiiiio.

Aaeaaioeaia o?aeoe?y iiaea iiiiaieoeny aeaeaie ?c ?iae?a Chenopodium,
Amaranthus, Malva, Veronica, Geranium, Euphorbia, ui aaea i?aaenoaaeai?
cia/iith e?euee?noth aeae?a o oei?? ?aa?iio. ?c aiio?oii? o?aeoe?? aei
neiaio?iiii? oei?e oa?eaeooue ?ineeie ?c ?iae?a Potentilla, Silene,
Polygonum, Rumex, Cirsium, Poa, Verbascum, ui iathoue aenieo aeieia?/io
ieanoe/i?noue ? eaaei aaeaioothoueny aei iiaeo oiia ?nioaaiiy.

A iaeaooiueiio niinoa??aaoeiaoueny oaiaeaioe?y aei ca?eueoaiiy o
oei?enoe/iiio niaeo?? ?aa?iio aae?io?o?a, ena?iiacio?o?a, oa?io?o?a.
Oae? ?ineeie iathoue a?eueoo aeieia?/io ieanoe/i?noue oa oaeaeoa
aaeaioothoueny aei ci?iaieo oiia.

AENIIAEE

Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio Aei?no?a iae?/o? 365
aeae?a, ui iaeaaeaoue aei 213 ?iae?a oa 54 ?iaeei. Aeey caoiaeo Oe?a?ie
aeyaeaii 6 iiaeo aeae?a.

A ?acoeueoao? aiae?co nenoaiaoe/ii? no?oeoo?e neiaio?iiii? oei?e
c’yniaai? ?? oa?aeoa?i? ?ene, ye? iieyaathoue a iiae?aiino? niaeo?a
i?ia?aeieo ?iaeei c aiaeia?/ieie oei?aie ?ioeo oa?eoi??e. Anoaiiaeaii,
ui a ?iaeiaiio niaeo?? ia?aaaaeathoue ?iaee c iaeith aeaeiaith
iane/ai?noth, ui oa?aeoa?ii aeey oei? oa?eoi??e c aeno?aiaeueieie
oiiaaie ?icaeoeo ?ineeiiiai iie?eao. A?aei?ii?noue a?ae niaeo?a
i?ia?aeieo ?iaeei neiaio?iiii? oei?e Oe?a?ie a?aeia?aaea? aeeeth/ii
aeniea iieiaeaiiy ?iaeeie Fabaceae – oeiiai? aeey Na?aaecaiiiii?nueei?
iaeano?, cia/io ?ieue ?iaeeie Chenopodiaceae, oa?aeoa?ii? aeey
ionoaeueieo oei?, a oaeiae aoiaeaeaiiy a ia?oo aeanyoeo ?iaeei Rosaceae
oa Polygonaceae, i?aaenoaaieee yeeo iioe?ai? a i?ai?/i?e iii??i?e
cii?.

Aiae?c a?iii?oieia?/ii? oa aeieia?/ii? no?oeoo?e neiaio?iiii? oei?e
na?ae/eoue i?i na?aaecaiiiii?nueeee oa?aeoa? oei?e ?aeiio
aeine?aeaeaiiy. Na?aae aeia?iii?o iaea?eueoa aae?io?o?a ?
iaciena?io?o?a, ui oa?aeoa?ii aeey ciie noai?a. Ia a?aei?io a?ae
?aa?iiaeueii? oei?e, o neiaio?iii?e oei?? ?aa?iio ca aaa?oonii oa
o?eaae?noth aeeooy ia?aaaaeathoue iiiiea?i?ee.

Iiaeaii eeaneo?eaoe?th neiaio?iiieo aeae?a ?ineei, ?ic?iaeaio ca
oi?i–oei?i–?noi?e/ieie e?eoa??yie ? iciaeaie iaoo?ae?caoe?? oa
aaeaioaoe??. Cai?iiiiiaaia nenoaia caaacia/o? aoaeoeai?oa i?iaaaeaiiy
aiae?co neiaio?iiii? oei?e.

Aeae?eaii aiio?oio oa aaeaaioeaio o?aeoe??. Ni?aa?aeiioaiiy i?ae ieie
noaiiaeoue 1:1, ui na?ae/eoue i?i cia/io ?ieue aaeaaioeaieo aeae?a o
oi?ioaaii? neiaio?iiii? oei?e, i?e/iio ca /anii i?iieeiaiiy
ia?aaaaeathoue a?oaio?oe, ca niiniaii ?i?a?aoe?? – enaiio?oe, a ca
nooiaiai iaoo?ae?caoe?? – aiaeio?oe. Iaoo?ae?caoe?y aaeaaioeaieo
aeae?a a?aeaoaa?oueny ia?aaaaeii a ?ineeiieo oa?oiiaaiiyo
o?ainoi?iiaaieo aeioii?a, a oie aea /an o i?e?iaeiiio ?ineeiiiio iie?ea?
/anoioa ?o iiyae ciaioo?oueny.

Aaia?ao?/ia no?oeoo?a aeine?aeaeoaaii? oei?e na?ae/eoue i?i oa, ui o
neeaae? neiaio?iiii? oei?e aa?oiuei? oa/?? aanaeio Aei?no?a
ia?aaaaeathoue /ioe?e oeie a?aae?a (aiea?eoe/iee, ?a?iac?aenueeee,
einiiiie?oe oa aai?einiiiie?oe). Cia/ia i?noea na?aae aiio?o?a
caeiathoue oe?ieia?aaeuei? aiea?eoe/i? oa ?a?iac?enuee? aeaee ?c
iacio?eueieo oa?oiiaaiue. Na?aae aaeaaioeaieo aeae?a ia?aaaaeathoue
einiiiie?oe oa aai?einiiiie?oe, ye? ? i?aaenoaaieeaie ena?io?eueieo
oa?oiiaaiue. Aiane?aeie neiaio?ii?caoe?? oei?a ?aa?iio caaaaoeeany
oei?enoe/ieie aeaiaioaie, aaia?ao?/ii iia’ycaieie c ena?e/ieie
oa?eoi??yie caiii? eoe?.

Oei?iaaiaoe/iee aiae?c aeo?aeieo a?aae?a na?ae/eoue i?i aenieo
iane/ai?noue neiaio?iiii? oei?e ?aa?iio aeine?aeaeaiiy aaeaaioeaieie
aeaeaie na?aaecaiiiii?nueeiai, ac?aonueeiai oa i?ai?/iiaia?eeainueeiai
iioiaeaeaiiy o ca’yceo c iiaioaaiiyi ieie iiaeo aio?iiiaaiii
ia?aoai?aieo aeioii?a.

Aena?oeaii ?noi??th iiyae oa iioe?aiiy ?yaeo aaeaaioeaieo ?ineei ia
oa?eoi??? Oe?a?ie. Ia iniia? neeaaeaieo ea?o a?aae?a aeyaeaii caaaeuei?
caeiiii??iino? i?a?aoe?? ia oa?eoi??? Oe?a?ie na?aaecaiiiii?nueeeo oa
ac?aonueeeo aeae?a a i?ai?/ii-cao?aeiiio iai?yieo.

?icaeyiooi oeiia? o?ainoi?iiaai? aeioiie, ui a?ae??ciythoueny ca
nooiaiai oa oa?aeoa?ii aio?iiiaaiiiai aieeao. Aeyaeaii a?aeiinii no?ee?
aeaee aei ??cieo oi?i ? o?eaaeino? aieeao aio?ii?/ieo i?an-oaeoi??a, ye?
iiaeooue a?ae?a?aaaoe ?ieue a?i?iaeeeaoi??a caaaeueieo ci?i o aeiae?ee?.

Ia i?aenoaa? oei?ieia?/ii? ioe?iee iane?aee?a neiaio?ii?caoe?? oa ?o
i?iya?a ia i?aai?ciiaiio, iiioeyoe?eiiio e aeinenoaiiiio ??aiyo
anoaiiaeaii, ui aiia a?aeaoaa?oueny a iai?yieao ci?ie no?oeoo?ii?
i?aai?caoe?? oei?e, aaeaioaoe?? iniaei e iiioeyoe?e aei ae??
aio?iiiaaiieo oaeoi??a ? ia?inoaiiy ?? aeeiai?/ieo oaiaeaioe?e.

Nienie iaoeiaeo i?aoeue, iioae?eiaaieo ca oaiith aeena?oaoe??

Neeaa TH.A. Neiaio?iiia oei?a aa?o?a’y aanaeio Aei?no?a oa i?iaeaie ??
eeaneo?eaoe?? // Iaoeia? caienee Oa?iii. aea?ae. iaae. oi-oo ?i.
Aieiaeeie?a Aiaothea. – Oa?iii?eue. – 1998.– 2 (4).– N.19-22.

Neeaa TH.A. Aeioiiieia?/ia i?eo?i/ai?noue neiaio?iiieo ?ineei // Iaoeia?
caienee Oa?i. aea?ae. iaae. oi-oo ?i. Aieiaeeie?a Aiaothea. –
Oa?iii?eue. – 1998. – 3 (4). – N.5-10.

Neeaa TH.A. ?iaac?y oa iaoo?ae?caoe?y aaeaaioeaieo ?ineei o aa?oi?e
oa/?? aanaeio Aei?no?a// Iaoeia? caienee Oa?i. aea?ae. iaae. oi-oo ?i.
Aieiaeeie?a Aiaothea. – Oa?iii?eue. – 1999. – 1(4). – N.7-11.

Neeaa TH.A. Iaeyae nenoai eeaneo?eaoe?? neiaio?iiii? oei?e oa i?iaeaie
?? aea/aiiy // Iaoeia? caienee: Ca??iee iaoeiaeo noaoae Iaoe. iaae.
oi-oo ?i. I.I. Ae?aaiiaiiaa – Ee?a: IIO, 1998.– Aei.2 – N. 165-172.

Neeaa TH.A. Aaeaaioeai? ?ineeie aa?oiuei? oa/?? aanaeio Aei?no?a
(Euea?anueea iae.) // I?iieneiaa aioai?ea: noai oa ia?niaeoeae
?icaeoeo (Iaoa??aee O?aouei? i?aeia?iaeii? eiioa?aioe??). – Aeiiaoeuee,
Aaainoai «Ioeueoei?an», 1998. – N.119-125

Neeaa TH.A., Iaaeai?iee A.A. No/aniee noai ?ineeiieo anioe?aoe?e oa
oaa?eiieo oa?oiiaaiue ia cae?cie/ieo aei?iaao I?eea?iaooy //
I?aeia?iaei? aniaeoe aea/aiiy oa ioi?iie a?i??ciiiai?ooy Ea?iao. –
?ao?a, 1997. – N. 188-191.

Iaaeai?iee A.A., Neeaa TH.A. Aeieia?/ia iiaeaeue i?noa — iaeia c oi?i
iiaeeaeaiiai aea/aiiy o/iyie oeeeeo i?e?iaeie/i-aaia?ao?/ieo aeenoeeie?i
// Oai??y ? i?aeoeea aea/aiiy i?e?iaeie/eo aeenoeeie?i o aeu?e oa
na?aaei?e oeie?. – Oiaiue, 1997.– N. 117-119.

Neeaa TH.A. ?noi?eei–a?ae?ia?ao?/iee iaeyae aea/aiiy neiaio?iiii? oei?e
aanaeio Aei?no?a // Iaoa??aee iaoeiaeo /eoaiue, i?enay/aieo 100-??//th
a?aee?eooy iiaea?eiiai caie?aeiaiiy o iie?eoiian?iieo ?ineei i?ioani?ii
oi?aa?neoaoo Nayoiai Aieiaeeie?a N.A.Iaaaoeiei. – E.: O?oinioe?ioeaio?,
1998. – N. 202-204.

Neeaa TH.A. Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio Aei?no?a
(Oe?a?ia).– ?oeiien.

Aeena?oaoe?y ia caeiaoooy a/aiiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe
c? niaoe?aeueiino? 03.00.05. – aioai?ea. Oeaio?aeueiee aioai?/iee naae
?iai? I.I. A?eoea IAI Oe?a?ie, Ee?a, 1999.

Aeena?oaoe?y i?enay/aia ieoaiiyi ?iaaioa?ecaoe?eiiai, oi?ieia?/iiai ?
no?oeoo?iiai aea/aiiy neiaio?iiii? oei?e aa?oiuei? oa/?? aanaeio
Aei?no?a. Neiaio?iiia oei?a aa?oiuei? oa/?? aanaeio Aei?no?a ia?aoiao?
365 aeae?a ?c 213 ?iae?a oa 54 ?iaeei. Anaa?/ii i?iaiae?ciaaii no/aniee
noai neiaio?iiii? oei?e aeaiiai ?aeiio. Aeae?eaii o?aeoe?? oei?e. Aeey
aeae?a ?c aaeaaioeaii? o?aeoe?? i?iaaaeaii aiae?c aeo?aeieo a?aae?a, a
oaeiae a?aeoai?aii ?noi??th ?i?a?aoe?? ? ?icnaeaiiy iecee aaeaaioeaieo
?ineei ia oa?eoi??? Oe?a?ie. Anoaiiaeaii aeieia?/io i?eo?i/ai?noue
neiaio?iiieo aeae?a aei aio?iiiaaiieo aeioii?a. Iaaaaeaii i?iaiic
iiaeeeaeo ci?i ?ineeiiiai iie?eao i?ae aieeaii ainiiaea?nueei?
ae?yeueiino?.

Eeth/ia? neiaa: Aei?noa?, neiaio?iiia oei?a, no?oeoo?a, eeaneo?eaoe?y,
aaeaaioe, aiio?oe, ?i?a?aoe?y, neiaio?ii?caoe?y.

Neeaa TH.A. Neiaio?iiiay oei?a aa?oiaai oa/aiey aannaeia Aeiano?a
(Oe?aeia).– ?oeiienue.

Aeenna?oaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeieiae/aneeo
iaoe ii niaoeeaeueiinoe 03.00.05 – aioaieea. Oeaio?aeueiue aioaie/aneee
naae ei. I.I. A?eoei IAI Oe?aeiu, Eeaa, 1999.

Aeenna?oaoeey iinayuaia aii?inai eiaaioa?ecaoeeiiiiai, oi?ieiae/aneiai
e no?oeoo?iiai eco/aiey neiaio?iiiie oei?u aa?oiaai oa/aiey aannaeia
Aeiano?a. Neiaio?iiiay oei?a aa?oiaai oa/aiey aannaeia Aeiano?a
ian/eouaaao 365 aeaeia ec 213 ?iaeia e 54 naiaenoa.

Ia iniiaa aiaeeca nenoaiaoe/aneie no?oeoo?u neiaio?iiiie oei?u
auyaeaiiua aa oa?aeoa?iua iniaaiiinoe, eioi?ua i?iyaeythoueny a
noiaenoaa niaeo?a ainiiaenoaothueo naiaenoa n aiaeiae/iuie oei?aie
ae?oaeo oa??eoi?ee. Onoaiiaeaii, /oi a ?iaeiaii niaeo?a i?aainoiaeyo
?iaeu n ieceie aeaeiaie ianuuaiiinoth, /oi oa?aoa?ii aeey oei? n
yeno?aiaeueiuie oneiaeyie ?acaeoey ?anoeoaeueiiai iie?iaa. Io niaeo?a
ainiiaenoaothueo naiaenoa neiaio?iiiie oei?u Oe?aeiu ioee/aaony aunieei
iieiaeaiea naiaenoaa Fabaceae – oeie/iie aeey N?aaeecaiiiii?neie
iaeanoe, cia/eoaeueiay ?ieue naiaenoaa Chenopodiaceae, oa?aoa?ioth aeey
ionouiiuo oei?, a oaeaea a ia?aoth aeanyoeo aioee naiaenoaa Rosaceae e
Polygonaceae, i?aaenoaaeoaee eioi?uo i?iec?inoatho a naaa?iie oia?aiiie
ciia.

Aiaeec aeiii?oieiae/aneie e yeieiae/aneie no?oeoo?u neiaio?iiiie oei?u
naeaeaoaeuenoaoao i n?aaeecaiiiii?neii oa?aeoa?a oei?u ?aeiia
enneaaeiaaiee. N?aaee yeiaeiii?o i?aiaeaaeatho aaeeioeou e
iaciena?ioeou, /oi oa?aeoa?ii aeey ciiu noaiae. A ioee/ea io
?aaeiiaeueiie oei?u a neiaio?iiiie oei?a ?aeiia enneaaeiaaiee ca
aaaeoonii e i?iaeieaeeoaeueiinoth aeecie i?aiaeaaeatho iiiiea?ieee.

Ananoi?iiia i?iaiaeece?iaaii nia?aiaiiia ninoiyiea neiaio?iiiie oei?u
aeaiiiai ?aaeiia. Auaeaeaiu o?aeoeee oei?u. Niioiioaiea iaaeaeo ieie
ninoaaeyao 1:1, /oi naeaeaoaeuenoaoao i cia/eoaeueiie ?iee aaeaaioeaiuo
aeaeia a oi?ie?iaaiee neiaio?iiiie oei?u, i?e /ai ca a?aiaiai
eiiea?aoeee i?aiaeaaeatho a?oaioeou, ca niiniaii eiiea?aoeee –
enaiioeou, a ca noaiaiueth iaoo?aeecaoeee –yiaeioeou.

Aaia?aoe/aneay no?oeoo?a enneaaeoaiie oei?u naeaeaoaeuenoaoao i oii, /oi
a ninoaaa neiaio?iiiie oei?u aa?oiaai oa/aiey aannaeia Aeiano?a
aeiieie?otho /aou?a oeia a?aaeiae/aneeo a?oii (aiea?eoe/aneay,
aa?aceaoneay, einiiiieeoiay e aaieeiniiiieeoiay). Cia/eoaeueiia ianoi
n?aaee aiioeoia caieiatho oe?ieia?aaeueiua aiea?eoe/iua e aa?aceeneea
aeaeu ec iacioeeueiuo niiauanoa. N?aaee aaeaaioeaiuo aeaeia
i?aiaeaaeatho einiiiieeou e aaieeiniiiieeou, eioi?ua anoue
i?aaenoaaeoaeyie ena?ioeeueiuo niiauanoa. A ?acoeueoaoa neiaio?iiecaoeee
ai oei?a ?aaeiia oaaee/eeinue eiee/anoai aeaeia aaia?aoe/anee
i?eo?i/aiiuie e oa??eoi?eyi n ena?e/aneeie oneiaeyie.

Aeey aeaeia ec aaeaaioeaiie o?aeoeee i?iaaaeai aiaeec enoiaeiuo a?aaeia.
Ii naeaeaoaeuenoaoao i?i aunieoth ianuuaiiinoue neiaio?iiiie oei?u
?aeiia enneaaeiaaiee aaeaaioeaiuie aeaeaie n?aaeecaiiiii?neiai,
aceaoneiai e naaa?iaia?eeaineiai i?ienoiaeaeaiey a nayce n iniaiaieai
eie iiauo aio?iiiaaiii i?aia?aciaaiiuo yeioiiia.

Ainnicaeaia enoi?ey eiiea?aoeee e ?annaeaiey ?yaea aaeaaioeaiuo
aeaeia ia oa??eoi?ee Oe?aeiu. Ia iniiaa ninoaaeaiiuo ea?o a?aaeia
onoaiiaeaiu iauea caeiiiia?iinoe iea?aoeee ia oa??eoi?ee Oe?aeiu
n?aaeecaiiiii?neeo e aceaoneeo aeaeia a naaa?i–caiaaeiii iai?aaeaiee.
Onoaiiaeaia yeieiae/aneay i?eo?i/aiiinoue neiaio?iiiuo aeaeia e
aio?iiiaaiiui yeioiiai.

Ia iniiaa oei?ieiae/aneie ioeaiee iineaaenoaee neiaio?iiecaoeee e eo
i?iyaeaiee ia i?aaieciaiiii, iiioeyoeeiiiii e yeinenoaiiii o?iaiyo
onoaiiaeaii, /oi iia i?ienoiaeeo a iai?aaeaiee ecieiaiey no?oeoo?iie
i?aaiecaoeee oei?u, aaeaioaoeee aeaeia e iiioeyoeee e aeaenoaeth
aio?iiiaaiiuo oaeoi?ia e ia?anoaiea aa aeeiaie/iuo oaiaeaioeee.

Eeth/aaua neiaa: Aeiano?, neiaio?iiiay oei?a, no?oeoo?a, eeanneoeeaoeey,
aaeaaiou, aiioeou, eiiea?aoeey, neiaio?iiecaoeey.

Skyba Yu.A. The synanthropic flora of the upper reaches of the Dnistro
(Dnister) River, Ukraine.– Manuscript.

Dissertation on pretence of candidate of biological scienses degree.
Fild of study 03.00.05 – botany. M.M. Grishko Central Botanical Garden
of National Academy of Sciences of Ukraina, Kiev, 1999.

Dissertation is devoted to inventarization, chorological and structural
study of synantrope Flora in the upper course of Dnister river.
Synanthropic flora of the region mentioned includes 365 species, 213
genuses and 54 families, its modern condition has been analysed.
Different flora of has been determined. Initialales of adventive
fraction’s species has been defined. Immigration and distribution time
has been found out for some adventive species. Synanthropic species has
been reffered to antropogenic ecotops. The forecast of probable
successive changes in flora under infuence of human activities is given.

Key words: Dnister river, synanthropic flora, stuctura, classification,
adventive plants, apophytos, immigration, synantropization.

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