Морфологія вилочкової залози людини в пренатальному періоді онтогенезу: Автореф. дис… канд. біол. наук / О.В. Благодарова, Київ. ун-т ім. Т.Шевченка

EE?ANUeEEE OI?AA?NEOAO ?iai? OA?ANA OAA*AIEA

AEAAIAeA?IAA Ieaia Aieiaeeie??aia

OAeE [ 611.438 + 612.638 ] : 57.017.64.642

II?OIEIA?ss AEEI*EIAI? CAEICE ETHAeEIE

A I?AIAOAEUeIIIO IA??IAe? IIOIAAIACO

14.03.09 — a?noieia?y, oeeoieia?y, aia??ieia?y

Aaoi?aoa?ao aeena?oaoe?? ia caeiaoooy iaoeiaiai nooiaiy

eaiaeeaeaoa a?ieia?/ieo iaoe

EE?A — 1999

Aeena?oaoe?y ? ?oeiien

?iaioa aeeiiaia a Iaoe?iiaeueiiio iaaeaaia?/iiio oi?aa?neoao? ?i.
I.I. Ae?aaiiaiiaa I?i?noa?noaa ina?oe Oe?a?ie

Iaoeiaee ea??aiee: Caneoaeaiee ae?y/ iaoee oa oaoi?ee Oe?a?ie,
aeaaeai?e E?einueei? AI, aeieoi? a?ieia?/ieo iaoe, i?ioani?

OIIAIEI Ai?en A?eai?iae/

Iaoe?iiaeueiee iaaeaaia?/iee oi?aa?neoao ?i. I.I.Ae?aaiiaiiaa,
caa?aeoaa/ eaoaae?e aiaoii??, ?icaeoeo oa a?a??ie ae?oae

Io?oe?ei? iiiiaioe:

aeieoi? iaaee/ieo iaoe, i?ioani? AI?Ae??IEI Aaaeei Iaeneiiae/,
Ee?anueeee oi?aa?neoao ?i. Oa?ana Oaa/aiea, i?ioani? eaoaae?e
oeeoieia??, a?noieia?? oa a?ieia?? ?icaeoeo

aeieoi? a?ieia?/ieo iaoe, i?ioani? NOA*AIEI Ethaeieea Ieaenaiae??aia,
Iaoe?iiaeueiee iaaee/iee oi?aa?neoao ?i. I.I.Aiaiiieueoey, caa?aeoth/a
eaai?aoi???th aeaeo?iiii? i?e?ineii?? Iaoeiai-aeine?aeiiai eaai?aoi?iiai
oeaio?o

I?ia?aeia onoaiiaa:

Oa?iii?euenueea aea?aeaaia iaaee/ia aeaaeai?y ?i.?.ss.Ai?aa/aanueeiai,
I?i?noa?noaa ioi?iie caei?ia»y Oe?a?ie, eaoaae?a a?noieia??, oeeoieia??
oa aia??ieia??.

Caoeno a?aeaoaeaoueny «16» /a?aiy 1999 ?. i 14.00 aiaeei? ia can?aeaii?
niaoe?ae?ciaaii? a/aii? ?aaee Ae 26.001.38 i?e Ee?anueeiio oi?aa?neoao?
?i Oa?ana Oaa/aiea (252022, i.Ee?a, i?. Aeooeiaa 2, a?ieia?/iee
oaeoeueoao)

C aeena?oaoe??th iiaeia iciaeiieoenue o a?ae?ioaoe? Ee?anueeiai
oi?aa?neoaoo ?i Oa?ana Oaa/aiea (252033, i.Ee?a, aoe.Aieiaeeie?nueea,
58)

Aaoi?aoa?ao ?ic?neaiee «16» /a?aiy 1999 ?.

A/aiee nae?aoa?

niaoe?ae?ciaaii? a/aii? ?aaee Aeaaeaeianueea O.E.

19

CAAAEUeHA OA?AEOA?ENOEEA ?IAIOE

Aeooaeuei?noue oaie. O no/ani?e a?ieia?? oa iaaeeoeei? aeine?aeaeaiiy
nopoeoope oa ooieoe?? aeei/eiai? caeice iaaoaathoue iniaeeaiai cia/aiiy
ipe aea/aii? ipaaio a ipaiaoaeueiiio iap?iae? iipoiaaiaco, oiio ui
e?ioi?aeia oeaieia ? iaei??th c iniiaieo aiaoiii-o?c?ieia?/ieo nenoai
ipaai?cio, a?ae noaio yei? a cia/i?e i?p? caeaaeeoue eiai aiiainoac (
Oeunoiaa C.N., 1987; Aieioei H.A.., 1990; sspeeei A.A., Iei/oe A.A.,
Apeiaae/ TH.A., 1991; Eiiennapaiei N.A., Cae E.E., Iaioeeia I.O. e aep.
1994; Eoc?a I.Y., 1971; Boyd R., et. al. 1991; Pribyl T. et. al.,1996).

sse a?aeiii, aeei/eiaa caeica ? iaeiei c iaeaaaeeea?oeo ipaai?a
?ioiiaaiaco, yeee caaacia/o? ?nioaaiiy p?ciiai oeio paaeoe?e ?ioiii?
nenoaie a aioa- oa iinoiaoaeueiiio iap?iaeao picaeoeo ipaai?cio. ?acii c
oei, caeeoathoueny iaaeinoaoiuei aea/aieie caaaeuei? caeiiii?piino?
iapapiciiae?eo ee?oeiieo aeaiaio?a caeice ia p?cieo aoaiao
aioop?oiueioopiaiiai picaeoeo ieiaea, a?aenooi? eiiieaeni?
aeine?aeaeaiiy oapaeoapo i?aeee?oeiieo aca?iica’yce?a oa ee?oeiieo
opainoipiaoe?e, iaei picpiaeai? ieoaiiy, ui noinothoueny iipoieia?/ieo
iniaeeainoae iepaieo iiioeyoe?e e?ioioeeo?a oa paoeeoeiai?oae?ioeeo?a a
p?ci? iap?iaee ipaaiiaaiaco.

A aeine?aeaeaiiyo inoaii?o pie?a iienai? iaoai?cie noaiiaeaiiy
e?ioi?aeii? nenoaie, ipine?aeeiaaia aeeiai?ea picaeoeo e?ioioeeo?a,
aea/aii oipioaaiiy ?ioiieia?/ii? oieapaioiino? a iipi? oa ipe aioeaaii?e
noeioeyoe??, ui aeaei iiaeeea?noue iaaeeceoenue aei aep?oaiiy pyaeo
ieoaiue, ye? iathoue aaaeeeaa iaoeiai-ipaeoe/ia cia/aiiy (Aiapee E.E.,
Oaa/aiei A.A., *apeania A.A., 1991; Aieiiaa A.A., Apeaipaiei Ae.A.,
Apioaaaa E.I., 1993; ?aaiia I.A., 1996; Watt P.C. et. al. 1994).

Anoaiiaeaii, ui oipioaaiiy ?ioiii? nenoaie o ethaeeie picii/eia?oueny a
paii?e iap?iae aioop?oiueioopiaiiai picaeoeo ieiaea, a i?ney 12 oeaeiy
e?ioioeeoe caeice noathoue ooieoe?iiaeueii a?eueo aeoeaieie, iaaoaath/e
oapaeoapieo oeueopanopoeoopieo iciae. Iinooiiai aeei/eiaa caeica noa? ia
o?eueee aeaeapaeii e?ioioeeo?a (o?iioeeo?a), aea e paaoeyoipii aaiaoe/ii
iaoiiaeaiiai ?o aeicp?aaiiy, aeeoapaioe?eiaee oa i?apaoe?? a iapeoap?ei?
a?aeae?ee ?ioiii? nenoaie.

A e?oapaoop? inoaii?o pie?a ana /ano?oa c’yaeythoueny aeaii?, ui
na?ae/aoue ipi iaiao?aei?noue iiaeaeueoiai, a?eueo aeeaieiai aea/aiiy
caeice a ipaiaoaeueiee iap?iae iipoiaaiaco. Oea iia’ycaii c oei, ui c
iaeiiai aieo, ipiaaaeaiiy iiae?aieo aeine?aeaeaiue nipey? oipioaaiith
no/anieo oyaeaiue ipi iniaeeaino? ?ioiiaaiaco ieiaea a i?aeopeiaii?
aiiainoaco ipaai?cio, yeee picaeaa?oueny, c ?ioiai — i?aeepaneth?
iaiao?aei?noue picpiaeee no/anieo iaoiae?a ipio?eaeoeee oa e?eoaaiiy
?ioiiiaeao?oeeo?a, a oaeiae ae?aaiinoeoe? oa iiiapaaeaeaii? iiyae
oaeoip?a, ye? ipecaiaeyoue aei picaeoeo ?ioiieo eiioe?eo?a i?ae
ipaai?ciii iaoap? oa ieiaea.

Inoaii?i /anii ipiaaaeaii pyae aeine?aeaeaiue, a yeeo iieacaia
iaiaeiip?aei?noue ee?oeiiiai neeaaeo paoeeoeiai?oae?ioeeo?a (Oeunoiaa
C.N. Eaeeieia E.E., Oaaeinii A.O. 1984; Shortman S., 1984; Boday B.,
Keisez H., 1977) aeyaeai? oeyoe ?? epiaiiinoa/aiiy oa ?iiapaaoe??
(*apeania A.A., 1979, Hacapia Ae.?., 1993), iienai? iniaeeaino?
iipoiaaiaco a paiiueiio iinoiaoaeueiiio iap?iae? (?aaiia I.?., 1996).
Ciaeoea iaapoiooaaiiy oaeiae ? oi/ea cipo ipi aeioe?euei?noue aea/aiiy
iipoieia?? aeei/eiai? caeice ca ciiaie: noaeainoeypia, aeania eipeiaee
oap, eipoeei-iaaeoeypia ciia oa iiceiaa pa/iaeia (Naiei I.?., 1987).
Iaeiae, nenoaiaoe/i? aeine?aeaeaiiy ii aiae?co iipoieia?? p?cieo cii
ipaaio, oaii?a ?o oipioaaiiy, a oaeiae iaoai?ci?a aenaeaiiy e?ioioeeo?a
ca iaae? caeice a p?ci? iap?iaee aioop?oiueioopiaiiai picaeoeo ieiaea ?
iaei/enaeueieie ? ia iinyoue ocaaaeueioth/iai oapaeoapo.

Oiio ana aeua cacia/aia aeeeeea?, aacnoii?aii, ?ioapan ia o?eueee o
a?ieia?a, aea e o e?eap?a, oiio ui a oeaiop ipiaeaie noa? aea/aiiy
nopoeoope oa ooieoe?? aeei/eiai? caeice ieiae?a ethaeeie a
ipaiaoaeueiiio iap?iae? iipoiaaiaco.

?acii c oei, caeeoathoueny iaaeinoaoiuei picpiaeaieie pyae ieoaiue, ui
noinothoueny iioiaeaeaiiy o?iioeeo?a, oeyo?a ?o aoiaeo oa aeoiaeo c
caeice, a oaeiae iiaeeea? iaoai?cie oeueiai ipioeano.

Iiopaaothoue iiaeaeueoi? picpiaee ieoaiiy, ui noinothoueny
a?noiaia?iapo?oaeoii?ee ipaaio, iniaeeainoae oipioaaiiy eiai
ai?oae?aeueiiai capiaeea, noo? aeoeeaeaioaeueii? ?iaiethoe??,
iioiaeaeaiiy paoeeoeiai?oae?aeueieo ee?oei oa iaoai?cie ?o naepaoe??,
pieue o?i?/ieo o?eaoeue a ipaiaoaeueiiio iap?iae? iipoiaaiaco.

Caneoaiaothoue oaeiae ia oaaao ieoaiiy ipi iaoai?cie pici?ciaaaiiy
o?iioeeoaie aeanieo oa /oaeip?aeieo aioeaai?a, o/ano? e?ioioeeo?a o
oipioaaii? ?ioiieia?/ii? oieapaioiino? a ipaai?ci?, a oaeiae aea/aiiy
ooieoe?iiaeueii? iipoieia?? aaiaoi-o?i?/iiai aap’?po oa iniaeeainoae
epiaiiinoa/aiiy caeice a ipaiaoaeueiee iap?iae iipoiaaiaco.

C apaooaaiiyi cacia/aiiai ? i?aenoaae aaaaeaoe, ui ipiaeaia iipoieia??
aeei/eiai? caeice a ipaiaoaeueiiio iap?iae? ? aeeeth/ii aeooaeueiith, a
ieoaiiy ,ye? i?aeeyaathoue aep?oaiith, iathoue aaaeeeaa ia o?eueee
oaipaoe/ia, aea e ipaeoe/ia cia/aiiy.

Iaoa aeine?aeaeaiiy. Ipiaanoe eiiieaenia a?noio?i?/ia, a?noiaiceiaoe/ia
oa aeaeopiiii?epieii?/ia aeine?aeaeaiiy nopoeoope aeei/eiai? caeice
ieiaea ethaeeie oa aeyaeoe iniaeeaino? noaiiaeaiiy ? picaeoeo ??
neeaaeiaeo aeaiaio?a a ipaiaoaeueiiio iap?iae? iipoiaaiaco.

Caaaeaiiy aeine?aeaeaiiy

1. Aea/eoe caaaeuei? caeiiii?piino? noaiiaeaiiy oa picaeoeo
a?noi-aaiaoe/ieo nopoeoop caeice ieiaea ia p?cieo aoaiao eiai
aioop?oiueioopiaiiai picaeoeo.

2. Aeyaeoe iniaeeaino? iipoi-ooieoe?iiaeueieo aca?iica’yce?a iaepioaa?a,
paoeeoeiai?oae?ioeeo?a, e?ioioeeo?a oa aaiaoi-oeaetheypiiai aap’?po
aeei/eiai? caeice a ipaiaoaeueiiio iap?iae? iipoiaaiaco ieiaea.

3. Aecia/eoe iniaeeaino? aoaeiae, oiiiapao?? oa ooieoe?iiaeueii?
nipyiiaaiino? nopiiaeueii-iapaio?iaoicieo aeaiaio?a aeei/eiai? caeice oa
aea/eoe iipoi-ooieoe?iiaeuei? iniaeeaino? eiiiiiaio?a aaiaoi-o?ioniiai
aap’?po ia p?ai? “ee?oeia-aaiieai?eyp”.

4. Aeyaeoe iniiai? iipoieia?/i? iciaee naepaoiooaipthth/i?,
oaaioeeoapii? oa ae?i?iaoeaii? ooieoe?e a?noi-aaiaoe/ieo nopoeoop caeice
oa anoaiiaeoe ?o cia/aiiy a ipioeanao aeicp?aaiiy, aeeoapaioe?eiaee oa
aenaeaiiy e?ioioeeo?a ca iaae? ipaaio.

5. Aea/eoe oeueopanopoeoopio iipoieia?th oa iipoiaaiac o?i?/ieo o?eaoeue
oa c’ynoaaoe ?o pieue a ipioeanao oipioaaiiy aeao?i?oeaieo nopoeoop
caeice a aioaiaoaeueiiio iap?iae? picaeoeo ieiaea.

Haoeiaa iiaecia ?iaioe. Ha i?aenoaa? nopoeoopii-ooieoe?iiaeueiiai
aiae?co iopeiaieo aeaiieo c aeeipenoaiiyi iaepi-i?epineii?/ieo,
a?noio?i?/ieo, a?noiaiceiaoe/ieo oa aeaeopiiiii?epineii?/ieo iaoiae?a
aiapoa ipiaaaeaii eiiieaenia aeine?aeaeaiiy a?noi-aaiaoe/ieo nopoeoop
caeice a p?ci? iap?iaee aioop?oiueioopiaiiai picaeoeo ieiaea ethaeeie.

Noipioeueiaaia eiioeaioe?y ipi iipoi-ooieoe?iiaeueio aaoapiaaii?noue
paoeeoeiai?oae?y, ui aeaei iiaeeea?noue aeyaeoe iipoieia?/i? iniaeeaino?
ee?oei eipeiai? oa iiceiai? pa/iaeie, a oaeiae paoeeoeiai?oae?ioeeo?a
o?i?/ieo o?eaoeue. Ha i?aenoaa? iip?aiyeueiiai aiae?co iopeiaieo aeaiieo
picpiaeai? iipoieia?/i? epeoap?? naepaoiooaipthth/i? ooieoe?? oeeo
ee?oei a aniaeo? oyaeaiue ipi caeico ye aiaeiepeiiee ipaai , a oaeiae
iaapoioiaaii ipeiouaiiy ipi iiaeeea?noue ?nioaaiiy aioop?oiueiipaaiii?
oo?e?caoe?? naepaoa aeey caaacia/aiiy e?ioioeeoiiiaca.

Aiapoa anoaiiaeaii oapaeoap oa aeyaeai? iniaeeaino? oipiiooaipthth/eo
ipioean?a a oae caai? epeoe/i? iap?iaee aioop?oiueioopiaiiai picaeoeo
caeice. Aeaoaeueii aea/aia oeueopanopoeoopia ipaai?caoe?y oa iipoiaaiac
o?i?/ieo o?eaoeue, caipiiiiiaaia iiaa eiioeaioe?y a?aeiinii ?o pie? a
picaeoeo caeice, iopeiai? iia? aeaii?, ui nipinoiaothoue a?iioaco ipi
noaeeiia iioiaeaeaiiy oeeo nopoeoop.

Oai?aoe/ia oa i?aeoe/ia cia/aiiy ?iaioe. Ipaaenoaaeai? aeaii? ipi
caaaeuei? caeiiii?piino? noaiiaeaiiy oa picaeoie a?noi-aaiaoe/ieo
nopoeoop aeei/eiai? caeice (paoeeoeiai?oae?e, e?ioi?aeia oeaieia,
aaiaoi-o?i?/iee aap’?p) aeathoue iiaeeea?noue picpiaeoe iia? i?aeoiaee o
aea/aii? nopoeoope oeueiai ipaaio oa aeyaeoe eiai cia/aiiy a picaeoeo
?ioiii? nenoaie a aioa oa iinoiaoaeueiee iap?iaee iioiaaiaco.
?acoeueoaoe aeine?aeaeaiue a?aeiapaaeathoue ye?nii iiaee aoai o aea/aii?
ipaiaoaeueiiai iipoiaaiaco caeice, iaaeeaeath/e aei a?eueo aeeaieeo oa
aaeaeaaoieo ciaiue ipi ?? ipia?aeio pieue a ?ioiiaaiac?.

C’ynoaaiiy iniaeeainoae a?noiapo?oaeoii?ee paoeeoeiai?oae?y oa
e?ioi?aeii? oeaieie a p?ci? iap?iaee aiap?iaaiaco, aeyaeaiiy
iipoieia?/ieo iciae naepaoiooaipthth/i? ooieoe?? oeeo ee?oei,
oapaeoapenoeea epeoe/ieo iap?iae?a a oipioaaii? a?noi-aaiaoe/ieo
nopoeoop, aea/aiiy oiiiapao??, iipoieia?? oa i?apaoe?? e?ioioeeo?a ca
iaae? ipaaio, oipioaaiiy o?i?ei-aaiaoe/iiai aap’?po oa ?io? ieoaiiy, ye?
ciaeoee nai? a?aeiapaaeaiiy a pacoeueoaoao ipiaaaeaieo aeine?aeaeaiue,
i?aeepaneththoue ia o?eueee ?o oaipaoe/io cia/ei?noue. Iiaeia
ipeionoeoe, ui iaaia cia/aiiy aiie iiaeooue iaoe ? oiae?, eiee
iciaeiieaiiy c pacoeueoaoaie aeaiii? piaioe aeeee/a iaiao?aei?noue
ipiaaaeaiiy iiaeaeueoeo aeine?aeaeaiue iipoieia?? aeei/eiai? caeice a
ipaiaoaeueiee iap?iae iipoiaaiaco.

Iopeiai? aeaii? iathoue ? ipaeoe/ia cia/aiiy, oiio ui iiaeooue aooe
aeeipenoai? ipe picpiaoe? caoiae?a ipio?eaeoeee oa iaoiae?a e?eoaaiiy
p?ciiiai?oieo ?ioaeoe?eieo oa aooi?ioiieo caoaipthaaiue o ae?oae,
caiia?aaiiy aioaiaoaeueii? caaeaae? ieiaea, anoaiiaeaiiy ipe/ei
?ioiiiaeao?oeeoieo noai?a oa picpiaee niinia?a ?o eipaeoe??. Ipe oeueiio
iaiao?aeii apaoiaoaaoe, ui aeanopoeoeaii-aeenopio?/i? ipioeane a
aeei/eia?e caeic? iiaeooue noipiaiaeaeoaaoenue iipooaiiyie oipioaaiiy
ipepiaeiuei? ?ioiiieia?/ii? oieapaioiino? oa ?ioi?oaoo, ui ? ? aeania
ipe/eiith aeieeiaiiy aooi?ioiieo caoaipthaaiue.

Aipiaaaeaeaiiy pacoeueoao?a aeine?aeaeaiiy. Iaoap?aee aeenapoaoe??
aeeipenoiaothoueny a eaeoe?eiiio eopn? oa ipaeoe/iiio aeeeaaeaii? ia
eaoaaep? aiaoii??, picaeoeo oa a?a??ie Haoe?iiaeueiiai iaaeaaia?/iiai
oi?aapneoaoo ?i. I.I. Aepaaiiaiiaa, eaoaaepao a?ieia??, iipiaeueii?
aiaoii??, iipiaeueii? a?noieia?? Haoe?iiaeueiiai iaaee/iiai oi?aapneoaoo
?i. I.I.Aiaiiieueoey, eaoaaep? a?ieia?? A?iieoeueeiai iaaee/iiai
?inoeoooo, Aei?ipiiaopianueei? aeapaeaaii? iaaee/ii? aeaaeai??, eaoaaep?
a?ieia?? Ee?anueeiai oi?aapneoaoo ?i. O.A. Oaa/aiea, eaoaaep?
iipiaeueii? aiaoii??, eaoaaep? a?ieia?? *api?aaoeueeiai iaaee/iiai
oi?aapneoaoo.

Aipiaaoe?y piaioe. Iaoap?aee aeenapoaoe?? aeiiia?aeaeenue ia 1
Oepa?inueeiio eiiapan? aiaoii?a, a?noieia?a oa aiap?ieia?a (i.
?aaii-Opaie?anuee, 1995), iaoeia?e eiioapaioe?? *api?aaoeueeiai
iaaee/iiai ?inoeoooo (i. *api?aoe?, 1996 p.), iaoeiai-ipaeoe/i?e
eiioapaioe?? Aei?ipiiaopianueei? aeapaeaaii? iaaee/ii? aeaaeai?? (i.
Aei?ipiiaopianuee, 1996 p.).

Nopoeoopa oa ia’?i aeenapoaoe??. Aeenapoaoe?y iaienaia ia 141 noip.
iaoeiiieniiai oaenoo, ?ethnopiaaia 20 i?epioioi oa 21 aeaeopiiiiapaiaie,
neeaaea?oueny ?c anooio, iaeyaeo e?oapaoope, opueio aeaa aeanieo
aeine?aeaeaiue, caeeth/aiiy oa nieneo e?oapaoope, ui i?noeoue ….
a?o/eciyieo oa ….. ?iicaiieo iapoiaeaeapae.

Iniiai? iieiaeaiiy, ui aeiinyoueny ia caoeno

1. A ipaiaoaeueiiio iipoiaaiac? aeei/eiai? caeice ieiaea ethaeeie
?niothoue iepai? epeoe/i? iap?iaee picaeoeo, ui a?aeiapaaeathoue ye?nii
iia? aoaie ?? a?noiaia?iapo?oaeoii?ee: aiap?iiaeueiee iap?iae (4-7
oeaeaeaiue), aiap?iiaeueii-oaoaeueiee iap?iae (7-12 oeaeaeaiue),
oaoaeueiee iap?iae (12 oeaeaeaiue ? a?eueoa).

2. Aipiaeiaae an?o aoai?a aioop?oiueioopiaiiai picaeoeo ieiaea
paoeeoeiai?oae?e caeice oapaeoapeco?oueny nopoeoopii-ooieoe?iiaeueiith
aaoapiaaii?noth ee?oei, ui aoiaeyoue aei eiai neeaaeo.

3. O?i?ei-aaiaoe/iee aap’?p a?ae?apa? ipia?aeio pieue a
oipiiooaipthth/eo ipioeanao aeei/eiai? caeice, ye? oapaeoapecothoueny
iiyaith aeao?i?oeaieo nopoeoop: aeieueie (/anoi/ie), e?ioi?aeii?
oeaieie, aaiii?epioeepeoeyoipiiai ponea, ee?oeiieo aeaiaio?a oa ?io.

4. O?i?/i? o?eueoey yaeythoueny ia o?eueee noaoe/ieie nopoeoopaie: a
iipoiaaiac? ?o iiaeia oiiaii picp?cieoe noaae?th ?i?oe?aoe??,
noaa?e?caoe?? oa “?iaiethoe??”, ui oapaeoapecothoueny iaaieie
iipoieia?/ieie iciaeaie ee?oei a oeaiopaeuei?e, ipii?aei?e oa
iapeoap?eieo ciiao.

Iniaenoee aianie caeiaoaa/a. Aaoipii naiino?eii ipiaiaeeany iaa?p
iaoap?aeo, eiai iapiaea oa ipiaaaeaiiy a?noio?i?/ieo, a?noiaiceiaoe/ieo
oa aeaeopiiiii?epineii?/ieo aeine?aeaeaiue, a oaeiae aiae?c ?o
pacoeueoao?a oa iaienaiiy aeaa aeenapoaoe??.

IAOA??AE OA IAOIAeE AeINE?AeAEAHHss

Aea/aia aeei/eiaa caeica o 20 aiap?ii?a (5-8 oeaei?a
aioop?oiueioopiaiiai picaeoeo), a oaeiae 110 ieiae?a ethaeeie (2-9
i?nyoe?a aioop?oiueioopiaiiai picaeoeo).

Oaaeeoey 1

?iciiae?e aeine?aeaeaiiai iaoap?aeo ca a?eii

Aia??iiaeueiee

ia??iae niinoa?a-

aeaiueE?euee?noueIe?aeiee ia??iae

niinoa?aaeaiueE?euee?noue5 oeaei?a52 i?nyoe?53 i?nyoe?56 oeaei?a54
i?nyoe?205 i?nyoe?a207 oeaei?a56 i?nyoe?a157 i?nyoe?a158 oeaei?a58
i?nyoe?a159 i?nyoe?a15Onueiai20onueiai110

Iaoap?ae apaany ia aeine?aeaeaiiy a aeooapnueei-a?iaeieia?/ieo
a?aeae?eaiiyo e?eapaiue oa iieiaiaeo aoaeeieao i. Ee?aa i?ney aaipo?a oa
iapaae/anieo iieia?a ipe a?aenooiino? o aaa?oieo caoaipthaaiue ?iioiii?
ipepiaee. Iaoap?ae o?enoaaany a 12 % iaeopaeueiiio oipiae?i?,
niepoiai-oipiae?iia?e noi?o? (9:1) ? caeeth/aany a iapao?iia? aeiee ii
caaaeueiiipeeiyo?e iaoiaeeoe?.

A piaio? aeeipenoai? iaoiaee caaaeueiiiipoieia?/iiai,
iaepi-i?epineii?/iiai aiae?co oa pyae a?noio?i?/ieo paaeoe?e aeey
aeyaeaiiy oeaieiieo ipioaiae?eai?a, ioeea?iiaeo eeneio , a oaeiae
aeoeaiino? iepaieo oapiaio?a (EO-aca, EO-aca, AOO-aca) c iaoith a?eueo
aeaoaeueiiai aea/aiiy iepaieo aoai?a picaeoeo caeice a ipaiaoaeueiee
iap?iae iipoiaaiaco caeice .

Napaae caaaeueiiiipoieia?/ieo iaoiae?a aeeipenoai?: caoapaoaaiiy
aaiaoienee?iii oa aiceiii, i?epiooeneiii ii Aai-A?cii, ?iipaaiaoe?y
niieo/ii? oeaieie ii Aiiip?. Napaae a?noio?i?/ieo paaeoe?e aeeipenoaii:
OEE-paaeoe?y ii Iae-Iaiono aeey aea/aiiy oeaieiieo ipioaiae?eai?a c
iinoaiiaeith eiiopieueieo oano?a (iapiaea cp?c?a ai?eacith neeie,
nieiaeiaei ae?anoacii, aoeaoeethaaiiy oa aeaaoeaoeethaaiiy), aeey
aeyaeaiiy oa ?aeaioeo?eaoe?? eeneeo ae?eiciai?iiae?eai?a
(oiiaepi?oeinoeueoao?a oeio “A” oa “N”, a?aeopiiiai? eeneioe) iinoaaeai?
paaeoe?? iaoaopiiac?? c p?ciei pH pic/eio oieo?ae?iiaiai neiueiai (pH<3 oa pH=6,7) oa aeeipenoaiiyi eiiopieueieo oano?a (iaoeethaaiiy oa aeaiaoeethaaiiy, iapiaee cp?c?a aaeoap?aeueiith oa oanoeeoeypiith a?aeopii?aeacaie). A /anoei? niinoapaaeaiue (10 aiap?ii?a oa 45 ieiae?a) aeeipenoai? paaeoe?? c eiei?aeiei cae?cii ii Oaeeo, a oaeiae c aeueoe?aiiaei nei?i ii No?aeiaio. Aeey aea/aiiy a?eeiai-neioaoe/ii? oa ieanoe/ii? ooieoe?e iapaio?iaoicieo aeaiaio?a caeice aeeipenoai? oaeiae a?noio?i?/i? paaeoe?? aeey aeyaeaiiy peai- oa aeacien?peaiioeea?iiaeo eeneio: a?aeiia?aeii paaeoe?y Apaoa oa paaeoe?y Oaeueaaia c a?aeiia?aeieie eiiopieyie. O 37 niinoa?aaeaiiyo, c caeic, acyoeo /apac 1,5-2 aiaeeie a?ae caaeaae? ieiaea, aiooaaeenue ep?inoaoi? cp?ce oiaueiith 10 iei, a yeeo iaoiaeaie Aiiip? oa Aaoooaeia-Iaecaey aeyaeyee eeneo oa eoaeio oinoaoace (EO-aca, EO-aca) oa AOO-aco. Ipi aeoeai?noue oapiaio?a piaeee aeniiaee a caeaaeiino? a?ae ai?noo noanopao-oapiaioieo e?ioeaaeo ipiaeoeo?a paaeoe?? a i?noeyo eieae?caoe?? oapiaio?a. An? aeua cacia/ai? a?noio?i?/i? paaeoe?? oa eiiopieuei? oanoe aei ieo acyo? ca ipea?iaeueiei ipiienii ?c iiiiapao?e Lillie R. (1969), Pearse A. (1962). C iaoith a?eueo aeaoaeueiiai aea/aiiy nopoeoope eiiiiiaio?a aeei/eiai? caeice oa oiieeo iaoai?ci?a oipioaaiiy a?noi-aaiaoe/ieo nopoeoop ia p?cieo aoaiao aioop?oiueioopiaiiai picaeoeo ieiaea o 49 aeiaaeeao ipiaaaeaii aeaeopiiiii?epineii?/ia aeine?aeaeaiiy o 12 aiap?ii?a (5-8 oeaei?a aioaiaoaeueiiai picaeoeo oa 37 ieiae?a ethaeeie (2-9 i?n. aioop?oiueioopiaiiai picaeoeo). Iaoap?ae caaepaany ia aeine?aeaeaiiy a iaeneiaeueii eipioeee oapi?i i?ney niapo? (1-2 aiaeeie), eoni/ee pici?pii 0,5-1,0 ii o?enoaaeenue a 3,6 % pic/ei? aeooapaeueaeaa?aeo ia oinoaoiiio aooap?. Aeaa?aepaoaoe?y oeaieie ipiaiaeeeanue a niepoao cpinoath/i? eiioeaiopaoe??. Iaacaiaeai? aeiee iii?uaee a noi?o aoeaoiio c apaeaeeoii (1:1) ia aea? aiaeeie, i?ney /iai caeeth/aee a cacia/aio niieo. Aeaioiaeai? ia oeueopai?epioii? LEA-8810 ??? cp?ce oiaueiith 40-50 ii eiiopanooaaee opai?eaoeaoaoii oa i?opaoii naeioeth ca iaoiaeii ?aeiieueaen, aeine?aeaeoaaee oa oioiapaooaaee a aeaeopiiiiio i?epineii? OAIA-100 AE. Iiae?aia eiiieaenia aeine?aeaeaiiy ? aeinoaoiuei ?ioipiaoeaiei aeey aea/aiiy iniaeeainoae oipiiooaipthth/eo ipioean?a a a?noi-aaiaoe/ieo nopoeoopao aeei/eiai? caeice ethaeeie ia p?cieo aoaiao aioop?oiueioopiaiiai picaeoeo ieiaea. ?ACOEUeOAOE AeINE?AeAEAHUe OA ?O IAAIAI?AIIss Iip?aiyeueiee aiae?c iopeiaieo a?noio?i?/ieo oa aeaeopiiiii?epineii?/ieo aeaiieo iieacaa, ui a ipaiaoaeueiiio iioiaaiac? ieiaea ethaeeie aeei/eiaa caeica ipioiaeeoue pyae epeoe/ieo iap?iae?a picaeoeo, eiaeiee c yeeo ia? iaai? iipoi-ooieoe?iiaeuei? iniaeeaino? aoaeiae p?ciiiai?oieo a?noi-aaiaoe/ieo nopoeoop. Anoaiiaeai? iap?iaee: ii/aoeiaee, aai aiap?iiaeueiee - 4-7 oeaei?a; ipii?aeiee, aai aiap?iiaeueii-oaoaeueiee - 7-12 oeaei?; i?ci?e, aai oaoaeueiee - 12 oeaeaeaiue oa a?eueoa oapaeoapecothoueny aeayeeie iipoieia?/ieie iciaeaie, ui a?ae?apaaeathoue iine?aeiai?noue picaeoeo a caeic? oipiiooaipthth/eo ipioean?a oa noaiiaeaiiy aeao?i?oeaieo nopoeoop. ?acoeueoaoe ipiaaaeaiieo aeine?aeaeaiue iieacothoue, ui aaaeeeaith iipoieia?/iith iciaeith iapoiai iap?iaeo picaeoeo ieiaea ? oa, ui ia 5 oeaei? aiap?iaaiaco aeei/eiaa caeica nii/aoeo ia? aoaeiao nooi ai?oae?aeueiiai ipaaio ? o?eueee i?ci?oa, oaea a e?ioe? 8 oeaeiy, a i?e iinooiiai c'yaeythoueny iapo? iiiaeeiie? e?ioioeeoe. Ai?oae?aeueiee capiaeie ipaaenoaaeaiee aeaiia oyaeaie aaaaoioapiaiai ai?oae?th, yeee picoaoiaaiee nii/aoeo a oeeiiio a?aeae?e?. A iiaeaeueoiio a?i picpinoa?oueny a aaiopaeueiiio iaipyieo. A ai?oae?ioeeoao oeaiopaeueieo a?aeae?e?a capiaeeo; ia a?aei?io a?ae ee?oei iapeoap?eiiai a?aeae?eo, c'yaeythoueny a?aepinoee , ui eaaeaoue o aeaeyae? nooe?eueiiai oapo ? iathoue oeee?iaep?/io oipio. Oeeoiieacia ee?oei aaaaoa peaiioeeaiipioa?aeaie, i?noeoue OEE-iiceoeai? ae?anoacipacenoaioi? apaioee, niinoap?aa?oueny i?aeaeuaiiy i?oioe/ii? aeoeaiino? ee?oei oa ca?eueoaiiy caaaeueiiai ai?noo yaeapii? AeHE. Noeoii?noue cacia/aieo iciae na?ae/eoue ipi aenie? iioaioe?? ai?oae?th ye c oi/ee cipo iiaeaeueoiai pinoo naii? caeice, oae ? a ca'yceo c picaeoeii ipioean?a naepaoiooaipaiiy. Ca iaoeie aeaieie, oeae ipioean c'yaey?oueny oaea ia 5 oeaei? aioop?oiueioopiaiiai picaeoeo ieiaea. sseui aeoiaeeoe c ipeiouaiiy iepaieo aaoip?a (Oeunoiaa C.N., 1987) ipi oa, ui a aiap?iaaiac? a caeic? c'yaey?oueny nii/aoeo aiaeiepeiia, a iio?i e?ioioeeoiiiaoe/ia ooieoe?y, oi iai caea?oueny eia?/iei ipeionoeoe, ui ? ipiaeoeo naepaoe?? nii/aoeo iiopaiey? a i?aeee?oeiiee ipino?p ai?oae?aeueii? caeeaaeee ipaaio, a iio?i oaea a caaaeueiee epiaioie, aieeaath/e iaaiei /eiii ia ooieoe?th ia/?iee - iapoiai epiaioaipiiai ipaaio a oeae iap?iae. ?acii c oei, iiyaa a e?ioe? eiai iapoeo e?ioioeeo?a, ye iai caea?oueny, piaeoue aaci?aenoaaiith naio iinoaiiaeo ieoaiiy ipi iine?aeiai?noue aeieeiaiiy ooieoe?e a aeei/eia?e caeic?. A oeueiio aeiaaeeo, iiaeeeai, a?eueo aeioe?eueiei aoei a picaeyaeaoe ?o ia a ieai? iapoi/apaiaino? picaeoeo a ye ipioeane aca?iii eiieopoth/?. Iipoieia?/i? iaapoiooaaiiy oe??? oace iiopaaothoue iiaeaeueoi? picpiaee, oiio ui a iniia? nai?e iathoue ipeiao nopoeoope iaae ooieoe??th, ui ia niinoap?aa?oueny a ipioeanao iipiaeueii? aeeoo?ae?yeueiino? ieiaea. A iapo?e iap?iae picaeoeo caeice aaaeeeaei ? oaeiae aea/aiiy ieoaiiy ipi iioiaeaeaiiy noiaaopiaeo ee?oei aai ee?oei - iiiapaaeiee?a O-e?ioioeeo?a, ye? i?ae aieeaii ipiaeoeo?a naepaoe?? iapaoaipththoueny a cp?e? O-ee?oeie (o?iioeeoe). A oeueiio ieoaii? ?niothoue ipioeeaaei? oi/ee cipo. Iaeia apoia aaoip?a aaaaea?, ui ia paii?o noaae?yo picaeoeo iie?iioaioi? noiaaopia? ee?oeie iinooiathoue a caeico c oieii epia?, oiaoi iathoue aaiaoiaaiia iioiaeaeaiiy (Eaeeieia E.E., 1983, Oeunoiaa C.N., 1987), ?ioa apoia ipeoiaeeoue aei aeniiaeo, ui a ca'yceo c a?aenooi?noth a oeae iap?iae noaeei, ee?oeie iinooiathoue a ipaai ipyii c iaaeieeoiuei? iacaio?ie, (Haar ?, 1974; Jenkinson O, 1981). Inoaiiy oi/ea cipo caea?oueny iai iaei eiia?piith, oiio ui a iaoeo aeine?aeaeaiiyo o ieiae?a 7,5 oeaei?a aioop?oiueioopiaiiai picaeoeo oaea c'yaeythoueny iapo? i?epinoaeeie, ye? iinooiiai aeiiaiththoueny apeopioeeoaie. A?eueoa oiai, aiae?c aeanieo aeaiieo iieaco?, ui ia ipioyc? iapoiai iap?iaeo a aeei/eia?e caeic? iino?eii niinoap?aathoueny iecueei aeeoapaioe?eiaai? ee?oeie, ye? ca nai?th oeueopanopoeoopith iiaeooue aooe a?aeianai? aei noiaaopiaeo ee?oei. A?aenooi?noue ?o a opaoueiio (i?ciueiio) iap?iae? picaeoeo ieiaea aeicaiey? ipeionoeoe oei/aniaee oapaeoap oeueiai ipioeano, ui iaoiiaeaii, iiaeeeai, iniaeeainoyie oipioaaiiy p?cieo eiiiiiaio?a caeice. Ca iaoeie aeaieie, oea ee?oeie aaeeeeo pici?p?a, c oeaiopaeueii picoaoiaaiei iooeei yaepii iaipaaeeueii iaaeueii? oipie. Eap?iieacia a ieo cia/ii ipina?oeaia, i?noeoue iacia/io e?euee?noue apoaeo apoaeie eiiaeainiaaiiai aaoapiopiiaoeio ?, ye ipaaeei, aeae?eueea yaeapaoeue. Aoopiiaoei aeeoocii picieeaiee a eap?iieaci? , iecueei? aeaeopiiii? u?eueiino?. Oeeoiieacia aaaaoa p?ciiiai?oieie aaeoieyie, iapaeiieie e?ciniiaie, iaaaeeeeo pici?p?a i?oioiiaep?yie. Ieanoei/anoee eiiieaen conop?/a?oueny p?aeei, aiaeiieaciaoe/iee paoeeoeoi iaaeinoaoiuei picaeiaiee. Iniaeea?noth aepoaiai (ipii?aeiiai, aai aiap?iiaeueii-oaoaeueiiai) iap?iaeo picaeoeo caeice ? ?ioaineaia, iniaeeai a iap?iae 10-12 oeaei?a, canaeaiiy ipaaio e?ioioeeoaie c ianooiiei iapapiciiae?eii ?o a oaea noipiiaaiiio eipeiaiio oa iiceiaiio oapao a caeaaeiino? a?ae nooiaiy aeeoapaioe?eiaee ee?oei . Oea icia/a?, ui a epaeia?e oa eipoeei-iaaeoeypi?e ciiao picoaoiaothoueny, a iniiaiiio iacp?e? (aeanoi?) oipie oeio aaeeeeo oa napaaei?o e?ioioeeo?a, a a iiceia?e pa/iaei? ciaoiaeyoueny iapaaaaeii cp?e?, aeeoapaioe?eiaai? oipie oeio iaeeo e?ioioeeo?a. Ca?eueoaiiy ieoiii? aaae e?ioi?aeii? oeaieie a oeae iap?iae noipiaiaeaeo?oueny iinooiiaei ciaioaiiyi ai?oae?aeueii? /anoeie ipaaio, ui caap?aa?oueny eeoa ii iapeoap?? capiaeea caeice o aeaeyae? aeania noaeainoeypii? ciie, ca paooiie yei? ? a?aeaoaa?oueny iiaeaeueoee pino ipaaio, oiio ui ooo iapaaaaeathoue ee?oeie c aenieei p?aiai i?oioe/ii? aeoeaiino?. Caapoa? ia naaa oaaao, ui iaeiei c iipoieia?/ieo ipiya?a oipiiooaipthth/eo ipioean?a a oeae iap?iae ? caiopiaoaaiiy ai?oae??i aeaiaio?a iacaio?ie pacii c epiaiiinieie i?epinoaeeiaie, oiaoi niinoap?aa?oueny oipioaaiiy aaiaoi-oeaetheypiiai aap'?po, yeee ia p?ai? aaiieai?eypa neeaaea?oueny c aiaeioae?th, aacaeueiiai oapo i?epinoaeei oa paoeeoeiai?oae?aeueieo ee?oei, ui picoaoiaai? ii oiaeo ?o ? iathoue aeanio aacaeueio iaiapaio. C iaeiiai aieo, aiia oipio? aeineoue /?oe? i?ae oeaieii? ipii?aeee, c ?ioiai - ia'?aeio? cacia/ai? aeaiaioe a ?aeeia oe?ea, oiaoi aap'?piee eiiieaen, ui iapaoeiaeaea? ipiieeiaiith o aioop?oi? napaaeiaeua ipaaio p?cieo iaepiiieaeoe, a oiio /ene? e oaeeo, ui ianooue aaiaoe/ii /oaeip?aeio ?ioipiaoe?th. A ipioean? picaeoeo caeice a oeae iap?iae noa? a?eueo aeaiiinopaoeaiei aca?iica'ycie paoeeoeiai?oae?aeueieo ee?oei oa e?ioioeeo?a. sse a?aei?/aeinue, a?i ipinoaaeo?oueny ua a iapoee iap?iae, iaeiae e?ioioeeoe a oeae /an ? iiiaeeiieeie, a ipioeane naepaoiooaipaiiy a paoeeoeiai?oae?ioeeoao ia/?oei aepaaeai?. Oiio a?eueo aaaeeeaei a aepoaee iap?iae picaeoeo ? ia no?eueee nopoeoopii aapeo?eiaai? ipiyae naepaoiooaipaiiy, ne?eueee nai iaoai?ci ipiieeiaiiy naepaoa (“o?i?/iiai oaeoipa”) a iaaeieeoi? i?aeee?oeiia napaaeiaeua, a a iiaeaeueoiio ? ca iaae? ipaaio. (Oeunoiaa C.N., Eaeeieia E.E., Oaaeinii A.O., 1993; Goya R.B. et. al., 1995). Haie ia iopeiaii iipoieia?/ii iaapoioiaaieo aeieac?a, ye? na?ae/eee a ipi iiaeeea?noue ipiieeiaiiy apaioe naepaoo aaciinapaaeiuei a ipina?o epiaiiiniiai eai?eypo aai iinoeai?eypii? aaioee. ?acii c oei, aeyaeai? aeoaea o?ni? eiioaeoe i?ae paoeeoeiai?oae?ioeeoaie oa no?ieith i?epinoaeei - ipioieai?eyp?a aai aoipeiieo eai?eyp?a, ipeioeeiiai? p?cieoe? i?ae yeeie iipoieia?/ii ia ?nio? (*apeania A.A., 1979). Oiio ? i?aenoaaa aeneiaeoe ipeiouaiiy, ui a?aenooi?noue o?iioeeo?a a neeaae? ai?oae?aeueiiai capiaeeo caeice ia paii?o noaae?yo (iapoee iap?iae) aiap?iaaiaco ? iiyaa ?o a cia/i?e e?eueeino? a aepoaee iap?iae ipe iayaiino? iia?/ieo iipoieia?/ieo iciae naepaoiooaipaiiy a paoeeoeiai?oae?ioeeoao na?ae/aoue ipi iiaeeea?noue aioop?oiueiipaaiii? oo?e?caoe?? ipiaeoeoa naepaoe??, oiaoi iiyao a ipaai? i?epiioi/aiiy, ui ?iaeoeo? e?ioioeeoiiiac oa aeeoapaioe?eiaeo O-ee?oei. Iipoieia?/iee aniaeo cacia/aii? i?aeee?oeiii? aca?iiae?? aea? iiaeeea?noue aecia/eoe ? aeaye? ?io? iciaee oipiiooaipthth/eo ipioean?a a aepoaee iap?iae aioop?oiueioopiaiiai picaeoeo ieiaea. Oae, iaie iino?eii niinoap?aaeenue paoeeoeiai?oae?ioeeoe (iapaaaaeii a iiceiaiio oap?), a oeeoiieaci? yeeo ciaoiaeeeenue iaci?iai? e?ioioeeoe. Ieaciieaie oeeo ee?oei /?oei eiioopothoueny, cia/iiai iioeiaeaeaiiy yaeap oa ipaaiae ?o ia anoaiiaeaii. Oe? iipoieiae/i? eapoeie ia aeeaaeathoueny a caaaeueiiipeeiyo? epeoap?? oaaioeeoapii? caeaoiino? ee?oei. Oiio, apaoiaoth/e /?oei niaoe?ae?ciaaio ooieoe?th e?ioioeeo?a caeice ye ?ioiieiiiaoaioieo ee?oei, ? i?aenoaaa aeneiaeoe ipeiouaiiy ipi iiaeeea?noue iapaaea/? a?aeiia?aeii? ?ioipiaoe?? naepaoipiei oeyoii. O?c?ieia?/i? aniaeoe oe??? ipiaeaie iiee-ui ia picpiaeai?. ?acii c oei, ia iniia? iopeiaieo aeaiieo a paii?e oa ipii?aeiee iap?iaee iipoiaaiaco caeice iiaeia picaeyaeaoe i?epieeaciaoic ye iaeei c iiaeeeaeo oeyo?a iapaaea/? ?ioipiaoe??. Iiae?aia ipeiouaiiy, cae/aeii, neoaeeoue ipaaeiaoii aeeneon??, oiio ui a iiae?aieo aeiaaeeao iiaa iiaea eoe o?eueee ipi iaeiioeii?noue nopoeoopieo ipiya?a ipioeano, ooieoe?iiaeueia cia/aiiy yeiai p?cia. Oae, opaaiaioe oeeoiieacie paoeeoeiai?oae?ioeeo?a, ye ? ipiaeoeoe poeioaaiiy yaeap e?ioioeeo?a, iino?eii niinoap?aathoueny ia ipioyc? anueiai aepoaiai iap?iaeo aioop?oiueioopiaiiai picaeoeo caeice. sseui aeoiaeeoe c oiai, ui ee?oeiiee aeaopeo iiaea aooe aeeipenoaiee aeey iiiaeaiiy oa picaeoeo iiaeo ee?oeiieo oipi, oi eia?/ii ipeionoeoe e iiaeeea?noue ?nioaaiiy iaaiiai iaoai?cio iapaaea/? a?aeiia?aeii? ?ioipiaoe?? ia ee?oeiiiio p?ai?. Aaaeeeao pieue a cacia/aieo ipioeanao, iiaeeeai, a?ae?apathoue iaepioaae, eiioaeoai yeeo c e?ioioeeoaie, ye a?aeiii, iaaea?oueny aaaeeeaa cia/aiiy a ?ioiieo paaeoe?yo ee?oeiiiai oa aoiipaeueiiai oeio (Carr I, 1978). Oaeei /eiii, a ipii?aeiee iap?iae oipiiooaipthth/? ipioeane a caeic? ? a?eueo neeaaeieie. Aiie oapaeoapecothoueny ?ioaineaiei ooaipaiiyi aeieueie (/anoi/ie), iiyaith eipeiai? oa iiceiai? pa/iaeie, ca?eueoaiiyi caaaeueiiai ai?noo e?ioioeeo?a oa ciaioaiiyi ieoiii? aaae ai?oae?aeueii? /anoeie ipaaio, iinooiiaei oipioaaiiyi aeao?i?oeaieo nopoeoop a neeaae? paoeeoeiai?oae?y, e?ioioeeo?a oa o?i?ei-aaiaoe/iiai aap'?po. A oeae iap?iae c'yaeythoueny iciaee, ui aeacothoue ia nopoeoopii-ooieoe?iiaeueio aaoapiaaii?noue paoeeoeiai?oae?y, a oaeiae iaiaeiip?aei?noue ee?oeiiiai neeaaeo e?ioi?aeii? oeaieie. I?ci?e (oaoaeueiee) iap?iae aioop?oiueioopiaiiai picaeoeo caeice oapaeoapeco?oueny iap?aiii?piei ? oaeeaiiae?aiei iapaa?aii oipiiooaipthth/eo ipioean?a. A oeae iap?iae niinoap?aa?oueny iinooiiaa cpinoaiiy aaae ipaaio ipe a?aeiini?e noaa?eueiino? a?eueoino? oaea noipiiaaieo a?noiaaiaoe/ieo nopoeoop: paoeeoeiai?oae?y, e?ioioeeo?a, o?i?ei-aaiaoe/iiai aap'?po, aaiii?epioeepeoeyoipiiai ponea, aeieueie (/anoi/ie) caeice. A oeae iap?iae niinoap?aa?oueny oaeaeeee p?no iiceiai? pa/iaeie oa, a?aeiia?aeii, ? e?eueeino? o?i?/ieo o?eaoeue. Aei inoaiiueiai /ano aaaaeaeinue, ui c iiceoe?e eeane/ieo oyaeaiue a iniia? ?o oipioaaiiy eaaeaoue aiaieuaa? neoi/aiiy paoeeoeiai?oae?ioeeo?a, ye? a iiaeaeueoiio, caciath/e a?aeiia?aeieo ci?i, oipiothoue nopoeoope o aeaeyae? eiioeaiope/ii picoaoiaaieo ipioape?a niethuaieo ee?oei c oeaiopaie ?c cpiaia?eeo aeaiaio?a. Iiae?aia oi/ea cipo, yea pai?oa caeaaaeany aaccaiapa/iith, iinooiiai ii/eia? aopa/aoe nai? cia/aiiy. Oae, iieacaii, ui a ooaipaii? o?eaoeue ipeeiathoue o/anoue oaeiae ? iaepioaae (Policard A, 1965). Jaroslow B.N. (1967) aeneiaea ipeiouaiiy, ui a o?i?/ieo o?eueoeyo aepiaey?oueny naepao, ui iiopaiey? iio?i a epiaiiinio oa e?ioaoe/io nenoaio. ?nio? oaeiae oi/ea cipo, ui o?i?/i? o?eueoey ? i?noeai neoi/aiiy caaeaeeo e?ioioeeo?a, aea a?aeaoaa?oueny ciaoeiaeaeaiiy oa aeaeaeaiiy aioeaaiieo noanoaioe?e (Blau I., 1973). Ep?i oiai, aiie iaaiei /eiii aieeaathoue ia picaeoie ai?oae?aeueiia capiaeeo caeice (Oeunoiaa C.N., 1987) ? iiaeooue noeioethaaoe ipioean cpiaia?iiy ee?oei (Mandel T., 1969). Ha iniia? aiae?co aeanieo aeaiieo ? i?aenoaaa picaeyaeaoe o?i?/i? o?eueoey ia o?eueee ye noaoe/i?, a ye ? iaaiith i?pith aeeiai?/i? nopoeoope, iiyaa yeeo ia iepaieo aoaiao iipoiaaiaco caeice ? aaiaoe/ii aeaoapi?iiaaiei ipioeanii, ui a?aeiapaaea? caaaeueiia?ieia?/i? caeiiii?piino? picaeoeo, pinoo oa aeeoapaioe?eiaee ee?oeiieo aeaiaio?a iiceiaiai oapo. Cacia/aia ipeiouaiiy iia?/ii i?aeoaapaeaeo?oueny ? aeaiieie e?oapaoope, ca?aeii yeeo anoaiiaeaia aeniea aeeiai?/i?noue ee?oeiiiai neeaaeo caeice, yeee (iniaeeai a iinoiaoaeueiee iap?iae) iiaea ci?ithaaoenue ipioyaii aeiae (Policard A., 1965). Iieacaia iiaeeea?noue oaeaeeiai cieeiaiiy oa iiyae iiaeo o?i?/ieo o?eaoeue, aeyaeaii p?cio e?euee?noue ?o ia ipioyc? anueiai iap?iaeo aioop?oiueioopiaiiai picaeoeo ieiaea anoaiiaeaii eieae?caoe?th ?o o?eueee a iiceia?e pa/iaei?, a oaeiae caapiaii oaaao ia iiyao aeineoue neeaaeieo ee?oeiieo eiiiapaoe?e a pacoeueoao? ?o ceeooy i?ae niaith o aeaeyae? aianoiiicoth/eo nopoeoop (Oeunoiaa C.N., 1987; Aipcye Y.E., Aieioei H.A., Eapcia I.A. oa ?io., 1990; Caepaoueyioe I.A., 1991 oa ?io.). C apaooaaiiyi cacia/aiiai iiaeia oiiaii aeae?eeoe ope iiaeeea? noaae?? picaeoeo o?eaoeue: a) noaae?y ?i?oe?aoe??, aai ii/aoeiaa noaae?y; a) noaae?y noaa?e?caoe??; a) “noaae?y ?iaiethoe??”. Eiaeia c iacaaieo noaae?e oapaeoapeco?oueny iaaieie iipoi-ooieoe?iiaeueieie iciaeaie oa aeayeeie iniaeeainoyie oeueopanopoeoope ee?oeiieo aeaiaio?a. Aiae?c ?o aea? iiaeeea?noue oiiaii aeae?eeoe a neeaae? o?eaoeue ope ciie: oeaiopaeueio, ipii?aeio oa iapeoap?eio. Ii/aoeiaa, aai ?i?oe?aeueia, noaae?y oapaeoapeco?oueny iiyaith eieaeueieo oapoiiaaiue c paoeeoeiai?oae?ioeeo?a, ui noipiaiaeaeo?oueny ciaioaiiyi i?aeee?oeiiiai ipinoipo, iaaoooyi ee?oeiaie aeaeiaaeaii? oipie, ci?iith eiio?aopaoe?? ?o yaeap, iap?aiii?piei iapapiciiae?eii a ieo ao- oa aaoapiopiiaoeio, picoepaiiyi oa opaaiaioaoe??th eaiaeueoe?a oeeoiieaciaoe/ii? n?oee, iiyaith a ee?oeiao iepaieo aaeoieae oa iaaaeeeeo apoi i?epiaaceeoe. Ha noaae?? noaa?e?caoe?? iienai? ci?ie a?eueo /?oei aepaaeai? ? iinyoue piciianthaeaeaiee oapaeoap. Ipe oeueiio a neeaae? o?eueoey ca?eueoo?oueny e?euee?noue ee?oei (aei 6-8), yaepa noathoue aeaeiaaeaii? oipie c iaaiipyaeeiaaiei picoaooaaiiyi aaeeeeo apoaeie aaoapiopiiaoeio. Oeeoiieacia paoeeoeiai?oae?ioeeo?a iinooiiai caoaeo?oueny, iaaoaa? aeaeyaeo nioaeee p?cii? oiaueie, picoaoiaaii? ii iapeiaopo yaepa. A oeeoiieaci? iepaieo paoeeoeiai?oae?ioeeo?a conop?/athoueny iaci?iai? e?ioioeeoe. Aiae?c iopeiaieo aeaieo aeicaiey? caiapa/oaaoe iaepioaaaeueio ooieoe?th paoeeoeiai?oae?aeueieo ee?oei, ye? aoiaeyoue aei neeaaeo o?i?/ieo o?eaoeue. ?acii c oei, a oeae iap?iae a ieo iino?eii niinoap?aathoueny iaepioaae, picoaoiaai? ia iaae? oeaiopaeueii? oa ipii?aeii? cii, a oaeiae iepai? e?ioioeeoe, ye? ia iathoue aea?peiai? eieae?caoe??. “?iaiethoeaia” noaae?y ia ? iine?aeiaiei aoaiii iiiapaaeiuei?, oiio ui oapaeoapi? aeey ia? iipoieia?/i? iciaee iiaeooue c'yaeyoeny ia naiiio ii/aoeo ?i?oe?aeueii? noaae??. Nipaaa a oiio, ui ana caeaaeeoue a?ae oaeaeeino? iiyae inoaiiuei? ?, c ?ioiai aieo - a?ae nooiaith aepaaeaiino? aeenopio?/ii-aeanopoeoeaieo ci?i a paoeeoeiai?oae?ioeeoao. ?ioeie neiaaie, oey noaae?y iiaea aooe p?ciiiai?oiith ca nai?i iipoieia?/ieie ipiyaaie a caeaaeiino? a?ae iapaaaaeaiiy a nopoeoop? o?i?/ieo o?eaoeue o??? /e ?ioi? ciie. Inoaii? iiaeooue aooe ipaaenoaaeai? o aeaeyae?: iapaaaaeii oeaiopaeueii? ciie, a ye?e ciaoiaeyoueny paoeeoeiai?oae?ioeeoe c iciaeaie aeenopio?/ii-aeanopoeoeaieo ci?i; ipii?aeii? ciie, ui oapaeoapeco?oueny iiyaith eiioeaiope/ii picoaoiaaieo niethuaieo ee?oei; iapeoap?eii? ciie, ui ipaaenoaaeaia picoaoiaaieie ii iapeiaopo o?eueoey ee?oeiaie iiceiai? pa/iaeie, ye? a?aep?ciythoueny a?ae aiaeia?/ieo ee?oei aeania caeice o?eueee a?eueo o?niei aca?iiei picoaooaaiiyi. Oaeei /eiii, o?i?/i? o?eueoey ? aaaeeeaeie nopoeoopith aeei/eiai? caeice, ye? iathoue iaaio aeeiai?eo picaeoeo, ipeeiath/e o/anoue a ipioeanao aeanopoeoe?? oa ae?i?iaoe?? ipiaeoeo?a piciaaeo e?ioioeeo?a. ?o o?nia aca?iiae?y c ?ioeie eiiiiiaioaie caeice ? a?ieia?/ii aeioe?eueiith, oiio ui c iaeiiai aieo a?aeiapaaea? p?aaiue ?ioiieia?/ii? aeoeaiino? ipaaio a oe?eiio, c ?ioiai - caaacia/o? eiipaeeioth/o ooieoe?th eiai a noaiiaeaii? oipiiooaipthth/eo ipioean?a ia p?cieo aoaiao aioop?oiueioopiaiiai picaeoeo ieiaea ethaeeie. AENHIAEE 1. ?icaeoie aeei/eiai? caeice a ipaiaoaeueiee iap?iae iipoiaaiaco ? aneiopiiiei, oaeeaiiae?aiei ipioeanii, ui oapaeoapeco?oueny ipiapanoth/ei ca?eueoaiiyi e?euee?nieo iieaciee?a pinoo (iane oa pici?p?a) c iiyaith ye?nii iiaiai noaio ipaaia, ui a?aeiapaaeathoue iinooiiaa oipioaaiiy eiai aeao?i?oeaieo nopoeoop. 2. Iepai? iap?iaee picaeoeo caeice a?aeiapaaeathoue ye?nii iia? aoaie oipioaaiiy ?? a?noiapo?oaeoii?ee ? iiaeooue aooe oiiaii aeae?eai? a caeaaeiino? a?ae iipoi-ooieoe?iiaeueieo iciae: paii?e (aiap?iiaeueiee) iap?iae 4-7 oeaei?a; ipii?aeiee (aiap?iiaeueii-oaoaeueiee) iap?iae - 7-12 oeaei?; i?ci?e (oaoaeueiee) iap?iae - c 12 oeaeiy oa aei iiiaioo iapiaeaeaiiy. 3. Iniiaiiith iciaeith iapoiai iap?iaeo ? aeicp?aaiiy oa aeeoapaioe?eiaea ee?oei ai?oae?aeueiiai capiaeeo c iiyaith iciae ?o naepaoipii? aeoeaiino?; aipiaeiaae aepoaiai iap?iaeo a?aeaoaa?oueny iinooiiaa canaeaiiy ipaaio e?ioi?aeieie aeaiaioaie c iaaoooyi ieie aeanoeainoae ?ioiieiiiaoaioieo ee?oei, ui iia?/ii i?aeoaapaeaeo?oueny iiyaith o ieo i?aeee?oeiieo eiioaeo?a c paoeeoeiai?oae??i oa iaepioaaaie; ia ipioyc? opaoueiai iap?iaeo niinoap?aa?oueny oipioaaiiy aeao?i?oeaieo a?noiaaiaoe/ieo nopoeoop, ui noipiaiaeaeo?oueny ipiapaneaiei ca?eueoaiiyi iane ipaaio. 4. Iniiaieie neeaaeiaeie eiiiiiaioaie caeice a ipaiaoaeueiee iap?iae iipoiaaiaco ? niaoe?ae?ciaai? ee?oeii? aeaiaioe (paoeeoeiai?oae?e, e?ioi?aeia oeaieia) oa o?i?ei-aaiaoe/iee aap'?p, yeee ia p?ai? aaiieai?eyp?a ipaaenoaaeaiee ai?oae?ioeeoaie, aacaeueiei oapii oa iniiaiith pa/iaeiith, c picoaoiaaieie a i?e iapeoeeoaie, a oaeiae paoeeoeiai?oae?ioeeoaie, ui iathoue aeanio aacaeueio iaiapaio. Aca?iica'ycee oeeo eiiiiiaio?a aecia/athoue ye iniaeeaino? nopoeoope naii? caeice, oae ? p?ciiiai?oi? ooieoe?? ?? ee?oeiieo aeaiaio?a ia p?cieo aoaiao aioop?oiueioopiaiiai picaeoeo ieiaea ethaeeie. 5. Ai?oae?aeueiee capiaeie caeice a iipoiooieoe?iiaeueiiio a?aeiioaii? oyaey? niaith aaoapiaaiio iiioeyoe?th ee?oei, ye? iathoue ai?oae?aeueia oa iacaio?iaeueia iioiaeaeaiiy. Ai?oae?aeueia ipepiaea ?o i?aeoaapaeaeo?oueny iniaeeainoyie oeueopanopoeoope (iayai?noue aeaniinii, oiiio?ap?e, aaeoieae), a oaeiae iia?/ieie iipoieia?/ieie iciaeaie ipioeano naepaoiooaipaiiy. Aioiaeapiaeueia iioiaeaeaiiy iepaieo noaiiioeyoe?e i?aeoaapaeaeo?oueny iayai?noth a neeaae? caeice paoeeoeypieo oaaioeeooth/eo ee?oei, ui iathoue oeiiao aoaeiao iaepioaa?a. Aiie caaacia/othoue ia o?eueee aioop?oiueiee?oeiia poeioaaiiy oa ae?i?iaoe?th ipiaeoeo?a piciaaeo ee?oei ca iaae? ipaaio, aea e (a oiiaao aioeaaiii? noeioeyoe??) iiaeeeaee iai?i a?aeiia?aeiith ?ioipiaoe??th oa o/anoue a eiiiapaoe?? c e?ioioeeoaie ipe picaeoeo ?ioiieo paaeoe?e. 6. ?aoeeoeiai?oae?aeuei? ee?oeie eipe oa iiceiai? pa/iaeie iathoue aeaye? iniaeeaino? oeueopanopoeoopii? aoaeiae yaepa oa oeeoiieacie, ui iia?/ii i?aeoaapaeaeo? ?o o/anoue a ipioeanao naepaoiooaipaiiy (ci?ia pici?p?a oa eiio?aopaoe?? yaepa, iaeiie/aiiy a oeeoiieaci? p?ciiiai?oieo aaeoieae oa aeaeopiiiiou?eueiaieo apaioe, aioop?oiueiee?oeiia a?iapieac?y aeaiaio?a ieanoei/anoiai eiiieaeno oa aiaeiieaciaoe/iiai paoeeoeoio c iaapiiaaeaeaiiyi a eaiaeueoeyo aep?aiicapienoiai oeieoeyoo. Aep?oaiiy ieoaiiy ipi oapaeoap cacia/aii? naepaoipii? ooieoe?? (aiaeiepeiii? aai e?ioiiiaoe/ii?), a oaeiae anoaiiaeaiiy iine?aeiaiino? ?o picaeoeo o?eueee ia i?aenoaa? oeueopanopoeoopieo aeaiieo ia oyaey?oueny iiaeeeaei. 7. Iniaeeaino? oiiiapao?? e?ioi?aeieo aeaiaio?a a?aeiapaaeathoue nipyiiaaii?noue ipioeano ?o aeicp?aaiiy: ipe oeueiio a eipeia?e pa/iaei? picoaoiaothoueny iacp?e? (aeanoi?) oipie ee?oei, a a iiceia?e pa/iaei? - a?eueo aeeoapaioe?eiaai? e?ioioeeoe. Iiae?a?i? aca?iia?aeiioaiiy aiapoa c'yaeythoueny ia aoai? canaeaiiy caeice e?ioioeeoaie ? a iiaeaeueoiio caap?aathoueny noaa?eueii ia ipioyc? anueiai aioop?oiueioopiaiiai iap?iaeo picaeoeo ieiaea. 8. ?aoeeoeiai?oae?e a neeaae? o?i?eiaaiaoe/iiai aap'?po caeice ia ? nooe?eueiei aipiaeiaae anueiai eai?eypiiai ponea, ui iaoiiaeth? iiyao o?nieo eiioaeo?a i?ae e?ioioeeoaie oa aacaeueiei oapii aaiieai?eyp?a aai iinoeai?eypieo aaioe. ?acii c oei, iipoieia?/ii ia aaea?oueny ipinoaaeeoe ipioean ?o i?apaoe?? /apac no?ieo cacia/aieo noaeei. A oie aea /an, aeoeaaoe?y aiaeioae?y (iaapye i?oioiiaep?e, a?iapieac?y aeaiaio?a ieanoei/anoiai eiiieaeno oa capienoi? oeeoiieaciaoe/ii? n?oee, iaeiie/aiiy a oeeoiieaci? i?epiaaceeoe oa aaeoieae, picaeoie i?epieeaciaoico oa ?io.) iia?/ii i?aeoaapaeaeo? o/anoue oeeo ee?oei a ipioeanao opaineai?eypiiai opainiipoo p?cieo pa/iaei oa iaoaaie?o?a. A ca'yceo c oeei iaia? i?aenoaa caiapa/oaaoe iiaeeea?noue opainaiaeioae?aeueii? aai e?ioiai?oae?aeueii? i?apaoe?? e?ioioeeo?a, ui iae/ano?oa niinoap?aa?oueny ia p?ai? iinoeai?eypieo aaioe a iiceiaiio oap? caeice. 9. O?i?/i? o?eueoey, aai o?eueoey Aannaey, iiaeia picaeyaeaoe ? ye aeeiai?/i?, a ia o?eueee noaoe/i? nopoeoope aeei/eiai? caeice. Aiapoa aiie c'yaeythoueny a e?ioe? 12 oeaeiy aioop?oiueioopiaiiai picaeoeo ieiaea, a a iiaeaeueoiio e?euee?noue ?o cia/ii aap'th?. A nai?io picaeoeo aiie oiiaii ipioiaeyoue ope noaae??: ?i?oe?aoe??, noaa?e?caoe?? oa “?iaiethoe??”, ye? a?aeiapaaeathoue iipoieia?/i? (a?noio?i?/i? oa oeueopanopoeoopi?) iniaeeaino? oipiiooaipthth/eo ipioean?a a caeic? a p?ci? iap?iaee aioop?oiueioopiaiiai picaeoeo ieiaea. 10. Aei neeaaeo o?i?/ieo o?eaoeue aoiaeyoue paoeeoeiai?oae?aeuei? ee?oeie, a oaeiae iaepioaae oa e?ioioeeoe, ui oipiothoue oeaiopaeueio , ipii?aeio oa iapeoap?eio ciie. A oe?eiio, paoeeoeiai?oae?th ia ipeoaiaii? oaaioeeoapi? aeanoeaino? oa o/anoue a ipioeanao naepaoiooaipaiiy. Haia? oaeiae i?aenoaa ipeioneaoe noaeeiia iioiaeaeaiiy o?eaoeue, oiio ui aeenopio?/ii-aeanopoeoeai? ci?ie ee?oei, iathoue i?noeaaa, oiaoi a?no?iaaiia, a ia aaiaoiaaiia iioiaeaeaiiy. NIENIE IOAE?EAOe?E CA OAIITH AeENA?OAOe?? 1. Aeanopoeoeey oeiioeeoia a aeei/eiaie aeaeaca ieiaeia /aeiaaea. Oace aeiiia?aeae ? iaoe?ii. eiiap. aiao. a?noie. aiap?ii?a Oepa?ie ?aaii - Opaie?anuee,1994, n. 15 2. Ii?oieia?y aeei/eiai? caeice a oaoaeueiiio iap?iae? iipoiaa- iaco. A ei. Aeooaeuei? ieoaiiy iipoiaaiaco. (iaoap?aee iaoeiai? eiioapaioe??). *api?aoe?, 1966, n. 392-393 (ni?aaao. Oiiaiei A.A.) 3. Oeueopanopoeoopa aeei/eiai? caeice a p?ci? iap?iaee aioop?oiueioopiaiiai picaeoeo. A ei.: Aeooaeuei? ieoaiiy iipoi- eia?? oa ee?i. iaae. (iaoeia? ipaoe?). Aei?ipiiaopianuee, 1966, n. 55-59. 4. Iipoieia?y aeei/eiai? caeice a aiap?iiaeueiiio iap?iae? iipoiaaiaco. A ei.: Aeo. ieoaiiy iipoieia?? oa ee?i. iaae. Aei?ipiiaopianuee 1966, n. 63-65. 5. Oeueopanopoeoopa aaiaoioeaetheypiiai aap'?po aeei/eiai? caei- ce ieiaea ethaeeie. // Oeno. - *api?aoe?. - 1977. - N 1, n. 50-56. 6. Iniaeeaino? a?noiaia?iapo?oaeoii?ee oeiona a ipaiaoaeueiiio iap?iae? iioiaaiaco. // Oeno. - *api?aoe?. - 1977. - N 1, N. 42-50. 7. I?epiiipoieia?y aeei/eiai? caeice a p?ci? iap?iaee aioop?oiueioopiaiiai picaeoeo ieiaea. //Ieoaiiy aaeaieia?? ? aeieia?? a opaae. oa iaopaae. iaae. (iaoeia? ipaoe?). - Aei?ipiiaopianuee. - "Iie?apao?no". - 1977. - N. 224-226. 8. Aeaye? iniaeeaino? oeueopanopoeoope aeei/eiai? caeice a ai- oaiaoaeueiee iap?iae. - Ieoaiiy aaeaie. ? aeieia?? a opaae. oa iaopaae. iaae. (iaoeia? ipaoe?). - Aei?ipiiaopianuee. - 1997. - N. 214-215. 9. Noaiiaeaiiy nopoeoope aeei/eiai? caeice ethaeeie a ipaia- oaeueiiio iap?iae? iioiaaiaco // Iaoap?aee ca?o. iaoe. eiio. aeeeaaea/?a OAeIO ?i. I.I. Aepaaiiaiiaa. - Ee?a: "Oieiea". - 1995. - 461-462. AIIOAOe?ss Aeaaiaeapiaa I.A. Ooieoe?iiaeueia iipoieia?y aeei/eiai? caeice ethaeeie a ipaiaoaeueiee iap?iae iipoiaaiaco. - ?oeiien. Aeenapoaoe?y ia caeiaoooy iaoeiaiai nooiaiy eaiaeeaeaoa a?ieia?/ieo iaoe ca niaoe?aeuei?noth 14.03.01. - iipiaeueia aiaoii?y. - Haoe?iiaeueiee iaaeaaia?/iee oi?aapneoao ?i. I.I. Aepaaiiaiiaa, Ee?a, 1977. Aeenapoaoe?y ipena'y/aia aea/aiith ooieoe?iiaeueii? iipoieia?? aeei/eiai? caeice ieiaea ethaeeie a p?ci? iap?iaee eiai aioop?oiueioopiaiiai picaeoeo. Aeyaeai? ye?nii iia? iipoi-ooieoe?iiaeuei? iciaee a?noiaaiaoe/ieo nopoeoop caeice, ye? aeathoue iiaeeea?noue picpiaeoe iia? i?aeoiaee a aep?oaii? ieoaiue, ui noinothoueny naepaoipii? ooieoe?? caeice, iioiaeaeaiiy oa i?apaoe?? e?ioioeeo?a, iniaeeainoyo aoaeiae aaiaoioeaeetheypiiai aap'?po ia p?ai? aaiieai?eypa, oipioaaii? o?i?/ieo o?eaoeue oa ?io. Ipaaenoaaeai? aeaii? ipi nopoeoopii-ooieoe?iiaeueio aaoapiaaii?noue paoeeoeiai?oae?y aeei/eiai? caeice, iienai? inia- eeaino? i?aeee?oeiieo aca?iia?aeiinei, aeneiaeaii ipeiouaiiy ipi cia/aiiy o?i?/ieo o?eaoeue a ipaiaoaeueiiio iap?iae? iipoiaaiaco. Eeth/ia? neiaa: aeei/eiaa caeica, iipoiaaiac, ieiae, a?noio?i?y, oeueopanopoeoopa, e?ioioeeo, paoeeoeiai?oae?ioeeo, aaiieai?eyp. AIIIOAOeEss Aeaaiaeapiaa A.A. Ooieoeeiiaeueiay iipoieiaey aeei/eiaie aeaeacu /aeiaaea a ipaiaoaeueiue iapeiae iipoiaaiaca. ?oeiienue. Aeennapoaoeey ia nieneaiea o/aiie noaiaie eaiaeeaeaoa aeiei- ae/aneeo iaoe ii niaoeeaeueiinoe 14.03.01 - iipiaeueiay aiaoiiey. Haoeeiiaeueiue iaaeaaiae/aneee oieaapneoao ei. I.I. Aepaaiiaiiaa. - Eeaa, 1997. Aeennapoaoeey iinayuaia eco/aieth ooieoeeiiaeueiie iipoieiaee aeei/eiaie aeaeacu ieiaea /aeiaaea a pacee/iua iapeiaeu aai aioopeoopiaiiai pacaeoey. Auyaeaiu ea/anoaaiii iiaua iipoi- ooieoeeiiaeueiua ipeciaee aenoiaaiaoe/aneeo nopoeoop aeaeacu, eioipua iicaieytho pacpaaioaoue iiaua iiaeoiaeu a paoaiee aiipinia, eanathueony naepaoipiie ooieoeee aeaeacu, ipienoiaeaeaiey e ieapaoeee eeioioeeoia, iniaaiiinoyo nopiaiey aaiaoi-oeaeetheypiiai aapueapa ia opiaia aaiieaieeypa, oipiepiaaiee oeie/aneeo oaeaoe e ip. Ipaaenoaaeaiu aeaiiua i nopoeoopii-ooieoeeiiaeueiie aaoapiaaiiinoe paoeeoeiaieoaeey aeei/eiaie aeaeacu, iienaiu ini- aaiiinoe iaaeeeaoi/iuo acaeiiioiioaiee, auneacaii ipaaeiieiaeaiea i cia/aiee oeie/aneeo oaeaoe a ipaiaoaeueiii iapeiaea iipoiaaiaca. Eeth/aaua neiaa: aeei/eiaay aeaeaca, iipoiaaiac, ieiae, aenoioeiey, oeueopanopoeoopa, eeioioeeo, paoeeoeiaieoaeeioeeo, aaii- eaieeyp. ANNOTATION Blagodarova E.V. Functional morphology of timus in prenatal period of morphogenesis. Manuscript. Dissertation for scientific degree of candidate of biological sciences 14.03.01. - normal anatomy. Kiev, 1997. It has been determiened a certain seguence of forming of definite structures of thymus of foetus in different periods of development. It has been revealed new morpho-functional features of thymus gystohaematic structures, wich give possibilities to solve problems concerning thymus secretory function, origin and migration of lymphocytes, pecularities of haemo-celluar barrier on the level of haemocapillar, forming of timus bodies etc. Data about structure-functional heterogenity of rethiculoepithely of intercellar relations are descriebed, the process of forming of thymus bodies and their importance in human foetus development is traced. Key words: thymus, morphogenesis, foetus, gystohaemic, ultrastructural, lymphocyte, rethiculoepitheliocyte, haemocapilare.

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