Поліморфізм гетерохроматинових районів акроцентричних хромосом свиней: Автореф. дис… канд. с.-г. наук / М.М. Івашура, УААН. Ін-т тваринництва. — Х.,

3

OE?A?INUeEA AEAAeAI?ss AA?A?IEO IAOE

?INOEOOO OAA?EIIEOeOAA

?aaoo?a Ia?eia Ieeiea?aia

OAeE 636.4.082

Iie?ii?o?ci aaoa?io?iiaoeiiaeo ?aeii?a

ae?ioeaio?e/ieo o?iiinii naeiae

03.00.15 — aaiaoeea

Aaoi?aoa?ao

Aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa n?euenueeiainiiaea?nueeeo iaoe

Oa?e?a – 1998

Aeena?oaoe??th ? ?oeiien

?iaioa aeeiiaia a eaai?aoi??? aaiaoeee ?inoeoooo

oaa?eiieoeoaa OAAI

Iaoeiaee ea??aiee: eaiaeeaeao n?euenueeiainiiaea?nueeeo iaoe,

noa?oee iaoeiaee ni?a?ia?oiee,

?icnioa Aieiaeeie? ?aaiiae/,

?inoeooo oaa?eiieoeoaa OAAI,

caa?aeoaa/ eaai?aoi???th aaiaoeee

Io?oe?ei? iiiiaioe: aeieoi? a?ieia?/ieo iaoe,
i?ioani?

Oaoaacia
Aaea??e Aaeiae/,

Oa?e?anueeiai aea?aeaaiiai oi?aa?neoao,

Caa?aeoaa/ eaoaae?ith aaiaoeee oa oe?oieia??

eaiaeeaeao
a?ieia?/ieo iaoe,

Aaaaoeueea Iaoae?y Aanee?aia

OIAe? ioi?iie caei?ia’y ae?oae oa i?aee?oe?a,

noa?oee iaoeiaee ni?a?ia?oiee

eaai?aoi??? iaaee/ii? aaiaoeee

I?ia?aeia onoaiiaa:

?inoeooo naeia?noaa OAAI, i. Iieoaaa

Caoeno a?aeaoaeaoueny ” 22 ”_a?oaeiy 1998 ?ieo i 10 aiaeei? ia
can?aeaii? niaoe?ae?ciaaii? ?aaee 65.356.01 i?e ?inoeooo?
oaa?eiieoeoaa OAAI, 312120, i. Oa?e?a, i/a Eoeeie/?.

C aeena?oaoe??th iiaeia iciaeiieoeny a a?ae?ioaoe? ?inoeoooo

oaa?eiieoeoaa OAAI, 312120, i. Oa?e?a, i/a Eoeeie/?

Aaoi?aoa?ao ?ic?neaiee “ 20 “_eenoiiaaea 1998 ?ieo

A/aiee nae?aoa? niaoe?ae?ciaaii? a/aii? ?aaee Oaaoia A. ?.

CAAAEUeIA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. Aoaeoeai?noue naeaeoe?eii? ?iaioe o oaa?eieoeoa?
iaaiith i??ith iaoiiaeth?oueny aaiaoe/iei iioaioe?aeii oaa?ei, ye?
aeei?enoiaothoueny aeey a?aeoai?aiiy noaaea. Eiio?ieue
a?aeoai?thaaeueii? caeaoiino? o?aaeeoe?eieie iaoiaeaie ia aeicaiey?
io?eioaaoe ae/a?iio ?ioi?iaoe?th i?i oaeoi?e, ye? aieeaathoue ia
caie?aeithaai?noue oa aia??iiaeueio aeeoo?caeaoi?noue. ?ioaineai?
oeeoiaaiaoe/i? aeine?aeaeaiiy aeyaeee ?yae iieaciee?a, iia’ycaieo c
i?iyaeaiiyie ?ai?iaeoeoeaii? ooieoe?? o oaa?ei.Aeei?enoaiiy iaoiae?a
aeeoa?aioe?eiiai caaa?aeaiiy o?iiinii iaoiiaeee aaaeeea? iai?yiee
oeeoiaaiaoe/ieo aeine?aeaeaiue. Niaoeeo?/i?noue iaiaeii??aeiiai
caaa?aeaiiy o?iiiniiiiai iaoa??aeo ia o?eueee noaea caniaii iaae?eii?
?aeaioeo?eaoe?? o?iiiniiieo ia?, aea e aeicaieeea aeyaeyoe iaai? oeie
o?iiiniiii? i?aai?caoe??: aoo?iiaoei oa aaoa?io?iiaoei. Iniaeeaino?
iieaeoey?iiai neeaaeo oa no?oeoo?e oeeo ?aeii?a o?iiinii, a oaeiae ?o
aca?iiae?y ia ??cieo aoaiao ee?oeiieo i?ioean?a ? iniiaith ?aae?caoe??
aaiaoe/ii? ?ioi?iaoe??. ?aeiie no?oeoo?iiai aaoa?io?iiaoeio
oa?aeoa?ecothoueny aenieith aa??aaaeuei?noth ye a iaaeao aeaeo, oae ? a
iaaeao iaeiiai aaiiio O no?oeoo?? aaoa?io?iiaoeiiaeo ?aeii?a o?aaiaioe
aaaaoiiiaoi?thaaieo iine?aeiaiinoae AeIE o?nii iia’ycai? c iao?eoeyie
aeayeeo aeeoo?ai aaaeeeaeo aai?a. Ca e?oa?aoo?ieie aeaieie,
aa??aaaeuei?noue aaoa?io?iiaoeiiaeo ?aeii?a iiaea aieeaaoe ia aeni?an?th
aai?a, cie?aia, a aia??iaaiac?.

Oiio aeine?aeaeaiiy iie?ii?oieo aa??aio?a o?iiinii oa iiooe ca’yce?a
i?ae oeeoiaaiaoe/ieie iieacieeaie oa ?ai?iaeoeoeaieie yeinoyie naeiae
ia? cia/aiiy ye aeey oai???, oae ? aeey i?aeoeee naeia?noaa.
Noaa?euei?noue ii?oieia?/ieo oei?a o?iiinii aeeeeea? ?ioa?an c oi/ee
ci?o ?nioaaiiy i?e?iaeieo o?iiiniiieo ia?ea??a. A ca’yceo c oeei,
oaeineiiaeaiiy ?nioth/eo iaoiae?a aeine?aeaeaiue oa ?ic?iaea iiaeo
i?aeoiae?a aei oeeeoiaaiaoe/iiai aiae?co ? aaaeeeaei oa iaiao?aeiei
aoaiii o aea/aii? iie?ii?o?cio o?iiinii.

Aeine?aeaeaiiy o?iiinii naeiae, caaa?aeaieo ye ?ooeiiei, oae ?
aeeoa?aioe?eiei N-iaoiaeii, iaaeathoue aaaeeeao ?ioi?iaoe?th i?i noai
aaiiio, yea iiaea aeei?enoiaoaaoenue i?e ae??oaii? i?iaeai i?aeoe/ii?
naeaeoe??.

Ca’ycie ?iaioe c iaoeiaeie i?ia?aiaie, ieaiaie, oaiaie. ?iaioa ?
neeaaeiaith /anoeiith iaoeiai-oaoi?/ii? eiiieaenii? i?ia?aie
“I?iaeiaieuenoai-95” caaaeaiiy I.04.02 “?ic?iaeoe aoaeoeaio nenoaio
eiio?ieth, ioe?iee, ni?yiiaaii? ci?ie oa caa?aaeaiiy aaiaoe/ii?
no?oeoo?e oa iioaioe?aeo iiioeyoe?e naeiae ia iniia? aeei?enoaiiy
?ioiiaaiaoe/ieo oa oeeoiaaiaoe/ieo iaoiae?a” ia 1991-1995 ??. (? aea?ae.
?a?no?aoe?? 0196U01390 ).

Iaoa oa caaea/? aeine?aeaeaiue. Aeine?aeaeaiiy N-iie?ii?o?cio o?iiinii
naeiae i?iaiaeeeinue c iaoith aecia/aiiy aca?iica’yceo i?ae
aa??aaaeuei?noth aaoa?io?iiaoeiiaeo ?aeii?a oa ?ai?iaeoeoeaiith
ooieoe??th oaa?ei oa aea/aiiy iiaeeeainoae aeei?enoaiiy oeeoiaaiaoe/ieo
iieaciee?a o i?aeoe/i?e naeaeoe??. Aeey aeinyaiaiiy iinoaaeaii? iaoe
ae??ooaaeenue oae? caaaeaiiy:

1. ?ic?iaeoe iiaeeo?eaoe?th aeeoa?aioe?eiiai N-iaoiaeo caaa?aeaiiy
o?iiinii naeiae.

2. ?ic?iaeoe e?eoa??? a?coaeueii? ioe?iee aaoa?io?iiaoeiiaeo ?aeii?a ia
ae?ioeaio?e/ieo o?iiiniiao naeiae.

3. Aea/eoe iiaeeea?noue aeine?aeaeaiiy aeeoa?aioe?eii caaa?aeaieo
o?iiinii c aeei?enoaiiyi eiii’thoa?ii? oaoi?ee.

4. Aea/eoe eieae?caoe?th oa iaae? aa??aaaeueiino? aaoa?io?iiaoeiiaeo
?aeii?a ia o?iiiniiao naeiae ii??ae aaeeea a?ea oa eaiae?an.

5. Aea/eoe aca?iica’ycie N-iie?ii?o?cio o?iiinii c ?ai?iaeoeoeaiith
ooieoe??th naeiae.

6. Ii??aiyoe /anoioo oa niaeo? niiioaiieo iooaoe?e o iin??a iie?ii?oieo
aa??aio?a o?iiinii.

7. Aea/eoe aieea aa??aaaeueiino? i?eoeaio?ii??iiai aaoa?io?iiaoeio ia
?aae?caoe?th anioe?aoe?e ae?ioeaio?e/ieo o?iiinii.

8. I?iaanoe iiooe ia?ea?ieo o?iiinii oa aea/eoe ?o oniaaeeoaaiiy.

Iaoeiaa iiaecia iaea?aeaieo ?acoeueoao?a. Aecia/aii iioeiaeuei? oiiae
N-naaiaioaoe?? ia o?iiiniiao naeiae. ?ic?iaeaii e?eoa???
iai?ae?euee?nii? ioe?iee ?ici???a N-aeie?a ia ae?ioeaio?e/ieo o?iiiniiao
ca 5-aaeueiith oeaeith. Ia iniia? ?ic?iaeaiiai iaoiaeo aeine?aeaeaii
iaae? aa??aaaeueiino? ?aeii?a i?eoeaio?ii??iiai N-aaoa?io?iiaoeio oa
aecia/aii /anoioo iie?ii?oieo aa??aio?a o?iiinii 13-18 o naeiae ii??ae
aaeeea a?ea oa eaiae?an. ?ic?iaeaii niin?a oa iaeaaeiaiiy aeey
a?coae?caoe?? oa eiii’thoa?ii? ia?iaee i?e?icia?aaeaiue o?iiinii.
Aeei?enoaiiy i?ia?aiii-aia?aoiiai eiiieaeno aeicaiey? ea??ioeioaaoe
o?iiiniie, caaa?aeai? ye ?ooeiiei, oae ? aeeoa?aioe?eieie iaoiaeaie.
Aeine?aeaeaii ca’ycie i?ae N-iie?ii?o?ciii o?iiinii oa
a?aeoai?thaaeueieie yeinoyie naeiae, a oaeiae aieea ??cieo eiia?iaoe?e
iie?ii?oieo aiiieia?a ia o?iiiniiio ianoaa?euei?noue o eaeeioeeoao
e?ia?.

I?aeoe/ia cia/aiiy io?eiaieo ?acoeueoao?a. ?acoeueoaoe aea/aiiy ca’yceo
N-iie?ii?o?cio o?iiinii c i?iyaaie ?ai?iaeoeoeaii? ooieoe?? naeiae, a
oaeiae c ianoaa?euei?noth aaiiio ia ee?oeiiiio ??ai?, ? aaaeeeaith
/anoeiith caaaeueiiai oeeoiaaiaoe/iiai oanooaaiiy
n?euenueeiainiiaea?nueeeo oaa?ei, ? iiaeooue aeei?enoiaoaaoenue i?e
ae??oaii? iaaieo naeaeoe?eieo caaea/. ?ic?iaeai? e?eoa??? a?coaeueiiai
aiae?co aeicaieythoue i?iaiaeeoe aeni?an-ioe?ieo iie?ii?oieo N-aa?eaio?a
o?iiinii. Aeei?enoaiiy iai?aaaoiiaoe/ii? nenoaie aeey ia?iaee
i?e?icia?aaeaiue o?iiinii oa ea??ioeioaiiy ? aoaeoeaiei iaoiaeii i?e
i?iaaaeaii? ianooaaieo aeine?aeaeaiue noaio aaiiio
n?euenueeiainiiaea?nueeeo oaa?ei, a oaeiae ? iaiao?aeiith oiiaith
noai?aiiy aaieo aeaieo ieai?iieo oaa?ei. ?ic?iaeaia iiaeeo?eaoe?y
N-iaoiaeo aeicaiey? io?eioaaoe ye?nia N-caaa?aeaiiy ia o?iiiniiao
naeiae aac ii?ooaiiy ii?oieia?? o?iiinii.

Iniaenoee aianie caeiaoaa/a. Aeena?oaio ia?aaaaeio /anoeio ?ia?o aeeiiaa
naiino?eii. Iaoiaeieia?y oa noaia aeine?aeaeaiue ?ic?iaeai? ?acii c
iaoeiaei ea??aieeii. ?c caaaeueieo iaoeiaeo aenia?eiaio?a oa ioae?eaoe?e
aeena?oaio aeei?enoaa ca caiaeith ni?aaaoi??a o?eueee naith /anoeio
?acoeueoao?a.

Ai?iaaoe?y ?acoeueoao?a aeena?oaoe??. Iniiai? iieiaeaiiy aeena?oaoe??
aoee i?aaenoaaeai? oa iaaiai?ai? ia A/ai?e ?aae? ?inoeoooo oaa?eiieoeoaa
OAAI, iaoeiai-i?aeoe/i?e eiioa?aioe?? iieiaeeo a/aieo oa niaoe?ae?no?a
“Iaoeia? ?ic?iaee aa?ii?iieneiaiai eiiieaeno” Ieaei? Ai?ioa, 1993;
ia?o?e iaoeia?e eiioa?aioe?? iieiaeeo a/aieo oa niaoe?ae?no?a
“Naeaeoe?eii-a?ioaoiieia?/i? iaoiaee aeei?enoaiiy aaiaoe/iiai
iioaioe?aeo n?euenueeiainiiaea?nueeeo oaa?ei”, Ee?a, 1994?.; ae?oa?e
i?aeia?iaei?e eiioa?aioe?? “Iieaeoey?ii-aaiaoe/i? ia?ea?e”, Ee?a,
1996?.; eiioa?aioe?yo iieiaeeo a/aieo oa niaoe?ae?no?a ( Oa?e?a,
1996?., Oa?e?a, 1998?).

Ioae?eaoe??. Iniiaiee ci?no aeena?oaoe?eii? ?iaioe aeeeaaeaii o 11
ae?oeiaaieo i?aoeyo. o oiio /ene? 3 noaooyo o oaoiaeo aeaeaiiyo,
caoaa?aeaeaieo AAE Oe?a?ie, oa 8 oacao aeiiia?aeae iaoeiai-i?aeoe/ieo
eiioa?aioe?e.

No?oeoo?a oa ianya aeena?oaoe??. Aeena?oaoe?th aeeeaaeaii ia 150
noi??ieao iaoeiiieniiai oaenoo, ai?uo? 18 oaaeeoeue oa 18 ?enoie?a.
Nienie e?oa?aoo?e neeaaea?oueny c 243 aeaea?ae, o oiio /ene? 162 ia
?iicaiieo iiaao.

IAOA??AE OA IAOIAeEEA AeINE?AeAEAIUe

Aeine?aeaeaiiy i?iaiaeeeenue o eaai?aoi??? aaiaoeee ?inoeoooo
oaa?eiieoeoaa OAAI ia ieai?iiiio iiaie?a’? naeiae ae/a “Oe?a?iea”,
Aoaeueii?anueeiai ieaicaaiaeo oa Aei?i?iiao?ianueeiai NAOe o ia??iae c
1991 ii 1995 ?. Eieae?caoe?th N-aeie?a aaoa?io?iiaoeio i?iaiaeeee ia
o?iiiniiao 25 eio??a, 27 naeiiiaoie oa 33 ii?inyo (a?eii aei 2
i?nyoe?a) ii??ae aaeeea a?ea oa eaiae?an. Caa?? e?ia?, eoeueoeaoaaiiy
eaeeioeeo?a oa aeaioiaeaiiy i?aia?ao?a o?iiinii i?iaiaeeee ca
caaaeueiii?eeiyoeie iaoiaeeeaie. Aeey aeyaeaiiy ?aeii?a no?oeoo?iiai
aaoa?io?iiaoeio ia o?iiiniiao naeiae aeei?enoiaoaaee N-iaoiae
caaa?aeaiiy o iiaeeo?eaoe?? . Ioe?ieo iie?ii?oieo N-aa??aio?a
ae?ioeaio?e/ieo, o?iiinii i?iaiaeeee ca ?acoeueoaoaie a?coaeueiiai
ea??ioeioaaiiy c aeei?enoaiiyi iai?ae?euee?niiai iaoiaeo.

Iae?e niiioaiieo o?iiiniiieo aaa?aoe?e i?iaiaeeee ia i?aia?aoao
o?iiinii, caaa?aeaieo ?ooeiiei niiniaii aa?aieeii A?ica, ca?aeii
iaoiaee/ieo ?aeiiaiaeaoe?e, ?ic?iaeaieo o eaai?aoi??? aaiaoeee (Iao.
?ae., Oa?e?a, 1990). A?ae eiaeii? oaa?eie aiae?coaaee ii 50 iaoaoac.
Aiaoiei?ae?y (N2) oa iie?iei?ae?y neeaaeaee ??aaiue aaiiiieo iooaoe?e.
No?oeoo?i? aaa?aoe?? aoee i?aaenoaaeai? ii?ooaiiyie o?iiiniiiiai oa
o?iiaoeaeiiai oeio, a oaeiae /enaeueiith o?aaiaioaoe??th o?iiinii.
Aeey aea/aiiy aca?iica’yceo N-iie?ii?o?cio o?iiinii c ?o anioe?aoeaiith
caeaoi?noth i?iaiaeeee aiae?c /anoioe oa niaeo?o anioe?aoe?e
ae?ioeaio?e/ieo o?iiinii (AAO) o ca’yceo c iayai?noth iie?ii?oieo
o?iiiniiieo aa??aio?a, a oaeiae ?o eiia?iaoe?yie. Aiae?c i?aia?ao?a
o?iiinii i?iaiaeeee o nenoai? na?oiaiai i?e?ineiio ”Jenaval” (Zeiss,
I?ia//eia) c ianeyiith ?ia?n??th i?e ca?eueooaaii? 10o100. I?e?iceiiea
ie?aieo iaoaoacieo ieao?aie i?iaiaeeeanue ia i?e?iie?aeo “I?e?ao-200”.
Aeey a?coae?caoe?? oa eiii’thoa?ii? ia?iaee i?e?icia?aaeaiue o?iiinii
aeei?enoiaoaaee ?ic?iaeaiee o eaai?aoi??? aaiaoeee i?ia?aiii-aia?aoiee
eiiieaen aeey ea??ioeioaaiiy o?iiinii n?euenueeiainiiaea?nueeeo
oaa?ei.Oeeo?iaee iaoa??ae ia?iaeyee iaoiaeaie aa??aoe?eii? noaoenoeee
i?e aeei?enoaii? iaeaoo i?ia?ai aiae?co aeaieo STADIA.

?ACOEUeOAOE AeINE?AeAEAIUe

1. Aeine?aeaeaiiy aoai?a aeeoa?aioe?eiiai N-aa?aeaiiy o?iiinii naeiae.
Aeine?aeaeaiiy o?iiinii c aeei?enoaiiyi iaoiae?a aeeoa?aioe?eiiai
caaa?aeaiiy ? aaaeeeaith /anoeiith oeeoiaaiaoe/iiai oanooaaiiy.
Aea/aiiy aca?iica’yceo N-iie?ii?o?cio o?iiinii c i?iaeoeoeaieie yeinoyie
oaa?ei, a oaeiae iiaeeea?noue aeei?enoaiiy i?e?iaeieo o?iiiniiieo
ia?ea??a, i?aaenoaaey?oueny aeooaeueiei iai?yieii aeine?aeieoeueei?
?iaioe. I?ioa, a ca’yceo c neeaaei?noth oa iecueeith a?aeoai?thaai?noth
?acoeueoao?a N-caaa?aeaiiy, oeae iaoiae ia io?eiaa iioe?aiiy o
oeeoiaaiaoe/ieo aeine?aeaeaiiyo n?euenueeiainiiaea?nueeeo oaa?ei.
Iaaeie?eii N-iaoiaeeee aaaaea?oueny cia/ia aeaiaoo?aoe?y o?iiiniiiiai
iaoa??aeo a ?acoeueoao? ae?? eoaeieo oa nieueiaeo ?ic/ei?a. Aeey
io?eiaiiy ye?niiai N-caaa?aeaiiy oa caa?aaeaiiy ii?oieia?? o?iiinii
aeiaiaeeoueny ?ii??e/ii i?aeae?aoe oiiae, ci?ithth/e ye o?i?/i?, oae ?
o?ce/i? iieacieee ae?th/eo aaaio?a. Oiio aeieeea iaiao?aei?noue
i?iaiae?coaaoe iniiai? aoaie N-iaoiaeo oa aecia/eoe iioeiaeuei? oiiae
N-caaa?aeaiiy o?iiinii naeiae.

Iaea?eueo aoaeoeaieie aeyaeeeny iine?aeiaiino? ia?iaie, ye? aeeth/aee
eoaeio ia?aaeia?iaeo i?aia?ao?a. I?iaaaeaiiy i’yei? aeaiaoo?aoe?? c
aeei?enoaiiyi a?ae?ieneaea aa??th aeicaieyei aeinyaiooe N-naaiaioaoe??,
oieeath/e neeueiiai ii?ooaiiy o?iiinii o ii??aiyii? c ?acoeueoaoaie
ia?iaee a?ae?ieneaeii iao??th. I?e ?ieoaaoe?? i?aia?ao?a o noaiaea?oiiio
nieueiaiio ?ic/ei? (SSC) ia?aaaaeia cia/aiiy iaa iieaciee pH. Aecia/aii,
ui ci?ia oeueiai iieacieea o eoaeio noi?iio (?H 8,5) aeyaeeany
iioeiaeueiith aeey io?eiaiiy ye?niiai N-caaa?aeaiiy ia o?iiiniiao
naeiae. Aeey iaee?aui? noaa?e?caoe?? o?iiiniiiiai iaoa??aeo
cai?iiiiiaaii aeei?enoaiiy aooa?ii-nie?oiai? noi?o?. ?ieoaaoe?y
i?aia?ao?a o nieueiaiio ?ic/ei? c aeiaeaaaiiyi 5 ie 96%-iiai aoaiieo ia
100 ie 2oSSC i?e pH 8,5 aeicaiey? cia/ii e?aua caa?aaoe ii?oieia?th
o?iiinii, iaa?oue i?e iaiao?aeiino? ca?eueoeoe /an ae?? aai oaiia?aoo?o
aooa?iiai ?ic/eio.

A ?acoeueoao? aeine?aeaeaiiy iniiaieo aoai?a N-iaoiaeeee aecia/aii
iaea?eueo aaaeeea? oaeoi?e, ye? aieeaathoue ia noai o?iiaoeio o ia??iae
ia?aaeia?iaee i?aia?ao?a, oa ?ic?iaeaii iiaeeo?eaoe?th oeueiai iaoiaea
aeey io?eiaiiy ye?niiai N-caaa?aeaiiy ia o?iiiniiao naeiae. Iniiaieie
iiiaioaie cai?iiiiiaaii? iine?aeiaiino? ia?iaie ? i’yea aeaiaoo?aoe?y
a?ae?ieneaeii aa??th (5%-iee Ba(OH)2, 500N) oa ?ieoaaoe?y o
aooa?ii-nie?oiaiio ?ic/ei? 2oSSC i?e pH 8,5 (2-4 a., 650N).

2. Aiae?c N-iie?ii?o?cio o?iiinii naeiae a?coaeueiei oa eiii’thoa?iei
niiniaii.O aeine?aeaeaiiyo N-iie?ii?o?cio o?iiinii aaaeeeaith i?iaeaiith
caeeoa?oueny ia’?eoeaia ioe?iea ?ici???a N-aeie?a aaoa?io?iiaoeio ia
o?iiiniiao naeiae. O i?aeoeoe? iaaee/ii? aaiaoeee aeey aiae?co
aa??aaaeueieo N-aeie?a aaoa?io?iiaoeio aeei?enoiaothoue iai?ae?euee?ni?
iaoiaee ioe?iee, yee aeicaieythoue aecia/aoe a?aeiini? ?ici??e N-aeie?a
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Aei??thaaiiy ii?oiiao?e/ieo ia?aiao??a aa??aaaeueieo ?aeii?a
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Aeie-noaia onoaiiaee aeey iai?aaaoiiaoe/iiai ea??ioeaoaaiiy. 1- eiiaeaini? c ina?oethaa/ai, 2 - eii?aeeiaoiee noieee i?e?ineiio, 3- i?aia?ao, 4- i?e?ineii, 5 - a?aeaieaia?a, 6- a?aeaiii?o, 7- ia?niiaeueiee eiii’thoa?. Nenoaia ciiioiaaia ia aac? i?e?ineiio “Ethiai-E2”. Iniiaiith neeaaeiaith /anoeiith ? a?aeaieaia?a 5, yea ca aeiiiiiaith a?aeaiii?oo 6 i?aeeeth/a?oueny aei IAII 7. Aeei?enoaiiy oeueiai iaeaaeiaiiy aeicaiey? aeine?aeaeoaaoe o?iiiniie, caaa?aeai? ye ?ooeiiei, oae ? aeeoa?aioe?eiei niiniaii. Cia?aaeaiiy o?iiinii ca aeiiiiiaith a?aeaieaia?e o?aineth?oueny c i?e?ii?aia?aoo aei IAII oa aeaiaeeoueny ia ae?ai iii?oi?o. I?ney ia?iaee cia?aaeaiue oi?io?oueny oaee ii/aoeiaeo aeaieo, a yeiio ciaoiaeyoueny a?ao?/i? ia?aiao?e i?e?iia’?eo?a. Aaaeeeaei aoaiii oeeoiaaiaoe/ieo aeine?aeaeaiue ? ea??ioeioaaiiy o?iiiniiiiai iaai?o. Aeey ae??oaiiy oe??? caaea/? iaie ni?eueii c o??iith “Iio?ii”(i.Ee?a) aoei ?ic?iaeaii i?ea?iaeueia i?ia?aiia caaacia/aiiy. ?ic?iaea i?ia?aie KARYOTYPE aoea i?eioeeiiai ni?yiiaaia ia ?ioa?aeoeaiee ?aaeei ?iaioe, ui aeicaiey? iia?aoi?o-aeine?aeieeo eiio?iethaaoe oa oaeaeei i?iaiaeeoe ei?aeoe?th an?o aoai?a aiae?co. Ea??ioeioaaiiy o?iiinii i?iaiaeeoueny o iai?aaaoiiaoe/iiio ?aaeei?. Ia?aaeaa/aia iiaeeea?noue ?aaeaaoaaiiy ea??ia?aie: ei?aeoe?y iieiaeaiiy oeaio?ii??e, i?aeai? aiiieia?/ieo o?iiinii. Aeei?enoaiiy eiii’thoa?ii? oaoi?ee o oeeoiaaiaoe/ieo aeine?aeaeaiiyo n?euenueeiainiiaea?nueeeo oaa?ei ia o?eueee aeicaiey? cia/ii i?aeaeueoe aoaeoeai?noue iaoiae?a i?e?iaeine?aeaeaiue, aea ? ? iaiao?aeiei i?e noai?aii? aaieo aeaieo ieai?iieo oaa?ei. I?ioa, ?ic?iaea iaoiae?a a?coaeueiiai aiae?co, ye? iiaeia aeei?enoiaoaaoe ia o?eueee o niaoe?ae?ciaaieo eaai?aoi??yo c eiooiaiei iaeaaeiaiiyi, oaeiae i?aaenoaaey?oueny aeioe?eueiith. 3. Aa??aaaeuei?noue ?aeii?a no?oeoo?iiai aaoa?io?iiaoeioia ae?ioeaio?e/ieo o?iiiniiao naeiae.Aiae?c e?oa?aoo?ieo aeaieo iieacaa, ui iie?ii?o?ci N-aeie?a aaoa?io?iiaoeio ia o?iiiniiao naeiae aeyaeaii o aaaaoueio aeine?aeaeaiiyo. I?ioa, oa?aeoa?enoeea ia?aiao??a oe??? i?ieeaino? iaiaaeaia, oiae? ye aa??aaaeuei?noue ?aeii?a N-aaoa?io?iiaoeio iaoiiaeth? no?oeoo?io iaiaeii??aei?noue o?iiiniiiiai iaoa??aeo, ui ne?ae a?aoe aei oaaae i?e ae??oaii? naeaeoe?eieo caaea/. Aeine?aeaeaiiy caaa?aeaieo ii N-iaoiaeo o?iiinii naeiae iieacaee, ui iaea?eueo aaeeeeie oa aa??aaaeueieie ? aeiee N-aaoa?io?iiaoeio ia ae?ioeaio?e/ieo o?iiiniiao. eei?enoiaoth/e e?eoa??? iai?ae?euee?niiai iaoiaeo ioe?iee i?iaaee aeine?aeaeaiiy aa??aaaeueiino? ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio ia ae?ioeaio?e/ieo o?iiiniiao naeiae ii??ae aaeeea a?ea oa eaiae?an. *anoioa iie?ii?oieo N-aa??aio?a o?iiinii 13 – 18 iaaaaeaia o oaae. 2. A ?acoeueoao? aiae?co /anoioe ii?oieia?/ieo N-oei?a ae?ioeaio?e/ieo o?iiinii aecia/aii, ui iae/ano?oa iie?ii?oi? aeiee N-aaoa?io?iiaoeio eieae?coaaeeny ia o?iiiniiao 13, 15, 16 oa 18. Iaea?eueoith aa??aaaeuei?noth N-aaoa?io?iiaoeiiaiai ?aeiio oa?aeoa?ecoaaeany o?iiiniia 16, /anoioa o?iiiniiieo aa??aio?a c aaeeeeie N-aeieaie ( 3-5 aae?a) neeaaeaea 45,8%.Oaaeeoey 2. *anoioa (%) N-aa??aio?a ae?ioeaio?e/ieo o?iiinii o naeiae (n=45, aaeeea a?ea oa eaiae?an). ??ici??e N-aeieo aaoa?io?iiao?ioo?iiiniie1 aae2 aaee3 aaee4 aaee5 aae?a135.5667.2625.0001454.4442.223.3300157.7822.2231.116.6701613.1940.2 834.0310.421.391785.5612.222.22001867.7814.447.785.560 Ii??aiyiiy /anoioe iie?ii?oieo N-aa??aio?a o?iiinii 13, 15, 16 oa 18 o naeiae ii??ae aaeeea a?ea (n = 36) oa eaiae?an (n = 34) iieacaei, ui ca aa??aaaeuei?noth N-aaoa?io?iiaoeia i?aaenoaaieee aeaio ii??ae i?aeoe/ii ia a?ae??ciythoueny. Iaaaeeea ??cieoey o /anoio? o?iiiniiieo aa??aio?a 16 oa 18 c iaeaiueeeie N-aeieaie (1 aae ) iiaea na?ae/eoe i?i ii/aoie iaeiie/aiiy iaaieo o?iiiniiieo N-aa??aio?a o aeine?aeaeoaaieo ii?iaeieo a?oiao oaa?ei (?<0,01). Aeine?aeaeaiiy ?iciiae?eo ii?oieia?/ieo N-oei?a o?iiinii 13, 15, 16 oa 18 o naeiiiaoie oa eio??a aeaio ii??ae oaeiae ia aecia/eee iii?oii? ??cieoe? ii /anoio? iie?ii?oieo N-aeie?a i?ae oaa?eiaie ??cii? noao?, ui aeicaiey? aaaaeaoe ooieoe?iiaeueia cia/aiiy aa??aaaeueieo ?aeii?a N-aaoa?io?iiaoeio ae?ioeaio?e/ieo o?iiinii iaeo?aeueiei o a?aeiioaii? aei noao? naeiae. A ?acoeueoao? aeine?aeaeaiue aecia/aii, ui aa??aoe?? N-aaoa?io?iiaoeiiai? /anoeie aaiiio iaoiiaeai? ia?aaaaeii iie?ii?o?ciii o?iiinii 13, 15, 16 oa 18. A?aenooi?noue cia/ii? ??cieoe? o ?iciiae?e? iie?ii?oieo N-aa??aio?a o?iiinii i?ae eio?aie oa naeiiiaoeaie ii??ae aaeeea a?ea oa eaiae?an aeicaiey? aaaaeaoe aa??aaaeuei?noue ?aeii?a N-aaoa?io?iiaoeio iaeo?aeueiith oa?aeoa?enoeeith ye o a?aeiioaii? aei noao? naeiae, oae ? a ii?iaeiiio aniaeo?. 4.Aca?iica’ycie N-iie?ii?o?cio ae?ioeaio?e/ieo o?iiinii c ?ai?iaeoeoeaiith ooieoe??th naeiae. Iie?ii?o?ci ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio niinoa??aa?oueny ia o?eueee a iaaeao iaeiiai aeaeo, aea e a iaaeao iaeiiai aaiiio. Aiiieia?/i? o?iiiniie, N-aeiee yeeo a?ae??ciyeeny ca ?ici??aie, aaaaeathoueny aaoa?iii?oieie, i?e oeueiio ?ic??cithaaee aaoa?iii?o?ci neaaeiai (1 aae) oa neeueiiai nooiaiy (2-3 aaee). I?iaaaeaii aiae?c iniiaieo iieaciee?a a?aeoai?thaaeueii? caeaoiino? naeiae (n=48) ii??ae o ca’yceo c iayai?noth o aaiii? iie?ii?oieo N-aa??aio?a o?iiinii 13, 15, 16 oa 18, a oaeiae eiia?iaoe?yie aaoa?iii?oieo aiiieia?a. Aeine?aeaeaiiy iieacaei, ui iin?? aaeeeiai N-aeieo (3-5 aae?a) ia o?iiinii? 16 oa?aeoa?ecoaaeenue a??ia?aeii aeuei (?<0,05) iieacieeii aaaaoiie?aeaey (10,00,23) o ii??aiyii? c iin?yie iaeaiueeeo N-aeie?a (9,00,61). Ii??aiyiiy a?aeoai?thaaeueii? caeaoiino? naeiae, ui oa?aeoa?ecoaaeenue aaoa?iii?o?ciii aiiieia?/ieo o?iiinii, aeyaeei ??cieoeth ca iieacieeaie e?eueeino? ii?inyo o 2 i?ny/iiio a?oe? oa caa?aaeai?noth i?eieiaeo, iia’ycaio c N-aaoa?iii?o?ciii aiiieia?a 16-i? ia?e(?<0,05). Aeine?aeaeaiiy ?ai?iaeoeoeaieo yeinoae eio??a oa naeiiiaoie c ??cieie eiia?iaoe?yie aaoa?iii?oieo aiiieia?a aeyaeee, ui oaa?eie c i?aeaeuaiith aa??aaaeuei?noth no?oeoo?iiai N-aaoa?io?iiaoeio oa?aeoa?ecothoueny a?eueo aenieith aeeoo?caeaoi?noth iauaaee?a. E?euee?noue ii?inyo o 2 i?nyoe? aoea a??ia?aeii aeuith o naeiiiaoie c ae?aaeaiei N-aaoa?iii?o?ciii o?iiiniiieo ia? (Oaae.3), i?aeaeuaiiy caa?aaeaiino? i?eieiaeo neeaaeaei 15,8%. Oaaeeoey 3. Iniiai? iieacieee ?ai?iaeoeoeaii? ooieoe?? naeiiiaoie o ca’yceo c N-aaoa?iii?o?ciii ae?ioeaio?e/ieo o?iiinii ?n,Nooi?iue aaoa?i-Aaaaoi-O 2 i?nyoe?A?oieaie?aii?o?cio, aae.Ie?aeaey, aie?ae?euee?noue ii?inyo, aie?acaa?aaeai?noue i?eieiaeo, %1180-19,40,547,20,40*71,83,80**292-310,40,418,80,47*84,13,40***- a oe?e oa ianooiieo oaaeeoeyo iaeiaeia? noia?ne?eiioe iathoue cia/aiiy, ui a?ae??ciythoueny c a??ia?aei?noth ia iaio 95%. Eio?e, a aaiii? yeeo aeyaeai? no?oeoo?ii iaiaeii??aei? aiiieiae, oaeiae oa?aeoa?ecoaaeenue i?aeaeuaiiyi caa?aaeaiino? i?eieiaeo (Oaae. 4), i?e oeueiio oa?oeeuei?noue aeinyaaea ??aiy 82,4%. Oaaeeoey 4. Iniiai? iieacieee ?ai?iaeoeoeaii? ooieoe?? eio??a o ca’yceo c N-aaoa?iii?o?ciii ae?ioeaio?e/ieo o?iiinii ?n,Nooi?iue aaoa?i-Aaaaoi-O 2 i?nyoe?a?oieaie?aii?o?cio, aae.Ie?aeaey, aie?ae?euee?noue ii?inyo, aie?acaa?aaeai?noue i?eieiaeo, %1100-19,90,297,80,4078,62,88*2112-39,60,418,40,2886,51,84* A ?acoeueoao? aiae?co ??aiy o?iiiniiii? ianoaa?eueiino? o ca’yceo c N-aaoa?iii?o?ciii aiiieia?a aeyaeaii ci?io ni?aa?aeiioaiiy no?oeoo?ieo oa aaiiiieo aaa?aoe?e ye o eio??a, oae ? o naeiiiaoie. (oaae. 5, 6)Iayai?noue o aaiii? o?iiiniiieo ia? c ae?aaeaiei no?oeoo?iei N-aaoa?iii?o?ciii noi?iaiaeaeoaaeanue cieaeaiiyi /anoioe aaiiiieo iooaoe?e (?<0,05). Oaaeeoey 5 Oeeoiaaiaoe/ia oa?aeoa?enoeea naeiiiaoie o ca’yceo c N-aaoa?iii?o?ciii ae?ioeaio?e/ieo o?iiinii ? n, Nooi?iue aaoa?i- ?OI,*anoioa no?oeoo?-*anoioa aaiiiieoCaaaeueiee aaea?oieaie?aii?o?cio, aae.%ieo aaa?aoe?e, %iooaoe?e, %N-aaoa?i-ii?o?cio1180-117,60,9112,12,466,50,83*2,20,20**292-317,23,5713 ,74,154,00,93*4,10,53** Oaaeeoey 6 Oeeoiaaiaoe/ia oa?aeoa?enoeea eio??a o ca’yceo c N-aaoa?iii?o?ciii ae?ioeaio?e/ieo o?iiinii ? a?oie n, aie?a Nooi?iue aaoa?i- ii?o?cio, aae. ?OI, %*anoioa no?oeoo?- ieo aaa?aoe?e, %*anoioa aaiiiieo iooaoe?e, %Caaaeueiee aae N-aaoa?i-ii?o?cio1100-116,02,564,30,88*11,82,19*1,90,32**2112-316,23,191 1,82,905,41,08*3,50,35**Aiae?c niaeo?o niiioaiieo iooaoe?e iieacaa i?iii?oe?eia ci?ithaaiiy an?o oei?a o?iiiniiieo ii?ooaiue, ca aeiyoeii aaa?aoe?e o?iiaoeaeiiai oeio, /anoioa yeeo iii?oii ia a?ae??ciyeany . A ?acoeueoao? aeine?aeaeaiue aecia/aii, ui eio?e oa naeiiiaoee c N-aaoa?iii?o?ciii aiiieia?/ieo o?iiinii aeua 2-io aae?a oa?eoa?ecoaaeenue i?aeaeuaiiyi caa?aaeaiino? i?eieiaeo. O naeiae, ye? iaee no?oeoo?ii iaiaeii??aei? aiiieiae ae?ioeaio?e/ieo ia?, aeyaeaii cieaeaiiy ??aiy aaiiiieo iooaoe?e, ye? iaaaoeaii aieeaathoue ia a?aeoai?thaaeuei? yeino? naeiae. Io?eiai? ?acoeueoaoe na?ae/aoue, ui no?oeoo?iee iie?ii?o?ci ii N-aeie?a aaoa?io?iiaoeio iiaea aooe iaeiei c oaeoi??a, ye? iaoiiaeththoue aenieo aeeoo?caeaoi?noue iauaaee?a. Oa?aeoa?enoeea N-iie?ii?o?cio o?iiinii ye neeaaeiaa /anoeia oeeoiaaiaoe/iiai oanooaaiiy naeiae iiaea aeei?enoiaoaaoenue i?e ?ic?iaoe? oeeoiaaiaoe/ieo e?eoa???a ?ai?iaeoeoeaii? caeaoiino? naeiae. 5. Aa??aaaeuei?noue N-aaoa?io?iiaoeiiaeo ?aeii?a o?iiinii oa ianoaa?euei?noue aaiiio a eaeeioeeoao e?ia? naeiae. Aeey aea/aiiy ca’yceo N-iie?ii?o?cio o?iiinii c ianoaa?euei?noth aaiiio naeiae i?iaaaeaii aiae?c /anoioe oa niaeo?o niiioaiieo iooaoe?e iin??a iie?ii?oieo N-aa??aio?a o?iiinii 13, 15, 16 oa 18. ?iciiae?e oei?a o?iiiniiieo ii?ooaiue i?aaenoaaeaiee ia iae. 3 oa 4. I?e iayaiino? o aaiii? aa??aioo o?iiinii 13 c aaeeeei N-aeieii niinoa??aaee cieaeaiiy ??aiy aaiiiieo iooaoe?e, yea a?aeaoaaeiny ca ?aooiie ciaioaiiy /enea iie?iei?aeieo ee?oei. I?aeaeuaiiy e?eueeino? N-aaoa?io?iiaoeio ia o?iiinii? 18 noi?iaiaeaeoaaeinue i?aeaeuaiiyi /anoioe no?oeoo?ieo aaa?aoe?e, ui iiaea aooe iia’ycaii c iaaeiaaeeia?noth o?iiiniiieo ii?ooaiue o N-aaoa?io?iiaoei? (?<0,05). I?e no?oeoo?i?e aaoa?iaaiiino? aiiieia?/ieo o?iiinii oaeiae aeyaeee iaai? ci?ie niaeo?o iooaoe?e. Ae?aaeaiee N-aaoa?iii?o?ci aiiieia?a 15-i? ia?e noi?iaiaeaeoaaany ye cieaeaiiyi ??aiy aaiiiieo iooaoe?e, oae ? i?aeaeuaiiyi ??aiy no?oeoo?ieo aaa?aoe?e, iaoiiaeaiiai o?aaiaioaoe??th o?iiinii (?<0,05). Cieaeaiith /anoioith aaiiiieo iooaoe?e oa?aeoa?ecoaaeeny iin?? aaoa?iii?oieo aiiieia?a 16-i? ia?e, oiae? ye no?oeoo?ia iaiaeii??aei?noue aiiieia?a 18-i? ia?e i?aeaeuoaaea /anoioo ii?ooaiue o?iiiniiiiai iaoa??aeo. Aeyaeai? ci?ie ni?aa?aeiioaiiy oa niaeo?o iooaoe?e aoee a??ia?aeieie ia ??ai? ?<0,05. I?aeaeuaia aa??aaaeuei?noue N-aaoa?io?iioaeiiaeo ?aeii?a o?iiinii 13, 15 oa 16 noi?iaiaeaeoaaeinue cieaeaiiyi ??aiy aaiiiieo iooaoe?e, oiae? ye i?e no?oeoo?i?e aaoa?iaaiiino? o?iiiniiieo ia? 15 oa 18 i?aeaeuoaaeanue /anoioa no?oeoo?ieo aaa?aoe?e. Aecia/ai? ci?ie niaeo?o iooaoe?e na?ae/aoue, ui iie?ii?o?ci ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio iiaea, o iaai?e i???, aieeaaoe ia i?iyae niiioaiieo iooaoe?e ia ee?oeiiiio ??ai?, i?e oeueiio iaai? oeie aaa?aoe?e iiaeooue ye i?aeaeuoaaoenue, oae ? cieaeoaaoenue. 6. Aca?iica’ycie iie?ii?o?cio ?aeii?a i?eoeaio?ii??iiai N-aaoa?io?iiaoeio c anioe?aoe?yie ae?ioeaio?e/ieo o?iiinii o naeiae. Aeey aea/aiiy ?ie? aa??aaaeueiiai N-aaoa?io?iiaoeio o oi?ioaaii? anioe?aoe?e i?iaaee aeine?aeaeaiiy anioeaoeaii? caeaoiino? ae?ioeaio?e/ieo o?iiinii o ca’yceo c eieae?caoe??th iie?ii?oieo N-aeie?a aaoa?io?iiaoeio. A ?acoeueoao? aiae?co aecia/aii, ui e?euee?noue i?eoeaio?ii??iiai N-aaoa?io?iiaoeio i?aeoe/ii ia aieeaaea ia anioe?aoeaio iiaaae?ieo o?iiinii, i?i ui na?ae/eoue a?aenooi?noue a??ia?aeii? ??cieoe? i?ae iin?yie iie?ii?oieo N-aa??aio?a o?iiinii ca iieacieeaie /anoioe ee?oei c anioe?aoe?yie ae?ioeaio?e/ieo o?iiinii (AAO), a oaeiae o/ano? o?iiiniiieo ia? o AAO. Aeyaeaii i?aeaeuaiiy /anoioe AAO i?e iayaiino? o aaiii? no?oeoo?ii aaoa?iaaiieo aiiieia?a. Ae?aaeaiee N-aaoa?iii?o?ci aiiieia?a 16-i? ia?e noi?iaiaeaeoaaany cia/iei i?aeaeuaiiyi iieacieea /anoioe AAO c 35,5% aei 53,1% (?<0,05), i?ioa ia aeyaeaii ia?aaaaeii? o/ano? 16-i? o?iiiniie o AAO. O iin??a aaoa?iii?oii? ia?e 15 aecia/aii iiceoeai? ei?aeyoe?? i?ae caaaeueiei aaeii N-aaoa?iii?o?cio oa /anoioith AAO (r1 =0,37; P<0,01), a oaeiae e?euee?noth AAO ia ee?oeio (r2=0,30; ?<0,05). Aeyaeeiny, ui iiceoeaiee ei?aeyoeaiee ca’ycie i?ae caaaeueiei aaeii N-aaoa?iii?o?cio aiiieia?a oa /anoioith AAO ? oa?aeoa?iei oaeiae aeey an?o aeine?aeaeaieo oaa?ei (r=0,29; ?<0,01). A ?acoeueoao? aeine?aeaeaiue aecia/aii, ui no?oeoo?ia aaoa?iaaii?noue aiiieia?/ieo o?iiinii noi?iaiaeaeo?oueny i?aeaeuaiiyi ??aiy AAO, oiae? ye e?euee?noue N-aaoa?io?iiaoeio ia ia? i?yiiai aieeao ia anioe?aoeaio caeaoi?noue ae?ioeaio?e/ieo o?iiinii. Io?eiai? aeai? na?ae/aoue i?i oa, ui aa??aaaeuei?noue ?aeii?a i?eoeaio?ii??iiai aaoa?io?iiaoeio iia’ycaia c oi?ioaaiiyi AAO, aea aieea iie?ii?oieo N-aeie?a aaoa?io?iiaoeio a?aeaoaa?oueny ia ??ai? oiieeo aca?iiae?e eiiiiiaio?a aaiiio. Aea/aiiy no?oeoo?ii-ooieoe?iiaeueieo ca’yce?a ia ee?oeiiiio ??ai? iaaea? ?ioi?iaoe?th i?i ae?yeuei?noue aaiiio, a oaeiae noai?th? iniiao aeey aecia/aiiy iciae, ye? o iiaeaeueoiio iiaeooue aooe aeeth/ai? o i?ioean ni?yiiaaii? naeaeoe??. 7. Ia?ea?i? o?iiiniie oa ?o oniaaeeoaaiiy Aea/aiiy iie?ii?o?cio aaoa?io?iiaoeiiaeo ?aeii?a i?aaenoaaey? ?ioa?an c oi/ee ci?o ?nioaaiiy i?e?iaeieo o?iiiniiieo ia?ea??a. I?enooi?noue o aaiii? o?iiinii, ia?e??oaaieo iie?ii?oieie N-aeieaie aeicaieythoue eiio?iethaaoe iioiaeaeaiiy o?iiinii. Ia?ea?i? N-aeiee (4-5 aae?a) aecia/aii o naeiae ii??ae aaeeea a?ea oa eaiae?an ia o?iiiniiao 15, 16 oa 18. Aiae?c iniiaieo iieaciee?a ?ai?iaeoeoeaii? caeaoiino? iin??a ia?ea?ieo N-aa??aio?a o?iiinii ia aeyaea iaoiaaiaoe/iiai ca’yceo i?ae i?aeaeuaiiyi e?eueeino? N-aaoa?io?iiaoeio oa ?ai?iaeoeoeaiith ooieoe??th naeiae. Aeey aea/aiiy oa?aeoa?o oniaaeeoaaiiy ia?ea?ieo o?iiinii i?iaaaeaii aiae?c iie?ii?oieo N-aa??aio?a o?iiinii o iauaaee?a (n=22) eio??a oa naeiiiaoie, a aaiii? yeeo aeyaeaii ia?ea?i? N-aeiee aaoa?io?iiaoeio. Aeyaeaii, ui ?iciiae?e ia?ea?ieo N-aa??aio?a o?iiinii ia a?aeoeeyany a?ae na?aa?aeiioaiiy 1:1. Oaeei /eiii, oniaaeeoaaiiy ia?ea?ieo o?iiinii c aaeeeeie N-aeieaie aaoa?io?iiaoeio a?aeaoaaeiny ca?aeii caaaeueiiaaiaoe/iiio i?eioeeio. O iaeiiai c iauaaee?a aeyaeee aaeee? N-aeiee ia iaio aiiieiaao 16-i? ia?e, oiae? ye o?eueee eio? aoa iin??i ia?ea?iiai aa??aioo o?iiiniie 16. I?eiouaiiy i?i iioiaeaeaiiy oeueiai iauaaeea a?ae ?ioeo aaouee?a i?aeoaa?aeea ?ioiiaaiaoe/iee aiae?c. A ?acoeueoao? aeine?aeaeaiue anoaiiaeaii aenieo noaa?euei?noue ii?oieia?/ieo oei?a o?iiinii c ia?ea?ieie N-aeieaie, a oaeiae iiaeeea?noue ?o ?aeaioeo?eaoe?? i?e a?coaeueiiio ea??ioeioaaii?. Ia?ea?i? o?iiiniie iiaeooue aeei?enoiaoaaoenue ye aeiaeaoeiaee e?eoa??e i?e aecia/aii? iioiaeaeaiiy oaa?ei. AENIIAEE 1. Ae?ioeaio?e/i? o?iiiniie naeiae oa?aeoa?ecothoueny aenieith aa??aaaeuei?noth ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio. Iie?ii?o?ci ?aeii?a i?eoeaio?ii??iiai N-aaoa?io?iiaoeio o?iiinii 13, 15, 16 oa 18 iaoiiaeth? no?oeoo?io aaoa?iaaii?noue aiiieia?/ieo o?iiinii. 2. Iie?ii?o?ci N-aaoa?io?iiaoeiiaeo ?aeii?a iia’ycaiee c ooieoe?yie aaiiio naeiae ye ia ee?oeiiiio ??ai?, oae ? ia ??ai? ae?yeueiino? i?aai?cio. Ia ee?oeiiiio ??ai? i?aeaeuaia aa??aaaeuei?noue N-aaoa?io?iiaoeia aiiieia?/ieo o?iiinii oa?aeoa?eco?oueny ci?iith ni?aa?aeiioaiiy no?oeoo?ieo oa aaiiiieo iooaoe?e o eaeeioeeoao e?ia?, oiae? ye ca’ycie c ooieoe?yie i?aai?cio i?iyaey?oueny o ??ai? ?ai?iaeoeoeaii? ooieoe?? naeiae. No?oeoo?ia aaoa?iaaii?noue aiiieia?a ae?ioeaio?e/ieo o?iiinii ii N-aaoa?io?iiaoeio o eio??a oa naeiiiaoie iia’ycaia c i?aeaeuaiiyi caa?aaeaiino? i?eieiaeo ia 10-15%. 3. Iie?ii?o?ci ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio aca?iiiia’ycaiee c i?iyaaie niiioaiieo o?iiiniiieo iooaoe?e o eaeeioeeoao e?ia? naeiae. I?aeaeuaia aa??aaaeuei?noue N-aaoa?io?iiaoeio ia o?iiiniiao 13, 15 oa 16 noi?iaiaeaeo?oueny cieaeaiiyi ??aiy aaiiiieo iooaoe?e, oiae? ye no?oeoo?iee iie?ii?o?ci o?iiiniiieo ia? 15 oa 18 iia’ycaiee c i?aeaeuaiiyi /anoioe no?oeoo?ieo aaa?aoe?e. 4. No?oeoo?ia aaoa?iaaii?noue aiiieia?a ae?ioeaio?e/ieo o?iiinii ii N-aeieao aaoa?io?iiaoeio ni?ey? oi?ioaaiith anioe?aoe?e AAO (r=0,29; P<0,01). Aeyaeaii ca’ycie N-aaoa?iii?o?cio 16-i? ia?e c i?aeaeuaiiyi /anoioe AAO (P<0,05), a oaeiae iiceoeaio ei?aeyoe?th caaaeueiiai aaea N-aaoa?iii?o?cia c iieacieeaie anioe?aoeaii? caeaoiino? o?iiinii (r1=0,37; P<0,01; r2=0,30; P<0,05) o iin??a aaoa?iii?oieo aiiieia?a 15-i? ia?e. 5. Naei? ii??ae aaeeea a?ea oa eaiae?an i?aeoe/ii ia a?ae??ciythoueny ca /anoioith iie?ii?oieo aa??aio?a o?iiinii 13, 15, 16 oa 18, iacia/ia ??cieoey o /anoio? ie?aieo N-oei?a o?iiinii 16 oa 18 iiaeooue na?ae/eoe i?i iaeiie/aiiy iaaieo o?iiiniiieo aa??aio?a o naeaeoe?iiiaaieo a?oiao oaa?ei. A?aenooi?noue cia/ii? ??cieoe? o aa??aaaeueiino? ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio i?ae eio?aie oa naeiiiaoeaie aeaio ii??ae aeicaieythoue aaaaeaoe N-iie?ii?o?ci ae?ioeaio?e/ieo o?iiinii iaeo?aeueiith oa?aeoa?enoeeith ye o a?aeiioaii? aei noao? naeiae, oae ? a ii?iaeiiio aniaeo?. 6. Ia?ea?i? N-aeiee aaoa?io?iiaoeio ? noaa?eueiei no?oeoo?ii-ii?oieia?/iei eiiiiiaioii o?iiiniie, oa?aeoa? oniaaeeoaaiiy ia?ea?ieo o?iiinii o naeiae oa?aeoa?eco?oueny ni?aa?aeiioaiiyi 1:1, ui a?aeiia?aea? caaaeueiiaaiaoe/iiio i?eioeeio. 7. A?coaeueiee aiae?c N-iie?ii?o?cio o?iiinii naeiae c aeei?enoaiiyi e?eoa???a iai?ae?euee?nii? ioe?iee aeicaiey? ca 5-aaeueiith oeaeith aecia/aoe a?aeiini? ?ici??e N-aeie?a aaoa?io?iiaoeio ia ae?ioeaio?e/ieo o?iiiniiao 13-18. 8. Oaeineiiaeaii iaoiae aeeoa?aioe?eiiai N-caaa?aeaiiy o?iiinii naeiae. Aeei?enoaiiy iiaeeo?eaoe?? N-iaoiaeo, iniiaieie aoaiaie yei? ? ia?iaea a?ae?ieneaeii aa??th (500N) oa ?ieoaaoe?y o aooa?ii-nie?oiaiio ?ic/ei? SSC i?e pH=8,5, aeiaiey? io?eioaaoe N-caaa?aeaiiy ia o?iiiniiao naeiae aac cia/ieo ii?ooaiue o?iiiniiiiai iaoa??aeo I?AEOE*I? I?IIICEOe?? 1. I?e a?coaeueiiio aiae?c? N-iie?ii?o?cio o?iiinii naeiae aeioe?eueii aeei?enoiaoaaoe e?eoa??? iai?ae?euee?nii? ioe?iee iie?ii?oieo N-aeie?a aaoa?io?iiaoeio ia ae?ioeaio?e/ieo o?iiiniiao 13-18. 2. O?aoiaoth/e aenieo aoaeoeai?noue oa iiaeeea?noue aaoiiaoecaoe??, i?e i?iaaaeaii? oeeoiaaiaoe/iiai aiae?co o?iiinii n?euenueeiainiiaea?nueeeo oaa?ei i?iiiio?oueny aeei?enoiaoaaoe i?ia?aiii-aia?aoiee eiiieaen aeey ea??ioeioaaiiy. 3. Aeey io?eiaiiy ye?niiai N-caaa?aeaiiy ia o?iiiniiao naeiae aeioe?eueii aeei?enoiaoaaoe iiaeeo?eaoe?th N-iaoiaeo, yea aeeth/a? ia?iaeo i?aia?ao?a a?ae?ieneaeii aa??th oa aooa?ii-nie?oiaei ?ic/eiii SSC i?e pH=8,5, ui caaacia/o? iiaeeea?noue i?iaaaeaiiy oeeoiaaiaoe/iiai aiae?co o naeia?noa?. 4. Oa?aeoa?enoeea N-iie?ii?o?cio o?iiinii ye neeaaeiaa /anoeia oeeoiaaiaoe/iiai oanooaaiiy naeiae ne?ae aeei?enoiaoaaoe o eiioaeno? i?ia?ai ii oaeineiiaeaiith a?aeoai?thaaeueii? caeaoiino? e?i?e oa ?iaeei naeiae. NIENIE IAOEIAEO ?IA?O CA OAIITH AeENA?OAOe?? 1.?innioa A.E., Eaaoo?a I.I. Acaeiinaycue N-iieeii?oecia o?iiinii n aini?iecaiaeeoaeueiuie ea/anoaaie naeiae //Iao/ii-oao. athe.. Einoeooo aeeaioiiaiaenoaa OAAI. – Oa?ueeia, 1996. - Ni 72. – N. 39-43. 2. ?aaoo?a I.I. Aa??aaaeuei?noue ?aeii?a i?eoeaio?ii??iiai aaoa?io?iiaoeio oa niiioaii? o?iiiniii? iooaoe?? o eaeeioeeoao e?ia? naeiae//Iao/ii-oao. athe.. Einoeooo aeeaioiiaiaenoaa OAAI. – Oa?ueeia, 1998. - Ni 73. – N. 27-30. 3. ?aaoo?a I.I. Aea/aiiy aca?iica’yceo no?oeoo?iiai iie?ii?o?cio o?iiinii c ?ai?iaeoeoeaiith ooieoe??th naeiae// A?niee: I?iaeaie cii?iaeaia??? oa aaoa?eia?ii? iaaeeoeeie. Iaoeiai-oai?. aeo?iae OCAE – Oa?ueeia, 1998. - Aei.3(27).-N.50-52. 4. Ai?iaeiaa I.I., ?o?iaiei E.E., ?icnioa A.?. Aea/aiiy o?iiiniiiiai iie?ii?o?cio o naeiae//Oace aeiiia. -i? I?aeia?. iaoeiai-i?aeo. eiio. “Naeaeoe?eii-a?ioaoiieia?/i? iaoiaee aeei?enoaiiy aaiaoe/iiai iioaioe?aeo n.-a. oaa?ei”.- Ee?a: ?inoeooo ?icaaaeaiiy oa aaiaoeee oaa?ei OAAI. – 1994.- N. 20. 5. Ai?iaueaaa I.I., ?innioa A.E., Aoeiaiei E.E. Iieeii?oeci N-aeieia o?iiinii naeiae// Oace aeiiia. I?aeia?. eiio. ”Oeyoe i?aeaeuaiiy ae?iaieoeoaa oa iie?ioaiiy yeino? naeieie”. – Oa?e?a: ?O OAAI. – 1995. – N. 33. 6. Ai?iaueaaa I.I., ?innioa A.E., Oeaeiaaa A.A. I?eiaiaiea oaeaaeceiiiiai eiiieaena i?e aiaeeca ecia?aaeaiee n i?aia?aoia o?iiinii // Oace aeiiia. I?aeia?. eiio.”Oeyoe i?aeaeuaiiy ae?iaieoeoaa oa iie?ioaiiy yeino? naeieie”. – Oa?e?a: ?O OAAI. – 1995. – N. 32. 7.Oaaoia A.E., ?innioa A.E., Aoeiaiei E.E. Ai?iaueaaa I.I., ?innioa E.A. Aiciiaeiinoe eniieueciaaiey oeeoiaaiaoe/aneeo iaoiaeia a naeaeoeee naeiae i?e ?aiiai auai?a ?iaeiia/aeueieeia caaiaeneeo eeiee// Oace aeiiia. I?aeia?. eiio.”Oeyoe i?aeaeuaiiy ae?iaieoeoaa oa iie?ioaiiy yeino? naeieie”. – Oa?e?a: ?O OAAI. – 1995. – N. 17. 8. ?aaoo?a I.I.,?icnioa A.?. Aeei?enoaiiy aeeoa?aioe?eiiai N-iaoiaeo aeey iioa?aoaaiiy o?iiinii naeiae// Oacenu aeiee. ?? Iaaeaeoia?. eiio. ii iieaeoey?ii-aaiaoe/aneei ia?ea?ai aeeaioiuo. Eeaa: Einoeooo aa?iyeieiaee e aeioaoiieiaee OAAI. – 1996. - N 82. 9. ?aaoo?a I.I.,?icnioa A.?., Aacoaeee I.Ae., Oeaeiaaa A.I., Aea/aiiy iiaeeeainoae iiaeo iaoiaee/ieo i?aeoiae?a aei ioe?iee N-iie?ii?o?cio o?iiinii naeiae// Oacenu aeiee. ?? Iaaeaeoia?. eiio. ii iieaeoey?ii-aaiaoe/aneei ia?ea?ai aeeaioi?o. Eeaa: Einoeooo aa?iyeieiaee e aeioaoiieiaee OAAI. – 1996. - N 83. 10. ?icnioa A.?., ?o?iaiei E.E., ?aaoo?a I.I. N-aeiee aaoa?io?iiaoeio ye i?e?iaei? o?iiiniii? ia?ea?e// Oacenu aeiee. ?? Iaaeaeoia?. eiio. ii iieaeoey?ii-aaiaoe/aneei ia?ea?ai aeeaioiuo. Eeaa: Einoeooo aa?iyeieiaee e aeioaoiieiaee OAAI. – 1996. - N 86. 11. ?icnioa A.?., ?aaoo?a I.I., ?o?iaiei E.E., A?aeaiei I.I. Aca?iica’ycie N-aaoa?iii?o?cio aiiieia?/ieo o?iiinii c a?aeoai?thaaeueieie yeinoyie naeiae// Oacenu aeiee. ?? Iaaeaeoia?. eiio. ii iieaeoey?ii-aaiaoe/aneei ia?ea?ai aeeaioi?o. Eeaa: Einoeooo aa?iyeieiaee e aeioaoiieiaee OAAI. – 1996. - N 87. AIIOAOe?? ?aaoo?a I.I. Iie?ii?o?ci aaoa?io?iiaoeiiaeo ?aeii?a ae?ioeaio?e/ieo o?iiinii naeiae. – ?oeiien. Aeena?oaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeoaoa n?euenueeiainiiaea?nueeeo iaoe ca niaoe?aeuei?noth 03.00.15. – aaoaoeea. – ?inoeooo oaa?eiieoeoaa OAAI, Oa?e?a , 1998. Aeena?oaoe?th i?enay/aii aea/aiith iie?ii?o?cia ?aeii?a no?oeoo?iiai aaoa?io?iiaoeio ia o?iiiniiao naeiae, a oaeiae ca’yceo aa??aaaeueiino? N-aaoa?io?iiaoeio c ?ai?iaeoeoeaieie ooieoe?yie oaa?ei. Aecia/aii, ui iin?? no?oeoo?ii aaoa?iaaiieo ii N-aaoa?io?iiaoeio aiiieia?/ieo o?iiinii, oa?aeoa?ecoaaeenue i?aeaeuaiith aeeoo?caeaoi?noth iauaaee?a. Ia ee?oeiiiio ??ai? aeyaeaii ca’ycie no?oeoo?iiai N-iie?ii?o?cio ae?ioeaio?e/ieo o?iiinii c ianoaa?euei?noth aaiiia a eaeeioeeoao e?ia? naeiae. I?aeaeuaia aa??aaaeuei?noue N-aaoa?io?iiaoeio iia’ycaia c ?aae?caoe??th iaaieo oei?a o?iiiniiieo ii?ooaiue, a oaeiae c oi?ioaaiiyi anioe?aoe?e o?iiinii. ?ic?iaeaii iaoiae oa iaeaaeiaiiy aeey aeine?aeaeaiiy o?iiinii eiii’thoa?iei niiniaii. Aeey a?coaeueiiai aiae?ca N-iie?ii?o?cio ae?ioeaio?e/ieo o?iiinii naeiae cai?iiiiiaaii e?eoa??? iai?ae?euee?nii? ioe?iee iie?ii?oieo N-aeie?a ca 5-aaeueiith oeaeith. ?ic?iaeaii iiaeeo?eaoe?th N-iaoiaeo, yea caaacia/o? noaa?e?caoe?th o?iiiniiiiai iaoa?eaeo i?e N-caaa?aeaii?. Eeth/ia? neiaa: no?oeoo?iee aaoa?io?iiaoei, aaoa?iii?o?ci aiiieia?/ieo o?iiinii, ianoaa?euei?noue aaiiio, aaa?aoe??, anioe?aoe?? ae?ioeaio?e/ieo o?iiinii, a?aeoai?thaaeueia caeaoi?noue naeiae. Eaaoo?a I.I. Iieeii?oeci aaoa?io?iiaoeiiauo ?aeiiia ae?ioeaio?e/aneeo o?iiinii naeiae. – ?oeiienue. Aeena?oaoeey ia nieneaiea iao/iie noaiaie eaiaeeaeaoa naeueneioicyenoaaiiuo iaoe ii niaoeeaeueiinoe 03.00.15. – aaoaoeea. – Einoeooo aeeaioiiaiaenoaa OAAI, Oa?ueeia , 1998. Aeena?oaoeey iinayuaia eco/aieth iieeii?oecia ?aeiiia no?oeoo?iiai aaoa?io?iiaoeia ia o?iiiniiao naeiae, a oaeiae acaeiinayce aa?eaaaeueiinoe N-aaoa?io?iiaoeia n ?ai?iaeoeoeaiuie ooieoeeyie aeeaioiuo. Onoaiiaeaii, /oi iineoaee no?oeoo?ii aaoa?iaaiiuo ii N-aaoa?io?iiaoeio aiiieiae/ieo o?iiinii, oa?aeoa?eciaaeenue iiauoaiiie aeecianiiniaiinoueth iioiieia. Ia eeaoi/iii o?iaia auyaeaia naycue no?oeoo?iiai N-iieeii?oecia ae?ioeaio?e/aneeo o?iiinii n ianoaaeeueiinoueth aaiiia a eaeeioeeoao e?iae naeiae. Iiauoaiiay aa?eaaaeueiinoue N-aaoa?io?iiaoeia naycaia n ?aaeecaoeeae ii?aaeaeaiiuo oeiia o?iiiniiiuo ia?ooaiee, a oaeaea n oi?ie?iaaieai anioeeaoeee o?iiinii. ?ac?aaioai iaoiae e iai?oaeiaaiea aeey enneaaeiaaiey o?iiinii eiiiuethoa?iui niiniaii. Aeey aecoaeueiiai aiaeeca N-iieeii?oecia ae?ioeaio?e/aneeo o?iiinii naeiae ?aeiiaiaeoaony eniieueciaaoue e?eoa?ee iieoeiee/anoaaiiie ioeaiee iieeii?oiuo N-aeieia ii 5-aaeueiie oeaea. ?ac?aaioaia iiaeeoeeaoeey N-iaoiaea, iaania/eaathuay noaaeeecaoeeth o?iiiniiiiai iaoa?eaea i?e N-ie?aoeaaiee. Eeth/aaua neiaa: no?oeoo?iue aaoa?io?iiaoei, aaoa?iii?oeci aiiieiae/iuo o?iiinii, ianoaaeeueiinoue aaiiia, aaa??aoeee, annioeeaoeee ae?ioeaio?e/aneeo o?iiinii, aini?iecaiaeeoaeueiay niiniaiinoue naeiae. Ivashura M.N. Polymorphism of heterochromatin regions of the pig acrocentric chromosomes.- Manuscript. Dissertation for the degree Candidate Science (Agriculture) in speciality 03.00.15.-The Institute of Animaj Science of UAAS, Kharkov, 1998. The dissertation deals with studying the polymorphism of structural heterochromatin regions on porcine chromosomes and C-heterochromatin variability relationship with the animal reproductive functions. It was established that carriers of the chromosomes structurally heterogeneous in C-heterochromatin were characterized by the progeny improved viability. The relatioship of the structural C-polymorphism of acrocentric chromosomes with genome instability in porcine blood leucocytes was found at the cell level. The incrized C-heterochromatin variability is connected to realization of certain chromosome aberrations and formation of associations. A method and equipment are developed to investigate chromosomes using a computer. For visual analysis of the C-polymorphism of porcine acrocentric chromosomes, criteria of semiquantitative evaluation of polymorphous C-blocks according to 5-point scoring were proposed. The C-method was modified to provide chromosome stabilization at C-staining. Key word: constitutive heterochromatin, heteromorphism of gomologous chromosomes, genome instability, aberration, association of acrocentric chromosomes, reproductive ability of pigs. Aaoi?aoa?ao A?aeiia?aeaeueiee ca aeione A.?.Oaaoia i?aeienaii aei aeioeo 17 eenoiiaaea 1998 ?. Ianya ae.a. Oe?aae 100 aec. Caiiaeaiiy ?495 Ae?eyiea iia?aoeaiiai ae?oeo ?inoeoooo oaa?eiieoeoaa OAAI 312120, Oa?e?a, i/a Eoeeie/?

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